Pub Date : 2026-01-14DOI: 10.1016/j.cretres.2026.106315
Maria Tekleva , Mário Miguel Mendes , Jiří Kvaček , Johanna H.A. Van Konijnenburg-van Cittert , Pedro Miguel Callapez , Zuzana Heřmanová
A new cheirolepidiaceous conifer pollen cone, Classostrobus amealensis sp. nov., is described from the Lower Cretaceous of Lusitanian Basin, in the Estremadura region of western Portugal. The new microsporangiate cone was collected in the Vale Cortiço open cast clay pit complex, near the small village of Ameal, from sedimentary deposits belonging to Santa Susana Formation and considered to be early Hauterivian in age. The new species is based on a single well-preserved, coalified microsporangiate cone, ca. 27 mm long and 14 mm wide, ovoid in shape, bearing ca. 20 helically arranged, imbricate microsporophylls. The microsporophyll bears four pollen sacs abaxially and it is smooth or finely papillate showing an acute apex fringed with short trichomes. The new microsporangiate cone is morphologically different from all Classostrobus species previously described, and we compare it with all other frenelopsid microsporangiate cones known to date. Pollen grains in situ show close similarity to dispersed pollen of the species Classopollis martinottii Reyre, 1970. Pollen details were studied using LM as well as scanning electron (SEM) and transmission electron microscopy (TEM). Pollen grains are spheroidal, ca. 26.1 μm in LM and 23.8 μm in SEM, characterized by an equatorial band with six to eight striations, a distal cryptopore, and a proximal tetrad scar. The exine sculpture is distinctly microechinate. The new microsporangiate cone is associated with fragments of vegetative shoots of Frenelopsis teixeirae Alvin et Pais 1978 and Pseudofrenelopsis dinisii M.M.Mendes et J.Kvaček et al., 2023.
在葡萄牙西部埃斯特雷马杜拉地区的卢西塔尼亚盆地下白垩世发现了一种新的针叶科针叶树花粉球果Classostrobus amealensis sp. nov.。新的微孢子体圆锥体是在靠近Ameal小村庄的Vale cortirado露天粘土坑群中收集到的,来自于属于Santa Susana组的沉积矿床,被认为是早Hauterivian时代。新种是基于一个保存完好的,联合的小孢子囊球果,长约27毫米,宽约14毫米,卵球形,长约20个螺旋排列的,叠瓦状的小孢子叶。小孢子叶背面有4个花粉囊,光滑或细乳头状,尖尖具短毛。新发现的小孢子囊球果在形态上不同于以前所描述的所有类球,我们将其与迄今为止已知的所有系蛙类小孢子囊球果进行了比较。原位花粉粒与Classopollis martinottii Reyre, 1970的分散花粉相似。利用LM、扫描电子显微镜(SEM)和透射电子显微镜(TEM)研究了花粉的细节。花粉粒呈球形,LM约26.1 μm, SEM约23.8 μm,具有6 ~ 8条条纹的赤道带,远端为隐孔,近端为四分体疤痕。外面的雕塑明显是微尖形的。新的小孢子球果与Frenelopsis teixeirae Alvin et Pais 1978和Pseudofrenelopsis dinisii m.m.d endes et j.k vaek et al., 2023的营养芽片段有关。
{"title":"The microsporangiate cone Classostrobus amealensis sp. nov. with in situ pollen from the Lower Cretaceous (lower Hauterivian) of Portugal: pollen ultrastructure and implications for frenelopsid species diversity","authors":"Maria Tekleva , Mário Miguel Mendes , Jiří Kvaček , Johanna H.A. Van Konijnenburg-van Cittert , Pedro Miguel Callapez , Zuzana Heřmanová","doi":"10.1016/j.cretres.2026.106315","DOIUrl":"10.1016/j.cretres.2026.106315","url":null,"abstract":"<div><div>A new cheirolepidiaceous conifer pollen cone, <em>Classostrobus amealensis</em> sp. nov., is described from the Lower Cretaceous of Lusitanian Basin, in the Estremadura region of western Portugal. The new microsporangiate cone was collected in the Vale Cortiço open cast clay pit complex, near the small village of Ameal, from sedimentary deposits belonging to Santa Susana Formation and considered to be early Hauterivian in age. The new species is based on a single well-preserved, coalified microsporangiate cone, ca. 27 mm long and 14 mm wide, ovoid