Modeling natural mortality for different plant densities in dendroenergetic trials

Simón Sandoval, Eduardo Acuña, J. Cancino, R. Rubilar
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引用次数: 1

Abstract

Mortality was modelled for three species (Acacia melanoxylon, Eucalyptus camaldulensis, Eucalyptus nitens) at three plantation densities (5000, 7500, and 10000 trees ha-1) in an trial of biomass production for purposes of dendroenergetic. One modelling based on individual tree level and two mortality modelling alternatives were evaluated: four survival probability equations and eight difference equations. The individual tree survival modelling considered a logistic model, is a linear combination of variables to individual tree at current time  and the previous time as estimator, being the main variables the variation of the competition index and the variation of basal area growth between the current growth period and the previous growth period. The survival probability alternative used state variables of the stand (age, dominant height, average square diameter) as predictors, whereas the difference equations were adjusted according to age-based changes only. The models to stand levels showed better result than individual tree models, and in general, the mortality models based on difference equations presented better indicators of precision and parsimony. The rate of relative mortality was constant, i.e., (dN/dE)/N, and varied between species, revealing greater mortality, consecutively, in E. nitens, A. melanoxylon, and E. camaldulensis. Although mortality tended to be higher at greater plantation densities, stand density did not significantly affect the parameters of the adjusted models. Highlights The mortality stand level models showed better results than the individual tree models for dendroenergetic crops, and in general, the mortality models based on difference equations presented better precision indicators and parsimony. The survival probability alternative involved state variables of the stand like age, dominant height, and average square diameter as predictors, while the difference equations were fitted according to age-based changes only. Mortality tended to be higher at greater plantation densities, however stand density did not significantly affect the parameters of the mortality equations.
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在树能试验中模拟不同植物密度的自然死亡率
在一项以树木能量为目的的生物量生产试验中,对三种树种(黑梭槐、桉树、nitens)在三种人工林密度(5000、7500和10000棵/公顷)下的死亡率进行了建模。评估了一种基于个体树水平的模型和两种死亡率模型替代方案:4个生存概率方程和8个差分方程。单株树生存模型被认为是一种logistic模型,是以单株树当前时间和前一时间为估计量的变量的线性组合,以竞争指数的变化和当前生长期与前一生长期的基面积生长的变化为主要变量。生存概率替代方案使用林龄、优势高度、平均平方直径等状态变量作为预测因子,而差异方程仅根据林龄变化进行调整。基于林分水平的模型优于单株模型,总体而言,基于差分方程的死亡率模型具有较好的精度和简洁性。相对死亡率(dN/dE)/N为恒定值,且不同种间存在差异,依次为nitens、melanoxylon和camaldulensis。人工林密度越大,死亡率越高,但林分密度对模型参数的影响不显著。结论树木能作物的林分水平死亡率模型优于单树模型,总体而言,基于差分方程的死亡率模型具有更好的精度指标和简洁性。生存概率替代方案以林龄、优势高度和平均平方直径等状态变量作为预测因子,而差异方程仅根据林龄变化拟合。人工林密度越大,死亡率越高,但林分密度对死亡率方程参数影响不显著。
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