Bryozoan assemblages of the Gulpen Formation (upper Campanian – upper Maastrichtian) in the Liège-Limburg area (Belgium, the Netherlands)

Oliver Kesselhut, Luc Goffings, Johan Vellekoop
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Abstract

Renewed interest in bryozoan assemblages from the various members of the Gulpen Formation (upper Campanian–upper Maastrichtian) in the Liège-Limburg area (southeast Netherlands and northeast Belgium) during recent years has resulted in the discovery of a number of species previously unrecorded from the area. Nineteen species are here recorded from the Zeven Wegen, Vijlen, Lixhe and Lanaye members (Gulpen Formation) in the study area, occurring in three distinct assemblages. The vast majority of the bryozoan taxa belong to the order Cheilostomata; only two taxa are assigned to the order Cyclostomata. The species recorded here are Clinopora aff. costulata, Disporella obvallata, ‘Vincularia’ (sensu lato) marssoniana, Herpetopora laxata, Heteroconopeum ovatum, Wilbertopora aff. oxyteichos, Biaviculigera sacerdotalis, Semiflustrella britannica, Aechmellina anglica, Escharifora papyracea, Onychocella cyclostoma, Onychocella cylindrica, Onychocella matrona, Rhebasia disparilis, Stichomicropora sicksi, Coscinopleura elegans, Coscinopleura lamourouxi, Pachydermopora pachyderma and Beisselina aviculifera. While not previously recorded from the Liège-Limburg area, the bryozoan taxa identified from the Gulpen Formation are well known from coeval Cretaceous strata elsewhere in Europe, North America and Russia, highlighting their broad palaeobiogeographical distribution.

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列日-林堡地区(比利时、荷兰)古尔彭地层(上元古界-上元古界)的浮游生物组合
近年来,人们对列日-林堡地区(荷兰东南部和比利时东北部)古尔彭地层(上元古界-上元古界)各岩层中的贝类组合重新产生了兴趣,并发现了许多以前从未在该地区记录到的物种。本文记录了来自研究地区 Zeven Wegen、Vijlen、Lixhe 和 Lanaye 组(Gulpen Formation)的 19 个物种,它们出现在三个不同的组合中。绝大多数的类囊动物都属于Cheilostomata目,只有两个类群被归入Cyclostomata目。这里记录的物种有 Clinopora aff. costulata、Disporella obvallata、'Vincularia' (sensu lato) marssoniana、Herpetopora laxata、Heteroconopeum ovatum、Wilbertopora aff.oxyteichos、Biaviculigera sacerdotalis、Semiflustrella britannica、Aechmellina anglica、Escharifora papyracea、Onychocella cyclostoma、Onychocella cylindrica、Onychocella matrona、Rhebasia disparilis、Stichomicropora sicksi、Coscinopleura elegans、Coscinopleura lamourouxi、Pachydermopora pachyderma 和 Beisselina aviculifera。虽然以前在列日-林堡地区没有发现过这些类群,但在欧洲、北美和俄罗斯其他地区的共生白垩纪地层中,古尔班地层中发现的这些类群已广为人知,这突出表明了它们在古生物地理学上的广泛分布。
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