Biostratigraphy of Bajocian–Oxfordian strata based on dinoflagellate cysts and ammonites from the Dalichai Formation, northeastern Iran

The Bajocian–Oxfordian Dalichai Formation of the Binalud Mountains (northeastern Iran) consists of marls, marly limestones and limestones that were sampled and studied in detail for their palynomorph and ammonite contents. Palynological studies enabled differentiation of three marine biozones (Cribroperidinium crispum, Dichadogonyaulax sellwoodii, and Ctenidodinium continuum) within the lower interval of the formation. The palynofacies analysis shows a gradual sea-level rise during the depositional period of the rock unit, while terrestrial palynomorphs (spores and pollen grains) revealed a predominantly tropical climate for the time interval. Furthermore, a large number of ammonites were recovered from the formation, based on which 12 families, 22 genera, 30 species and 12 biozones (Zigzag, Aurigerus, Retrocostatum, Discus, Bullatus, Gracilis, Anceps, Coronatum, Athleta, Transversarium, Bifurcatus, and Bimammatum) were differentiated. When comparing the dinoflagellate cyst zones with the ammonite zones, the Dichadogonyaulax sellwoodii dinoflagellate cyst zone was correlated with the ammonite zones of Zigzag, Aurigerus, Retrocostatum, Discus, Bullatus, and Gracilis and the Ctenidodinium continuum dinocyst zone was correlated with the Gracilis and Anceps ammonite zones. The dinocyst and ammonite assemblages of the Binalud Mountains are similar to assemblages described from northern and northeastern Iran (Alborz and Koppeh-Dagh basins) as well as Northwest Europe. This suggests marine connections between these sedimentary basins during the Bajocian–Oxfordian. Consequently, the integrated dinocyst and ammonite zonation established here can be utilized also in other parts of the northern Tethyan Realm. The diachroneity of the upper boundary of the Dichadogonyaulax sellwoodii Zone between the Binalud Mountains and Northwest Europe is however revealed by comparison with the ammonite zones.