in shape, bearing ca. 20 helically arranged, imbricate microsporophylls. The microsporophyll bears four pollen sacs abaxially and it is smooth or finely papillate showing an acute apex fringed with short trichomes. The new microsporangiate cone is morphologically different from all <em>Classostrobus</em> species previously described, and we compare it with all other frenelopsid microsporangiate cones known to date. Pollen grains <em>in situ</em> show close similarity to dispersed pollen of the species <em>Classopollis martinottii</em> Reyre, 1970. Pollen details were studied using LM as well as scanning electron (SEM) and transmission electron microscopy (TEM). Pollen grains are spheroidal, ca. 26.1 μm in LM and 23.8 μm in SEM, characterized by an equatorial band with six to eight striations, a distal cryptopore, and a proximal tetrad scar. The exine sculpture is distinctly microechinate. The new microsporangiate cone is associated with fragments of vegetative shoots of <em>Frenelopsis teixeirae</em> Alvin et Pais 1978 and <em>Pseudofrenelopsis dinisii</em> M.M.Mendes et J.Kvaček et al., 2023.</div></div>","PeriodicalId":55207,"journal":{"name":"Cretaceous Research","volume":"181 ","pages":"Article 106315"},"PeriodicalIF":1.7,"publicationDate":"2026-01-14","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"146037899","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
Pub Date : 2026-01-13DOI: 10.1016/j.cretres.2026.106313
Joel Crothers , Jaelyn Eberle , Donald Brinkman , Alyssa Wurtz , Andrew B. Heckert , ReBecca K. Hunt-Foster , John R. Foster , Ida C. Dirkes , Renee Dunn
The Williams Fork Formation (WFF) of northwestern Colorado preserves an understudied freshwater biota from the Campanian/Maastrichtian boundary. Here we describe a diverse actinopterygian-dominated fish assemblage from the ReBecca's Hollow locality of Rio Blanco County. Chondrichthyans are rare from this site, but include a hybodontid, Lonchidion, and the hemiscyllid Chiloscyllium. A fragmentary element may represent a pycnodontiform. Chondrosteans are represented by a tentative acipenserid. Holosteans include the lepisosteids Atractosteus and an unnamed taxon, and the amiids Melvius, Cyclurus and Palaeolabrus. Teleosteomorphs at this site include Belonostomus, Paralbula casei, Coriops, Estesesox foxi, Acronichthys, hiodontids, and acanthomorphs. This locality yields at least 17 fish taxa, most of which have not been described previously from the WFF. Although the WFF is temporally correlative with the St. Mary River and Horseshoe Canyon Formations in Alberta and the Prince Creek Formation in Alaska, the ReBecca's Hollow fish assemblage is markedly different from its northern contemporaries. Specifically, it contains several warm-climate taxa such as Lonchidion, Chiloscyllium, Melvius, Atractosteus, and Paralbula, and lacks higher latitude taxa such as Holostean A, documented from Alberta. The differences in these broadly contemporaneous fish assemblages supports the hypothesis that there was provincialism amongst freshwater fishes in Laramidia at the Campanian/Maastrichtian boundary. The fish fauna thus resembles that of Campanian localities in Utah, New Mexico, and Texas, and Maastrichtian (Lancian) localities in Montana, North Dakota, South Dakota, and Wyoming, suggesting that the boundary between these provincial zones remained north of the WFF locality throughout these periods of climate change.
