Evidence for the Dual Role of Floral Secretory Cells in Bulbophyllum

Plants of the genus Bulbophyllum Thouars have some of the most intricate flowers and complex pollination mechanisms to be found amongst the Orchidaceae (van der Pijl and Dodson, 1969; van der Cingel, 2001). For an up-to-date, overall account of the genus, readers are referred to Pridgeon et al. (2014). Many of its members are pollinated by dipterans (van der Pijl and Dodson, 1969; van der Cingel, 2001) attracted by foodrewards and fragrance. However, to date, these food-rewards have been investigated for relatively few species, mostly Neotropical and African taxa, and have generally been found to be lipid-rich and produced by palisade-like epidermal cells of the