Seed Hardening: A Way to Tolerate Against Abiotic Stress in Rainfed Areas

L. Jain, M. Verma, N. Ram, A. Choudhary, H. P. Parewa
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Abstract

Pre-sowing hardening or priming of seeds is one of the best methods for altering the biochemical and physiological process of seeds for favourable condition to induce drought tolerance in emerging seedlings. The recurrent phenomenon in rainfed areas i.e. soil moisture stress immediately after sowing affects seeds germination and their establishment. Under sub-optimal environmental conditions like rainfed farming, poor seed germination followed by poor field establishment is a common phenomenon. Seed hardening enables seedlings to survive this early moisture stress. hydration of seeds for short term before planting greatly benefits stand establishment along with the uses of chemicals like potassium or sodium phosphate would give additional advantage. It is defined as the process of soaking seeds in chemical solution for specified time and then drying to induce tolerance to drought. Various seed priming/hardening techniques have been developed including hydro-priming, halo-priming, osmo-priming and hormonal priming. Micronutrient seed treatments, which include seed priming and seed coating, are an attractive and easy alternative. Seed hardening promotes early establishment provides more strengthen to plant and it became competitive against infections and diseases is related to gene regulation and expression under stress conditions. The discovery of plant hormones in the 1920s underlined the crucial role of these compounds in seed desiccation tolerance, reserve mobilization, as well as cell division and cell elongation occurring during germination. The seed hardening is considered as low cost technology and is the most important requirement for pre-monsoon sowing. Seed hardening offer an attractive option for resource poor farmers. During seed hardening, seeds are subjected to partial hydration followed by dehydration before sowing.
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种子硬化:一种在雨养地区抵御非生物胁迫的方法
播前硬化或催熟是改变种子生化和生理过程的最佳方法之一,为苗期抗旱创造有利条件。在雨养地区,播种后土壤水分胁迫会影响种子的萌发和形成。在雨养耕作等次优环境条件下,种子发芽率低,接着是田间条件差是一种普遍现象。种子硬化使幼苗能够在早期的水分胁迫下存活下来。在播种前对种子进行短期的水化处理,对种植受益匪浅,同时使用钾肥或磷酸钠等化学品也会带来额外的好处。它是将种子在化学溶液中浸泡一定时间,然后干燥以诱导其抗旱能力的过程。各种种子启动/硬化技术已经发展起来,包括氢启动、晕启动、渗透启动和激素启动。微量营养素种子处理,包括种子灌浆和种子包衣,是一种有吸引力和简单的替代方案。种子硬化促进了种子的早期形成,为植物提供了更强的力量,并在逆境条件下具有对抗感染和疾病的竞争力,与基因调控和表达有关。20世纪20年代植物激素的发现强调了这些化合物在种子萌发过程中对干燥耐受性、储备动员以及细胞分裂和细胞伸长的重要作用。种子硬化被认为是低成本的技术,是季前播种的最重要要求。种子硬化为资源贫乏的农民提供了一个有吸引力的选择。在种子硬化期间,种子在播种前遭受部分水化,随后脱水。
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