GEOMETRIC MORPHOMETRIC APPROACH TO ESTABLISH PHYLOGENETIC AFFINITIES OF ENIGMATIC PTEROSAUR SPECIMENS FROM THE LOWER CRETACEOUS OF SOUTH KOREA

Chan-gyu Yun
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Abstract

Fragmentary but scientifically important pterosaur fossils have been reported in Lower Cretaceous strata such as the Jinju and Hasandong formations of South Korea. Recently, several large teeth and a second wing phalanx found in these strata have been suggested to have affinities with Boreopteridae, a clade in which has so far been reported only in the Yixian Formation of China. Here, the phylogenetic affinities of these proposed boreopterid specimens are reassessed, using two-dimensional geometric morphometric shape analyses. The morphometric analyses found the South Korean pterosaur teeth in different morphospaces from the Yixian Fm. boreopterids, and plot them closer to anhanguerians. An isolated second wing phalanx does plot close to a boreopterid Zhenyuanopterus longirostris in a morphospace defined by first two principal component axes, but taxa from different lineages are also plotted as closely, or even closer. Thus, there seems to be no clear evidence that these specimens belong to Boreopteridae, and the tooth fossils appear to be very different from those of established members of this clade. Therefore, the hypothesis that some South Korean pterosaur fossils can be referred to boreopterids, is here tentatively rejected.
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用几何形态计量学方法确定韩国下白垩统神秘翼龙标本的系统发育亲缘关系
在下白垩统地层(如韩国的晋州地层和哈山东地层)中发现了一些零碎但具有重要科学价值的翼龙化石。最近,在这些地层中发现的几颗大牙齿和第二翼趾骨被认为与蝶龙科(Boreopteridae)有亲缘关系。在此,我们利用二维几何形态分析重新评估了这些拟议的膛翅目标本的系统发育亲缘关系。形态分析发现,南朝鲜翼龙的牙齿与黟县地层的镗翼龙牙齿处于不同的形态空间,并将它们与安汉古龙类划定为更接近的类群。在由前两个主成分轴定义的形态空间中,一个孤立的第二翼趾骨确实与北缘翼龙(Zhenyuanopterus longirostris)接近,但来自不同类群的类群也与之接近,甚至更接近。因此,似乎没有明确的证据表明这些标本属于伯劳蝶科,而且这些牙齿化石似乎与该支系已有成员的牙齿化石非常不同。因此,南韩的一些翼龙化石可归属于膛翅目的假说在此被暂时否定。
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CHALLENGES IN ORBITOLINID IDENTIFICATION AND BIOSTRATIGRAPHIC CONSEQUENCES: AN APTIAN EXAMPLE BELOPTERA LONGA NAEF, 1922 (CEPHALOPODA: COLEOIDEA) FROM THE LATE EOCENE (PALAEOGENE) OF UKRAINE PALYNOSTRATIGRAPHIC AND PALAEOENVIRONMENTAL INVESTIGATIONS OF THE MAASTRICHTIAN FROM OARDA DE JOS (SOUTHWESTERN TRANSYLVANIAN BASIN) GEOMETRIC MORPHOMETRIC APPROACH TO ESTABLISH PHYLOGENETIC AFFINITIES OF ENIGMATIC PTEROSAUR SPECIMENS FROM THE LOWER CRETACEOUS OF SOUTH KOREA NEW MIDDLE TRIASSIC BELL-SHAPED NASSELLARIAN RADIOLARIA FROM ALPINE AND CARPATHIAN AREAS
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