{"title":"An actinopterygian-dominated fish fauna from the Upper Cretaceous Williams Fork Formation, northwestern Colorado, and evidence for provinciality across Laramidia at the Campanian/Maastrichtian boundary","authors":"Joel Crothers , Jaelyn Eberle , Donald Brinkman , Alyssa Wurtz , Andrew B. Heckert , ReBecca K. Hunt-Foster , John R. Foster , Ida C. Dirkes , Renee Dunn","doi":"10.1016/j.cretres.2026.106313","DOIUrl":"10.1016/j.cretres.2026.106313","url":null,"abstract":"<div><div>The Williams Fork Formation (WFF) of northwestern Colorado preserves an understudied freshwater biota from the Campanian/Maastrichtian boundary. Here we describe a diverse actinopterygian-dominated fish assemblage from the ReBecca's Hollow locality of Rio Blanco County. Chondrichthyans are rare from this site, but include a hybodontid, <em>Lonchidion</em>, and the hemiscyllid <em>Chiloscyllium</em>. A fragmentary element may represent a pycnodontiform. Chondrosteans are represented by a tentative acipenserid. Holosteans include the lepisosteids <em>Atractosteus</em> and an unnamed taxon, and the amiids <em>Melvius</em>, <em>Cyclurus</em> and <em>Palaeolabrus</em>. Teleosteomorphs at this site include <em>Belonostomus</em>, <em>Paralbula casei</em>, <em>Coriops</em>, <em>Estesesox foxi</em>, <em>Acronichthys</em>, hiodontids, and acanthomorphs. This locality yields at least 17 fish taxa, most of which have not been described previously from the WFF. Although the WFF is temporally correlative with the St. Mary River and Horseshoe Canyon Formations in Alberta and the Prince Creek Formation in Alaska, the ReBecca's Hollow fish assemblage is markedly different from its northern contemporaries. Specifically, it contains several warm-climate taxa such as <em>Lonchidion</em>, <em>Chiloscyllium</em>, <em>Melvius</em>, <em>Atractosteus</em>, and <em>Paralbula</em>, and lacks higher latitude taxa such as Holostean A, documented from Alberta. The differences in these broadly contemporaneous fish assemblages supports the hypothesis that there was provincialism amongst freshwater fishes in Laramidia at the Campanian/Maastrichtian boundary. The fish fauna thus resembles that of Campanian localities in Utah, New Mexico, and Texas, and Maastrichtian (Lancian) localities in Montana, North Dakota, South Dakota, and Wyoming, suggesting that the boundary between these provincial zones remained north of the WFF locality throughout these periods of climate change.</div></div>","PeriodicalId":55207,"journal":{"name":"Cretaceous Research","volume":"182 ","pages":"Article 106313"},"PeriodicalIF":1.7,"publicationDate":"2026-01-13","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"146071113","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
Pub Date : 2026-01-10DOI: 10.1016/j.cretres.2026.106312
Daniel Sedorko
The Lower Cretaceous deposits of the Sanfranciscana Basin (Brazil) were predominantly shaped by continental processes. Despite this, thin chert layers bearing marine microfossils indicate brief marine incursions. Various hypotheses have been proposed to explain these fossils, ranging from storm-induced remobilization to episodic incursions controlled by faulting. The short-lived nature of these events has been suggested by the absence of sedimentological and ichnological evidence of sustained marine influence in the basin's interior. To explore whether these incursions left additional evidence beyond microfossil assemblages, this study integrates ichnological and sedimentological analyses in the southern Sanfranciscana Basin. Fieldwork focused on the transitional interval between the Quiricó and Três Barras formations. Notably, three sections within the basal Três Barras Formation preserve a marine ichnoassemblage, consisting of whitish, fine-grained stratified sandstones, with horizontally concentric-filled (Asterosoma), plug-shaped (Bergaueria), and spreiten-bearing (Teichichnus) burrows. The low-diversity ichnoassemblage suggests impacted salinity conditions, corresponding to a polyhaline-brackish setting. The studied successions show dominance of low angle stratified sandstone interbedded with trough cross-bedded sandstones and wave ripples, suggesting high energetic settings, locally reworked by waves. However, restricted exposition area and thickness, as well as low diverse marine signatures allow to identify a marine connection to the sea, precluding definition of an architectural element or depositional scenario. This record suggests that the marine incursions were long enough to allow benthic colonization of the substrate, as evidenced by at least 12 bioturbated beds. This data argues against episodic, short-lived colonization driven by storm events, at least for the here studied sections.
