Sorellevania Engel的第二个物种,2006(Evanioidea:Evaniidae),来自白垩纪中期的缅甸琥珀

IF 1.9 Q2 ENTOMOLOGY Palaeoentomology Pub Date : 2023-02-28 DOI:10.11646/palaeoentomology.6.1.2
C. Jouault
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引用次数: 2

摘要

Evanioidea超科的代表很容易与所有其他现存和已灭绝的膜翅目区分开来,因为它们的交代体(有时是圆形的)高高地铰接在蜂胶上,远高于元壳纲(例如,Goulet和Huber,1993)。最近对Evanioidea超科的系统发育研究表明,即使包括化石分类群,也强烈支持其单系性和Evaniidae的单系性(例如,Li等人,2018;Parslow等人,2020;Jouault等人,2022)。根据最新的分析,该超家族出现在上三叠纪或下侏罗纪。尽管如此,其最早的物种记录在更年轻的中侏罗纪,其冠群代表在下白垩纪(Jouault et al.,2022:图8)。据估计,茎Evaniidae起源于上侏罗纪,而它们的冠群起源更近,可能在白垩纪-古近纪边界附近。现存的伊凡尼黄蜂很常见,几乎是世界性的,并且适度多样化(21属约580种现存物种),即使这种多样性被低估了(Deans,2005;Mullins等人,2012年)。对现存的伊凡虫的生物学知之甚少,但它们的幼虫被认为是卵鞘科蟑螂卵的捕食者(Huben,1995)。Evaniidae有着良好的化石记录,其已知的最古老的物种可以追溯到豪特里阶晚期,记录到中新世的许多物种,以及缅甸琥珀中的重要多样性(例如,Rasnitsyn等人,1998年;Nel等人,2002年;Deans等人,2004年;Jennings等人,2012年;Shih等人,2020年)。尽管如此,他们过去的多样性仍然被低估。
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The second species of Sorellevania Engel, 2006 (Evanioidea: Evaniidae) from the mid-Cretaceous Burmese amber
The representatives of the superfamily Evanioidea are easily distinguishable from all other extant and extinct Hymenoptera because of their metasoma (sometimes rounded) articulated high on the propodeum and well above the metacoxae (e.g., Goulet & Huber, 1993). Recent phylogenetic studies on the superfamily Evanioidea have shown strong support for its monophyly and for the monophyly of the Evaniidae even when fossil taxa are included (e.g., Li et al., 2018; Parslow et al., 2020; Jouault et al., 2022). According to the most recent analysis, the superfamily arose during either the Upper Triassic or the Lower Jurassic. Still, its earliest species are recorded in the younger Middle Jurassic, and its crown-group representatives during the Lower Cretaceous (Jouault et al., 2022: fig. 8). The stem Evaniidae are estimated to arise during the Upper Jurassic while their crown group has a more recent origin, likely around the Cretaceous-Paleogene boundary. Extant evaniid wasps are common, nearly cosmopolitan, and moderately diversified (about 580 extant species in 21 genera) even if this diversity is underestimated (Deans, 2005; Mullins et al., 2012). Little is known about the biology of extant evaniids, but their larvae are considered predators of cockroach eggs in oothecae (Huben, 1995). The Evaniidae have a good fossil record with the oldest species known from the late Hauterivian, numerous species documented up to the Miocene, and an important diversity in Burmese amber (e.g., Rasnitsyn et al., 1998; Nel et al., 2002; Deans et al., 2004; Jennings et al., 2012; Shih et al., 2020). Nevertheless, their past diversity is still underestimated.
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