Silurian and earliest Devonian birkeniid anaspids from the Northern Hemisphere

H. Blom, T. Märss, C. Miller
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引用次数: 53

Abstract

ABSTRACT The sculpture of scales and plates of articulated anaspids from the order Birkeniida is described and used to clarify the position of scale taxa previously left in open nomenclature. The dermal skeleton of a well-preserved squamation of Birkenia elegans Traquair, 1898 from the Silurian of Scotland shows a characteristic finely tuberculated sculpture over the whole body. Rhyncholepis parvula Kiær, 1911, Pterygolepis nitida (Kiær, 1911) and Pharyngolepis oblonga Kiær, 1911, from the Silurian of Norway show three other sculpture types. Northern Hemisphere disarticulated scales and plates are described here, supporting a new anaspid taxonomy that includes both articulated and disarticulated material. The diversity, distribution, evolutionary trends and biostratigraphy of anaspids are described in the context of this new taxonomy, which includes six families (two are new) subdivided into 16 genera (10 are new) and 22 species (15 are new). New taxa among Birkeniidae Traquair, 1898 are Birkenia robusta sp. nov. and Hoburgilepis papillata gen. et sp. nov.. Rhyncholepididae Kiær, 1924 includes Rhyncholepis butriangula sp. nov., Silmalepis erinacea gen. et sp. nov., Vesikulepis funiforma gen. et sp. nov., Maurylepis lacrimans gen. et sp. nov., and the previously described Schidiosteus mustelensis Pander, 1856 and Rytidolepis quenstedtii Pander, 1856. Tahulalepididae fam. nov. is represented by Tahulalepis elongituberculata gen. et sp. nov. and the revised T. kingi (Woodward, 1947). Septentrioniidae fam. nov. contains Septentrionia lancifera gen. et sp. nov., S. mucronata gen. et sp. nov., S. dissimilis gen. et sp. nov., S. seducta gen. et sp. nov., Liivilepis curvata gen. et sp. nov., Spokoinolepis alternans gen. et sp. nov. and Manbrookia asperella gen. et sp. nov. The family level position of Ruhnulepis longicostata gen. et sp. nov. is uncertain. Pterygolepididae Obruchev, 1964 and Pharyngolepididae Kiær, 1924 remain monogeneric.
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北半球志留纪和最早的泥盆纪birkeniids
本文描述了Birkeniida目铰接无肢纲动物的鳞片和板片的雕刻,并用于澄清以前在开放命名法中留下的鳞片分类群的位置。摄于1898年苏格兰志留纪的一具保存完好的秀丽白kenia elegans Traquair鳞片的皮肤骨架,整个身体上都有一个典型的精细的结核状雕塑。来自挪威志留纪的Rhyncholepis parvula Kiær, 1911年,Pterygolepis nitida (Kiær, 1911年)和Pharyngolepis oblonga Kiær, 1911年显示了另外三种雕塑类型。北半球分离的鳞片和板在这里描述,支持一个新的无肢纲分类,包括关节和分离的材料。该分类包括6科(2新科),16属(10新科)和22种(15新科)。Birkenia robusta sp. nov.和Hoburgilepis papillata gen. et sp. nov.是1898年在Birkenia科发现的新分类群。kiekæ r, 1924,包括布三角舌蝗科,灰尾蝗科,长尾蝗科,长尾蝗科,长尾蝗科,长尾蝗科,长尾蝗科,长尾蝗科,长尾蝗科,长尾蝗科,长尾蝗科,长尾蝗科,长尾蝗科,长尾蝗科,长尾蝗科,长尾蝗科。Tahulalepididae家人。11 .以Tahulalepis elongituberculata gen. et sp. 11 .和经修订的T. kingi (Woodward, 1947)为代表。Septentrioniidae家人。11 .包含七叶蛾(Septentrionia lancifera gen. et nov)、长叶蛾(s.m.mucronata gen. et nov)、异叶蛾(s.m.dissimilis gen. et nov)、绢叶蛾(s.m.seducta gen. et nov)、曲叶蛾(liviilepis curvata gen. et nov)、花叶蛾(spokinoolepis alternans gen. et nov)和曲霉(Manbrookia asperella gen. et nov)。Pterygolepididae Obruchev, 1964和Pharyngolepididae Kiær, 1924仍然是单属的。
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