{"title":"Marine incursion in the lower Cretaceous Sanfranciscana Basin evidenced by ichnological data","authors":"Daniel Sedorko","doi":"10.1016/j.cretres.2026.106312","DOIUrl":"10.1016/j.cretres.2026.106312","url":null,"abstract":"<div><div>The Lower Cretaceous deposits of the Sanfranciscana Basin (Brazil) were predominantly shaped by continental processes. Despite this, thin chert layers bearing marine microfossils indicate brief marine incursions. Various hypotheses have been proposed to explain these fossils, ranging from storm-induced remobilization to episodic incursions controlled by faulting. The short-lived nature of these events has been suggested by the absence of sedimentological and ichnological evidence of sustained marine influence in the basin's interior. To explore whether these incursions left additional evidence beyond microfossil assemblages, this study integrates ichnological and sedimentological analyses in the southern Sanfranciscana Basin. Fieldwork focused on the transitional interval between the Quiricó and Três Barras formations. Notably, three sections within the basal Três Barras Formation preserve a marine ichnoassemblage, consisting of whitish, fine-grained stratified sandstones, with horizontally concentric-filled (<em>Asterosoma</em>), plug-shaped (<em>Bergaueria</em>), and spreiten-bearing (<em>Teichichnus</em>) burrows. The low-diversity ichnoassemblage suggests impacted salinity conditions, corresponding to a polyhaline-brackish setting. The studied successions show dominance of low angle stratified sandstone interbedded with trough cross-bedded sandstones and wave ripples, suggesting high energetic settings, locally reworked by waves. However, restricted exposition area and thickness, as well as low diverse marine signatures allow to identify a marine connection to the sea, precluding definition of an architectural element or depositional scenario. This record suggests that the marine incursions were long enough to allow benthic colonization of the substrate, as evidenced by at least 12 bioturbated beds. This data argues against episodic, short-lived colonization driven by storm events, at least for the here studied sections.</div></div>","PeriodicalId":55207,"journal":{"name":"Cretaceous Research","volume":"181 ","pages":"Article 106312"},"PeriodicalIF":1.7,"publicationDate":"2026-01-10","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"146037898","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
Pub Date : 2026-01-07DOI: 10.1016/j.cretres.2025.106311
Zsolt Vallner , Tamás Müller , Dávid Bajnai , Ottilia Szives , Lajos Ádám Csicsek , Gábor Héja , Bernát Heszler , Norbert Zajzon , József Pálfy
The OAE 1d is one of the oceanic anoxic events that punctuate the Cretaceous and represent Earth system-wide changes prominently captured by carbon isotope excursions (CIE). Although the positive CIE near the Albian/Cenomanian boundary (ACB) is known from multiple localities, many of these records disagree in the shape of the δ13C curve. Interpretations of OAE 1d and a broader ACB Event (ACBE) remain ambiguous, and their duration is insufficiently constrained. Here we present new analyses on high-resolution samples from Core Jásd-42. The >450 m thick Pénzeskút Marl Fm. provides an expanded record across the ACBE. We generated elemental and stable isotope geochemical data and used these time series for cyclostratigraphic analyses. The derived astrochronologic age model establishes a depositional duration of 4.4–4.83 Myr and is integrated with ammonite and planktonic foraminifera biostratigraphy. This high-resolution chemostratigraphy provides a new, calibrated record of OAE 1d and the ACBE that we correlate with other reference sections, including the GSSP. A minimum estimate of 1.23 Myr and 2.28 Myr is obtained for OAE 1d and the ACBE, respectively. For the latter, a minimum of 602 kyr is assigned to the rising limb, 748 kyr to the culminating interval, and 931 kyr to the falling limb in the δ13Ccarb curve. The δ13C record through OAE 1d appears orbitally controlled, where the positive peaks are paced by long eccentricity. Considering the differences between seasonality extremes, we suggest a model for depositional cyclicity in a synorogenic basin with a high sedimentation rate of 9–15 cm/kyr.
