American Beautyberry for Borrow Pit Reclamation in South Carolina

H. Martín, G. Sick
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Abstract

surface-mined sites. A beautyberry (Callicarpa americana) is a hardy, drought-tolerant shrub native to the southeastern United States from southern Maryland west to Tennessee and Oklahoma, south to Texas, Mexico, Florida, and the West Indies (Dirr, 1990; Foote and Jones, 1989; Radford et al., 1968). It is most common in the sandy lowlands of the southeastern Coastal Plain (Radford et al., 1968). In Florida and.adjacent states it is a "characterizing" shrub of the upland hardwood hammock plant association that is common over large areas of north and central Florida, southern Alabama and Georgia, and coastal areas of the Carolinas. ("Characterizing," according to the Soil Conservation Service, means that "this species so commonly occurs in a community that you would expect to see it there at most locations supporting that community" (Soil Conservation Service, 1989).) The upland hardwood hammock is a climax community with high species-diversity where few pines occur and hardwood trees dominate (Gano, 1917; Soil Conservation Service, 1989). Two associations at the Savannah River Site in South Carolina where our project was conducted are upland hardwood hammocks--the white oak-dogwood-pipsissewa and the white oak-post oak forest communities (Jones et al., 1981). Beautyberry also grows in the longleafpine/turkey-oak sand-hill association, in mixed hardwood and pine forests, in south Florida flatwoods, in cabbage palm flatwoods, and in wetland hardwood hammock plant communities in Florida and adjacent states (Soil Conservation Service, 1989). These diverse forest types provide a wide variety of feeding, nesting, and escape habitats for several animal species (Gilbert and Stout, 1983). In natural landscapes, beautyberry is usually found at the edges of woods (Brown & Tighe, 1991) and in forest openings resuiting from tree-fall and other natural disturbances. Beautyberry is also a common component of managed landscapeg such as hedgerows and roadsides (Martin et al., 1951). Beautyberry is in the Verbenaceae, the Verbena family, to which lantana (Lantana camara) and golden dewdrol5 (Durantarepens) also belong (Zona, 1994). American beautyberry is one of about 140 species of Callicarpa native to the tropics and mild-temperate zones of both the eastern and the western hemispheres (Zona, 1994). A number of observations and studies indicate that beautyberry has considerable value for wildlife. Beautyberry twigs and foliage are a minor component of the diet of white-tailed deer (Odocoileus virginianus) (Martin et al., 1951; Stribling, 1988). The fruit, a small, bright-violet, fleshy drupe (Radford et al., 1968), is a favored food of the robin (Turdus migratorius ) , mockingbird ( Mimus polyglottos ) , catbird (Dumatella carolinensis), brown thrasher ( Toxostoma rufum ) (Martin et al., 1951), and wild turkey (Meleagris gallo. pavo) (Kennamer et al., 1980), and is a mi-
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美国美丽的浆果在南卡罗来纳借坑开垦
surface-mined网站。美莓(calicarpa americana)是一种耐寒、耐旱的灌木,原产于美国东南部,从马里兰州南部到田纳西州和俄克拉何马州,南至德克萨斯州、墨西哥、佛罗里达州和西印度群岛(Dirr, 1990;富特和琼斯,1989;Radford et al., 1968)。它最常见于东南沿海平原的沙质低地(Radford et al., 1968)。在佛罗里达和。它是一种高地硬木吊床植物协会的“特征”灌木,在佛罗里达州北部和中部、阿拉巴马州南部和佐治亚州以及卡罗莱纳州沿海地区的大片地区都很常见。(根据土壤保持局的说法,“特征化”意味着“该物种在一个群落中如此普遍地出现,以至于你会期望在支持该群落的大多数地点看到它”(土壤保持局,1989)。)山地硬木吊床是一个物种多样性高的顶极群落,松树很少,硬木为主(Gano, 1917;土壤保持处,1989)。我们的项目是在南卡罗来纳州萨凡纳河遗址进行的,两个协会是高地硬木吊床——白橡树-茱萸-pipsissewa和白橡树-柱橡树森林社区(Jones et al., 1981)。美莓也生长在长叶松/火鸡橡树的沙丘丛中,在混合硬木和松林中,在南佛罗里达的平原林中,在卷心菜棕榈平原林中,在佛罗里达州和邻近州的湿地硬木吊床植物群落中(土壤保持局,1989年)。这些不同的森林类型为一些动物物种提供了各种各样的觅食、筑巢和逃生栖息地(Gilbert和Stout, 1983)。在自然景观中,美莓通常生长在树林的边缘(Brown & Tighe, 1991)以及由于树木倒下和其他自然干扰而形成的森林开口中。美丽的浆果也是一个常见的组成部分管理景观,如树篱和路边(马丁等人,1951)。美莓属马鞭草科,马鞭草科,马鞭草属(lantana camara)和金露珠(Durantarepens)也属于该科(Zona, 1994)。美洲美莓是大约140种原产于东半球和西半球热带和温和温带的美莓属植物之一(Zona, 1994)。许多观察和研究表明,美莓对野生动物具有相当大的价值。白尾鹿(Odocoileus virginianus)的饮食中,美莓树枝和叶子只占很小的一部分(Martin et al., 1951;Stribling, 1988)。这种果实是一种小的、亮紫色的肉质核果(Radford et al., 1968),是知更鸟(Turdus migratorius)、反舌鸟(Mimus polyglottos)、猫鸟(Dumatella carolinensis)、棕脱毛鸟(Toxostoma rufum) (Martin et al., 1951)和野生火鸡(Meleagris gallo)最喜欢的食物。pavo) (Kennamer et al., 1980),并且是一个mi-
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