Biodiversity and evolutionary phases of Lochkovian (Lower Devonian) conodonts in the Pyrenees: A comparative study

This paper aims to evaluate the zone-by-zone changes in Lochkovian conodont biodiversity in the Pyrenees region and compare the results with previous studies. Additionally, the paper seeks to identify evolutionary phases within the Lochkovian. We analyse the stratigraphic range data from eight classical Pyrenean Lochkovian sections. A total of 48 taxa at the species level, primarily confined to the Lochkovian, are included in the analysis. We employ both qualitative and quantitative approaches, including Clark's method for the conodont evolution index and Foote's parameters for measuring taxonomic diversity and rates. The zonal scheme employed in this study follows the global three-fold subdivision of the Lochkovian stage, with additional subdivisions based on local records. The zone-by-zone analysis of stratigraphically well-controlled successions contributes to our understanding of the conodont diversity dynamics in this region. The results are consistent with previous studies, but also highlight the unique aspects of the Pyrenean conodont record. Their highest diversity occurs in the middle Lochkovian within the kutscheri-pandora beta Zone, succeeded by an extinction event towards the end of this interval. Following a brief proliferation of species at the onset of the upper Lochkovian, another extinction event concludes this Stage. Results from all different rate measurements applied to Pyrenean Lochkovian conodonts are in disagreement with the prevailing perspective suggesting an escalation in the proportion of singletons and a concomitant decline in per-taxon rates with an increase in interval length. Comparative analysis of conodont records across various regions (Carnic Alps, Prague Synform, Central Nevada, and the Pyrenees) during the Lochkovian reveals significant differences in biodiversity, taxonomic composition, and zonal distribution.