About the Age of Fossil Flora of the Kygyltasian Regional Stage of the Upper Paleozoic in the Verkhoyansk Region, Northeastern Russia

L.G. Porokhovnichenko, R. Kutygin, A. Kilyasov
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Abstract

The Kygyltasian Regional Stage is widespread throughout the Verkhoyansk Region, where it reflects a large transgressive-regressive cycle of the Verkhoyanian terrigenous sedimentation. Over the years, a huge amount of material has been accumulated on biostratigraphic division, facies features, and paleontological characteristics of this large stratigraphic unit, but its age is still debatable. The main fossils in the upper part of the Kygyltasian Regional Stage are higher plants, the remains of which were studied in the last century using outdated taxonomy. This study is devoted to revision of the collection of fossil plants from the type locality of the Kygyltasian Regional Stage (Western Verkhoyanie) in order to clarify the taxonomic composition and the age of the Upper Kygyltasian floristic association. The dominant cordaite leaves in the association were studied using the morphological and microstructural methods. This made it possible to clarify the previous paleontological definitions of the genus Noeggerathiopsis and bring them in line with modern taxonomy. The species Noeggerathiopsis heteroclitus Tolstych and N. khorokytica Tolstych were validated for type specimens. The following species were identified from the Upper Kygyltas Subformation of the Khorokyt section: Cordaites heteroclitus (Tolstych), C. cf. neuburgae Gluchova, C. cf. zalesskyi Durante, C. krychtofovichii (Radczenko), Cordaites ? khorokytica (Tolstych), Cordaites sp. sp. sp. sp. Rufloria ( Praerufloria ) Sphenophyllum cf. Sphenophyllum ? sp., Sphenopteris sp., Paracalamites sp. Based on the results of the study, the Upper Kygyltasian floristic association of the Western Verkhoyansk Region was compared with floristic associations of the Alykaeva Formation (Kuzbass) and the Upper Kata Subformation (Tunguska basin). From the above, the Late Carboniferous age of the upper part of the Kygyltasian Regional Stage follows. The results obtained are in good agreement with the correlation of the Verkhoyanian Carboniferous-Permian deposits for marine invertebrates.
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俄罗斯东北部维尔霍扬斯克地区上古生界Kygyltasian区域阶段化石植物群的时代
吉尔吉斯区域阶段广泛分布于维尔霍扬斯克地区,反映了韦尔霍扬期陆源沉积的一次大型海侵-退回旋回。多年来,人们对这一大型地层单元的生物地层划分、相特征、古生物学特征等方面积累了大量资料,但其年代仍有争议。吉尔吉斯区域阶段上部的主要化石为高等植物,这些植物的遗存是在上个世纪使用过时的分类方法进行研究的。本文对Kygyltasian Regional Stage (Western Verkhoyanie)模式地区的植物化石标本进行了修订,以明确Kygyltasian Upper flora association的分类组成和时代。用形态学和显微结构的方法研究了该组合中的优势叶片。这使得澄清以前对Noeggerathiopsis属的古生物定义成为可能,并使它们与现代分类学保持一致。以Noeggerathiopsis heteroclitus Tolstych和N. khorokytica Tolstych为模式标本进行了验证。在Khorokyt剖面上Kygyltas亚组中鉴定到的种有:Cordaites heteroclitus (Tolstych), C. cfneuburgae Gluchova, C. cfzalesskyi Durante, C. krychtofovichii (Radczenko), Cordaites ?花椰菜(Tolstych), Cordaites sp. sp. Rufloria (Praerufloria)在此基础上,将西部Verkhoyansk地区上Kygyltasian的区系组合与库兹巴斯地区Alykaeva组和通古斯盆地上Kata亚组的区系组合进行了比较。由此推断出晚石炭世为吉尔吉斯区域阶段上半部分。所得结果与Verkhoyanian石炭-二叠纪海洋无脊椎动物沉积对比结果吻合较好。
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