{"title":"Integrated astrochronology, isotope and biostratigraphy of the Albian-Cenomanian transition and expression of the OAE 1d in an expanded record of a rapidly subsiding Tethyan synorogenic basin (Core Jásd-42, Transdanubian range, Hungary)","authors":"Zsolt Vallner , Tamás Müller , Dávid Bajnai , Ottilia Szives , Lajos Ádám Csicsek , Gábor Héja , Bernát Heszler , Norbert Zajzon , József Pálfy","doi":"10.1016/j.cretres.2025.106311","DOIUrl":"10.1016/j.cretres.2025.106311","url":null,"abstract":"<div><div>The OAE 1d is one of the oceanic anoxic events that punctuate the Cretaceous and represent Earth system-wide changes prominently captured by carbon isotope excursions (CIE). Although the positive CIE near the Albian/Cenomanian boundary (ACB) is known from multiple localities, many of these records disagree in the shape of the δ<sup>13</sup>C curve. Interpretations of OAE 1d and a broader ACB Event (ACBE) remain ambiguous, and their duration is insufficiently constrained. Here we present new analyses on high-resolution samples from Core Jásd-42. The >450 m thick Pénzeskút Marl Fm. provides an expanded record across the ACBE. We generated elemental and stable isotope geochemical data and used these time series for cyclostratigraphic analyses. The derived astrochronologic age model establishes a depositional duration of 4.4–4.83 Myr and is integrated with ammonite and planktonic foraminifera biostratigraphy. This high-resolution chemostratigraphy provides a new, calibrated record of OAE 1d and the ACBE that we correlate with other reference sections, including the GSSP. A minimum estimate of 1.23 Myr and 2.28 Myr is obtained for OAE 1d and the ACBE, respectively. For the latter, a minimum of 602 kyr is assigned to the rising limb, 748 kyr to the culminating interval, and 931 kyr to the falling limb in the δ<sup>13</sup>C<sub>carb</sub> curve. The δ<sup>13</sup>C record through OAE 1d appears orbitally controlled, where the positive peaks are paced by long eccentricity. Considering the differences between seasonality extremes, we suggest a model for depositional cyclicity in a synorogenic basin with a high sedimentation rate of 9–15 cm/kyr.</div></div>","PeriodicalId":55207,"journal":{"name":"Cretaceous Research","volume":"181 ","pages":"Article 106311"},"PeriodicalIF":1.7,"publicationDate":"2026-01-07","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"146037875","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
Pub Date : 2025-12-31DOI: 10.1016/j.cretres.2025.106310
Jiří Kvaček , Fritz Messner , Franz Bernhard
Upper Cretaceous flora (upper Santonian - lower Campanian) is described from the Kainach Gosau Group sediments in Styria, Austria, in the Central Eastern Alps. The flora is associated with thin coal seams. It consists of angiosperms dominated by monocotyledon foliage Pandanites trinervis, which is typical for coal-bearing strata of Gosau. The second most abundant angiosperm foliage in the locality of Geistthal is platanoid foliage Ettingshausenia geistthalensis sp. nov. Twelve dicotyledon leaves are described from the area of Kainach Gosau, including three new species Juglandiphyllites kainachensis sp. nov., J. graefii sp. nov. and J. roemaskogelensis sp. nov. Ferns and conifers (Geinitzia reichenbachii) are present, but in a distinct minority. The preserved flora is interpreted as vegetation of three types: 1) alluvial plain vegetation dominated by platanoids, 2) coal swamp vegetation dominated by pandanoids, 3) upland vegetation dominated by probable juglandoids. The terrestrial vegetation grew on the tectonically driven orogenic island, a part of the Adriatic Plate, in the northern part of the Tethys Sea. The massive occurrence of the Pandanites monocot is important not only as a palaeoecological indicator, but also for phytogeographical interpretations. As a thermophilous plant, it was tectonically delivered from the African coast to the southern coast of Europe by tectonic plate movement.
{"title":"Upper Cretaceous flora of the Gosau Group of Kainach, Styria, Austria","authors":"Jiří Kvaček , Fritz Messner , Franz Bernhard","doi":"10.1016/j.cretres.2025.106310","DOIUrl":"10.1016/j.cretres.2025.106310","url":null,"abstract":"<div><div>Upper Cretaceous flora (upper Santonian - lower Campanian) is described from the Kainach Gosau Group sediments in Styria, Austria, in the Central Eastern Alps. The flora is associated with thin coal seams. It consists of angiosperms dominated by monocotyledon foliage <em>Pandanites trinervis</em>, which is typical for coal-bearing strata of Gosau. The second most abundant angiosperm foliage in the locality of Geistthal is platanoid foliage <em>Ettingshausenia geistthalensis</em> sp. nov. Twelve dicotyledon leaves are described from the area of Kainach Gosau, including three new species <em>Juglandiphyllites kainachensis</em> sp. nov., <em>J. graefii</em> sp. nov. and <em>J. roemaskogelensis</em> sp. nov. Ferns and conifers (<em>Geinitzia reichenbachii</em>) are present, but in a distinct minority. The preserved flora is interpreted as vegetation of three types: 1) alluvial plain vegetation dominated by platanoids, 2) coal swamp vegetation dominated by pandanoids, 3) upland vegetation dominated by probable juglandoids. The terrestrial vegetation grew on the tectonically driven orogenic island, a part of the Adriatic Plate, in the northern part of the Tethys Sea. The massive occurrence of the <em>Pandanites</em> monocot is important not only as a palaeoecological indicator, but also for phytogeographical interpretations. As a thermophilous plant, it was tectonically delivered from the African coast to the southern coast of Europe by tectonic plate movement.</div></div>","PeriodicalId":55207,"journal":{"name":"Cretaceous Research","volume":"180 ","pages":"Article 106310"},"PeriodicalIF":1.7,"publicationDate":"2025-12-31","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"145980853","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
