[土壤生态系统进化的早期步骤]。

Pub Date : 2013-11-01
A G Ponomarenko
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引用次数: 0

摘要

土壤化石早在前寒武纪就已为人所知,远早于地球上高等陆生植物的出现。陆地和大陆水体的原始生物群落是漂浮的和底部的原核生物垫层和薄膜,它们产生了大部分生物量,在比生产力方面不低于任何其他光合作用。在高等植物出现之前,侵蚀非常强烈,导致地势平坦,没有永久性溪流,主要是流河和地表径流;所有的水体都是泥泞的。当洪水发生时(这是经常发生的),泥水流的粘土颗粒使底垫与阳光隔绝,然后它们的大部分就消失了。结果土壤不是一个统一的生物惰性体,而是一个由未氧化的烧焦有机物和粘土层组成的“泡芙饼”。未氧化有机物的埋藏使大气中的氧气富集。蚯蚓和节肢动物在寒武纪时期出现在陆地和大陆水体中,它们将有机质和矿物成分混合在一起,大大加强了土壤的形成过程。现代类型的土壤是在泥盆纪高等植物扩张并取代浅水底垫后出现的。这一时期存在的土壤动物在进化水平上与现代土壤动物没有太大的不同。
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[Earlier steps of the soil ecosystem evolution].

Fossil soils are known since early Praecambrian, long before the occurrence of higher terrestrial plants on the Earth. Primeval biocoenoses on the land and in continental water bodies were floating and bottom prokaryotic mats and films which produced the majority of biomass and with regard to specific productivity were not inferior to any other photosynthetics. Before the occurrence of higher plants, erosion was very strong, resulting in flat relief, absence of permanent streams, domination of wandering rivers and surface runoff; all water bodies were muddy. When floods occurred, which was quite so often, clay particles of muddy water streams isolated bottom-mats from the light and then their considerable part perished. The result was not soil as a uniform bioinert body but rather a "puff pie" consisted of layers of unoxidized charred organic matter and clay prolayers. The burial of unoxidized organic matter contributed to enrichment of the atmosphere with oxygen. Worms and arthropods, which came out to the land and continental water bodies during Cambrian period, mixed up the organic matter with mineral components strengthening the process of soil forming considerably. Soils of the modern type appeared after higher plants expanded in Devonian and displaced bottom-mats in shallow waters. The soil fauna that existed at this time was not so different from the modern one with regard to its evolutionary level.

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