Euboeus mimonti Boieldieu, 1865, the oldest record of an extant species of Tenebrionidae (Coleoptera) and notes on other species identified as darkling beetles from the Late Pliocene of Willershausen (Germany)

Abstract

Several fossil beetles from the Late Pliocene (Piacenzian) of Willershausen (Lower Saxony, Germany) previously identified as Tenebrionidae and Staphylinidae are reviewed. One species, formerly determined as “Staphylinoidea” is identified as Euboeus mimonti Boieldieu, 1865 (Tenebrionidae: Helopini). This species is the oldest record of an extant species of darkling beetles, which occurs now in the Balkans and Anatolia. The discovery of Euboeus mimonti in the Late Pliocene of Germany indicates that the range of this species was much wider, and the climate in the Late Pliocene in the modern territory of Lower Saxony was much milder. One species formerly determined as “Alleculidae gen. sp.” belongs to the subfamily Alleculinae (Tenebrionidae) and it is identified here as Pseudocistela aff. ceramboides. Two species, misidentified by a previous author as Tenebrionidae belong to Elateridae and Cerambycidae respectively. Generic and species composition, as well as the ratio of extinct and extant beetle taxa in the Willershausen Fossil Lagerstätte are discussed.