Pub Date : 2025-12-31DOI: 10.1016/j.cretres.2025.106309
Rodrigo A. Otero , Sergio Soto-Acuña , Raúl Ugalde , Patricio Sepúlveda , Alexander O. Vargas , Guillermo Aguirrezabala , Héctor Ortiz
Teeth of bramble sharks (Echinorhinidae) are one of the scarcest chondrichthyan elements reported to date in the Upper Cretaceous of central Chile and in general, South America. Prior to this research, its local record was restricted to only four available specimens, previously considered to be equivalent to teeth from the upper Maastrichtian of Argentinean Patagonia, being these referred to the species Echinorhinus maremagnum. This contribution presents two new elements, including the first complete large tooth from central Chile, showing the presence of a distally projected basal cusplet and a complex mesial, serrated cutting edge with the primordium of two clear cusplets, this last being a trait commonly present in several Cenozoic species. These new features remained unnoticed in the previous material from central Chile, either due to informative portions missing or the immature growth stage of the smaller known sample. The dental features of the new specimen are unique among known species of the genus Echinorhinus; moreover, the crown traits represent an ancestral condition with respect to those features commonly present in Cenozoic species. With the new evidence, all the known Echinorhinus teeth from the lower Maastrichtian of central Chile are now referred to Echinorhinus taverai sp. nov. The occurrence of the studied material directly associated with an elasmosaur skeleton is consistent with other findings of the genus in the Southern Hemisphere, suggesting that Upper Cretaceous austral Echinorhinus spp. were opportunistic scavengers in the shallow coastal waters of southern South America.
{"title":"A new upper Cretaceous bramble shark from central Chile reveals austral diversity among Weddellian echinorhinids","authors":"Rodrigo A. Otero , Sergio Soto-Acuña , Raúl Ugalde , Patricio Sepúlveda , Alexander O. Vargas , Guillermo Aguirrezabala , Héctor Ortiz","doi":"10.1016/j.cretres.2025.106309","DOIUrl":"10.1016/j.cretres.2025.106309","url":null,"abstract":"<div><div>Teeth of bramble sharks (Echinorhinidae) are one of the scarcest chondrichthyan elements reported to date in the Upper Cretaceous of central Chile and in general, South America. Prior to this research, its local record was restricted to only four available specimens, previously considered to be equivalent to teeth from the upper Maastrichtian of Argentinean Patagonia, being these referred to the species <em>Echinorhinus maremagnum.</em> This contribution presents two new elements, including the first complete large tooth from central Chile, showing the presence of a distally projected basal cusplet and a complex mesial, serrated cutting edge with the primordium of two clear cusplets, this last being a trait commonly present in several Cenozoic species. These new features remained unnoticed in the previous material from central Chile, either due to informative portions missing or the immature growth stage of the smaller known sample. The dental features of the new specimen are unique among known species of the genus <em>Echinorhinus</em>; moreover, the crown traits represent an ancestral condition with respect to those features commonly present in Cenozoic species. With the new evidence, all the known <em>Echinorhinus</em> teeth from the lower Maastrichtian of central Chile are now referred to <em>Echinorhinus taverai</em> sp. nov. The occurrence of the studied material directly associated with an elasmosaur skeleton is consistent with other findings of the genus in the Southern Hemisphere, suggesting that Upper Cretaceous austral <em>Echinorhinus</em> spp. were opportunistic scavengers in the shallow coastal waters of southern South America.</div></div>","PeriodicalId":55207,"journal":{"name":"Cretaceous Research","volume":"180 ","pages":"Article 106309"},"PeriodicalIF":1.7,"publicationDate":"2025-12-31","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"145928914","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
Pub Date : 2025-12-27DOI: 10.1016/j.cretres.2025.106308
Carlos A. Bueno-Cebollada , Jiří Kvaček , Eduardo Barrón
Late Albian floras from eastern Iberia are mainly dominated by conifers with xeromorphic adaptations and include well-diversified fern and angiosperm communities. Some of these coniferous taxa are believed to be related to a global mass resin production period known as the Cretaceous Resin Interval (CREI), which ultimately led to the development of amber sites.
