Death by ammonite: fatal ingestion of an ammonoid shell by an Early Jurassic bony fish

Abstract A remarkable specimen of the actinopterygian fish Pachycormus macropterus from the Early Jurassic (Toarcian) Posidonienschiefer Formation of Germany exceptionally preserves an unusually large ammonite inside its gut. The ammonite was swallowed by the fish, likely by accident, and represents the first direct evidence for an actinopterygian fish consuming an ammonoid. Exceptional aragonite preservation of the conch retaining partial nacreous lustre, combined with only minor acid etching of the shell, strongly indicates that the ammonite was ingested immediately prior to and was directly responsible for the fish’s death. The fish’s stomach provided a microenvironment protecting the aragonite from chemical dissolution.