Endothyrella Zilch, 1960 in Bhutan (Gastropoda: Pulmonata: Plectopylidae), with a description of three new species

Abstract

Samples of the genus Endothyrella from Bhutan are dealt with, starting from the modern review of this genus by Páll-Gergely et al. (2015). Four species are recognized, 3 of which are new to science: E. bhutanensis, E. spirostriata, and E. pemagatshel. The record of Endothyrella blanda nearly 100 km southeast of its well documented range in Sikkim, India, puts some isolated unconfirmed records from India in a new light. Apart from shell size and shape, the number of spiral rows of hairs and the structure of the parietal and palatal callosities on the shell wall turned out to be important for identification. The species were found at altitudes of 335–2300 m a.s.l. in the Indo-Malayan part of Bhutan.