This research presents a study of late Albian macro- and mesofloras of eastern Iberia from descriptive, palaeoecological, and palaeoclimatological perspectives. The studied material was collected from four outcrops: the Cañada del Hoyo outcrop, located in the Cuenca Basin; and the Arroyo de la Pascueta, Cortes de Arenoso, and San Just outcrops, located in the Maestrazgo Basin. The collected taxa were grouped into four palaeoecological associations (PA-1 to PA-4), representing different biocoenoses from the coastline to the hinterland: Supra-to intertidal vegetal communities (PA-1), Nearshore woodlands with Araucariaceae (PA-2), Drought-adapted hinterland vegetation (PA-3), and Local riparian vegetal communities (PA-4). The dominance of taxa assigned to the genera Frenelopsis, Dammarites, and Eretmophyllum indicates that most of the assemblages are related to supratidal plant communities, a typical feature of the mid-Cretaceous Tethyan coast.
Furthermore, we reinforce the hypothesis of an Araucariaceae affinity for some of the upper Albian amber of the Maestrazgo Basin based on the presence of Brachyphyllum cf. obesum Heer (1881) and Rabagostrobus cf. hispanicus Kvaček et al. (2018). We also relate the lack of upper Albian amber-bearing strata in the Cuenca Basin to the sedimentary environment dynamics rather than to the nature of the palaeobotanical assemblages.
来自伊比利亚东部的晚阿尔巴尼亚植物区系主要以适应旱形的针叶树为主,包括多样化的蕨类和被子植物群落。其中一些针叶分类群被认为与白垩纪树脂间隔(CREI)的全球大规模树脂生产时期有关,该时期最终导致了琥珀遗址的发展。本研究从描述、古生态和古气候学的角度对伊比利亚东部的晚阿尔巴尼亚宏观和中型植物区系进行了研究。研究材料收集自四个露头:位于昆卡盆地的Cañada del Hoyo露头;以及位于Maestrazgo盆地的Arroyo de la Pascueta、Cortes de arennoso和San Just露头。将收集到的分类群划分为4个古生态群落(PA-1 ~ PA-4),代表了从海岸线到内陆不同的生物群落:超潮至潮间带植被群落(PA-1)、近岸带龙葵科林地(PA-2)、适应干旱的内陆植被群落(PA-3)和当地河岸植被群落(PA-4)。其中,Frenelopsis属、Dammarites属和Eretmophyllum属占主导地位,表明大多数组合与潮上植物群落有关,这是中白垩世特提斯海岸的典型特征。此外,基于Brachyphyllum cf. obesum Heer(1881)和Rabagostrobus cf. hispanicus kva eket al.(2018)的存在,我们加强了Araucariaceae与Maestrazgo盆地上部Albian琥珀的亲和性假设。我们还将昆卡盆地上亚系琥珀地层的缺乏与沉积环境动力学联系起来,而不是与古植物组合的性质有关。
{"title":"Late Albian floras of eastern Iberia: Insights into the palaeoenvironments and palaeoecology of fossil plant communities associated with amber-bearing sedimentary successions","authors":"Carlos A. Bueno-Cebollada , Jiří Kvaček , Eduardo Barrón","doi":"10.1016/j.cretres.2025.106308","DOIUrl":"10.1016/j.cretres.2025.106308","url":null,"abstract":"<div><div>Late Albian floras from eastern Iberia are mainly dominated by conifers with xeromorphic adaptations and include well-diversified fern and angiosperm communities. Some of these coniferous taxa are believed to be related to a global mass resin production period known as the Cretaceous Resin Interval (CREI), which ultimately led to the development of amber sites.</div><div>This research presents a study of late Albian macro- and mesofloras of eastern Iberia from descriptive, palaeoecological, and palaeoclimatological perspectives. The studied material was collected from four outcrops: the Cañada del Hoyo outcrop, located in the Cuenca Basin; and the Arroyo de la Pascueta, Cortes de Arenoso, and San Just outcrops, located in the Maestrazgo Basin. The collected taxa were grouped into four palaeoecological associations (PA-1 to PA-4), representing different biocoenoses from the coastline to the hinterland: Supra-to intertidal vegetal communities (PA-1), Nearshore woodlands with Araucariaceae (PA-2), Drought-adapted hinterland vegetation (PA-3), and Local riparian vegetal communities (PA-4). The dominance of taxa assigned to the genera <em>Frenelopsis</em>, <em>Dammarites</em>, and <em>Eretmophyllum</em> indicates that most of the assemblages are related to supratidal plant communities, a typical feature of the mid-Cretaceous Tethyan coast.</div><div>Furthermore, we reinforce the hypothesis of an Araucariaceae affinity for some of the upper Albian amber of the Maestrazgo Basin based on the presence of <em>Brachyphyllum</em> cf. <em>obesum</em> Heer (1881) and <em>Rabagostrobus</em> cf. <em>hispanicus</em> Kvaček et al. (2018)<em>.</em> We also relate the lack of upper Albian amber-bearing strata in the Cuenca Basin to the sedimentary environment dynamics rather than to the nature of the palaeobotanical assemblages.</div></div>","PeriodicalId":55207,"journal":{"name":"Cretaceous Research","volume":"180 ","pages":"Article 106308"},"PeriodicalIF":1.7,"publicationDate":"2025-12-27","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"145928913","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
