Early Cretaceous dinosaur, bird and turtle tracks from the Lanzhou-Minhe Basin, Gansu Province, Northwest China

The Early Cretaceous avian ichnofauna of Laurasia, particularly in East Asia, is remarkably abundant. The northwestern region of China is the most productive area for bird tracks. Recently, four avian-dominated track sites have been discovered in the Cretaceous Lanzhou-Minhe Basin of Gansu, where the shorebird track Koreanaornis, the ankylopollexian track Caririchinium and the turtle track Chelonipus occur. The Kongjiasi site yields a new type of the fluvio-lacustrine Chelonipus ichnocoenosis related to birds, which was previously defined to include only non-avian theropods and turtles. The site is associated with a waterfront foraging site of a Cretaceous shorebird which might be due to the invertebrate-rich substrate. The sole association of both bird and other theropod tracks with Chelonipus ichnocoenosis may indicate a difference in the appetite of avian and non-avian theropods for littoral foraging sites. And a review of the global turtle track-related ichnofauna and ichnocoenosis may offer new insights into the qualitative speculation of palaeobathymetry in riparian environments.