MORPHOLOGICAL CRITERIA FOR THE DIFFERENTIATION OF AGE STATUS OF PTILOSTEMON ECHINOCEPHALUS (WILLD.) GREUTER (LAMYRA ECHINOCEPHALA (WILLD.) TAMAMSCH.)

E. Zadneprovskaya, L. P. Vakhrusheva
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Abstract

Ptilostemon echinocephalus (Willd.) Greuter (Lamyra echinocephala (Willd.) Tamamsch.) is the only species of the genus Ptilostemon from the Asteraceae family that grows on the Crimean Peninsula. The relic area of Ptilostemon echinocephalus distributed by the Crimea, the Caucasus, Asia Minor. The species is included in the Red Book of the Republic of Crimea, Krasnodarskiy and Stavropolskiy Krays of the Russian Federation. The article is the result of a long-term study of morphological criteria of Ptilostemon echinocephalus age status. Thus, the ontogenesis of individuals of the species Ptilostemon echinocephalus includes 12 age states, the total during of ontogenesis lasts 15–18 years. The main characters of the pregenerative period differentiation of development are the number of leaves and the differentiation degree of the leaves lamina. During the study, another important feature was identified to indicate different age stages, the ratio of vegetating and dried leaves. Generative plants were subdivided into: immature generative (g1), adult (mature) generative (g2) and old generative (g3). For the generative stage of development, the ratio of generative and vegetative rosettes, the ratio of dried and vegetative shoots, and the number of baskets were indicative. Differentiation of subsenile and senile age states is conducted according to the same criteria that are used for generative individuals, the main emphasis is on the ratio of dried and vegetative parts.The article also notes the specificity of the course of ontogenesis depending on the ecological-cоenotic conditions in the petrophytic steppes and phrygana communities of the Crimea.
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野生棘头子茎年龄分化的形态学标准棘头层膜(野生)TAMAMSCH)。
棘头天竺葵(野生)格鲁特(野生刺头拉米)(Tamamsch.)是菊科中唯一的一种,生长在克里米亚半岛。棘头火车头遗迹分布于克里米亚、高加索、小亚细亚一带。该物种被列入俄罗斯联邦克里米亚共和国,克拉斯诺达尔斯基和斯塔夫罗波尔斯基克的红皮书。本文是对棘头天竺葵年龄状况的形态学标准进行长期研究的结果。因此,棘头拟南棘个体的个体发生经历了12个年龄阶段,个体发生的总时间为15 ~ 18年。发育前生殖期分化的主要特征是叶片数量和叶片层分化程度。在研究过程中,确定了指示不同年龄阶段的另一个重要特征,即植被和干叶的比例。生殖植物分为:未成熟生殖(g1)、成年(成熟)生殖(g2)和老生殖(g3)。在生殖发育阶段,生殖莲座与营养莲座之比、干枝与营养枝之比、花篮数具有指示性。亚衰老和老年状态的区分依据与生殖个体相同的标准进行,主要重点是干燥部分和营养部分的比例。文章还指出了个体发生过程的特殊性,这取决于克里米亚岩生草原和芦苇群落的生态-生态条件。
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