New Taxa of Damselflies (Protozygoptera) from the Permian of the East European Platform

Abstract

A new classification system for Protozygoptera is proposed. It is based on mass material from the Isady, Chekarda and Soyana localities. The new system is based on a set of wing venation characters. The hypothesis on the division of Protozygoptera based on their body structures is supplemented and confirmed (Nel et al., 2012). A three-segmented tarsal structure has been established for the archizygopteran family Kennedyidae. The infraorder Permagrionomorpha infraordo nov. is established. An additional character has been proposed to separate archizygopterans and permagrionomorphs based on the structure and morphology of the thorax. Lodeviidae is synonymized with Permagrionidae (junior synonym). Two new families, four new genera and 14 new species are described. A new family Progoneuridae fam. nov. is described based on a significant reduction in the wing longitudinal and transverse venation. It includes the genera Progoneura Carpenter, 1931; Azaroneura Nel et al., 2012; Progophlebia Felker, 2021, and Issadoneura gen. nov. (with a new species Issadoneura marilevorum gen. et sp. nov.). Based on re-examination of the type material from Chekarda, the genus Engellestes Nel et al., 2012 is assigned to a new family Engellestidae fam. nov. The following voltzialestids are described from the Upper Permian of the Vologda Region: Prototerskeja dubia gen. et sp. nov., Permosticta elegans gen. et. sp. nov., and Permosticta parva gen. et sp. nov. The diagnosis of the family Voltzialestidae is changed because of the inclusion of new genera. The genus Sushkinia Martynov, 1930 is transferred to Permagrionidae. The following permagrionids are described: Permagrion sharovi sp. nov, Permolestes obscurus sp. nov., P. vjatkensis sp. nov., Salagoulestes martynovi sp. nov., Solikamptilon aequus sp. nov., Sushkinia angulata sp. nov., Triadolestes sakmarensis gen. et sp. nov., and T. mutovkensis gen. et sp. nov. The new material from Isady and Kargala suggests closer relationships of the Permian Permagrionomorpha and Triassic Triadophlebiomorpha. In the family Kennedyidae, Kennedya sylvensis sp. nov. and K. volatica sp. nov. are described from Chekarda, and K. ivensis Nel et al., 2012 is redescribed from Soyana.