Pub Date : 2025-12-23DOI: 10.1016/j.cretres.2025.106306
Marianella Talevi , Soledad Brezina , Darío G. Lazo
Research on ecological successions in recent skeletal remains, such as those of whales, has revealed specific colonization patterns and four distinct succession phases. These succession patterns can also be inferred in the fossil record from evidence such as bioerosion traces, bone encrustation, and associated body fossils. The study of ecological successions in fossil bones of Mesozoic marine reptiles not only helps to understand the phases of degradation and entombing in the past but also contributes to our knowledge of the communities associated with these carcasses during that time. In this work, the presence of sclerobionts (represented by micro and macroborings) in bone remains of a plesiosaur from the upper Maastrichtian of Antarctica was identified and described, interpreting the phases of ecological succession in a high-latitude marine environment. Macroborings were assigned to Trypanites and Osedacoides. The microborings, similar to Wedl tunnels, were attributed to cyanobacteria and fungi, indicating that the bones were exposed for a long time. Furthermore, framboid pyrite in the bones shows anaerobic stages associated with bacterial activity. The four known stages of ecological succession were documented: the mobile scavenger stage, identified by bite marks; the opportunistic enrichment stage by Osedacoides; the sulfophilic stage, marked by modified vascular channels filled with pyrite; and the reef stage, by Trypanites traces, which indicate colonization by organisms before final burial. These findings enhance our understanding of the interactions between microorganisms and mineralization in ancient marine vertebrates, and aid in reconstructing paleocommunities associated with marine reptile falls, offering valuable insights into past ecosystem dynamics.
{"title":"Successional stages of a Maastrichtian plesiosaur fall community from Antarctica","authors":"Marianella Talevi , Soledad Brezina , Darío G. Lazo","doi":"10.1016/j.cretres.2025.106306","DOIUrl":"10.1016/j.cretres.2025.106306","url":null,"abstract":"<div><div>Research on ecological successions in recent skeletal remains, such as those of whales, has revealed specific colonization patterns and four distinct succession phases. These succession patterns can also be inferred in the fossil record from evidence such as bioerosion traces, bone encrustation, and associated body fossils. The study of ecological successions in fossil bones of Mesozoic marine reptiles not only helps to understand the phases of degradation and entombing in the past but also contributes to our knowledge of the communities associated with these carcasses during that time. In this work, the presence of sclerobionts (represented by micro and macroborings) in bone remains of a plesiosaur from the upper Maastrichtian of Antarctica was identified and described, interpreting the phases of ecological succession in a high-latitude marine environment. Macroborings were assigned to <em>Trypanites</em> and <em>Osedacoides</em>. The microborings, similar to Wedl tunnels, were attributed to cyanobacteria and fungi, indicating that the bones were exposed for a long time. Furthermore, framboid pyrite in the bones shows anaerobic stages associated with bacterial activity. The four known stages of ecological succession were documented: the mobile scavenger stage, identified by bite marks; the opportunistic enrichment stage by <em>Osedacoides;</em> the sulfophilic stage, marked by modified vascular channels filled with pyrite; and the reef stage, by <em>Trypanites</em> traces, which indicate colonization by organisms before final burial. These findings enhance our understanding of the interactions between microorganisms and mineralization in ancient marine vertebrates, and aid in reconstructing paleocommunities associated with marine reptile falls, offering valuable insights into past ecosystem dynamics.</div></div>","PeriodicalId":55207,"journal":{"name":"Cretaceous Research","volume":"181 ","pages":"Article 106306"},"PeriodicalIF":1.7,"publicationDate":"2025-12-23","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"145939330","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
In this work, 47 isolated small theropod teeth from different microsites of “El Gallo” Formation, Baja California, Mexico are studied. Isolated theropod teeth constitute important evidence to analyse theropod diversity, since they are quite common in the fossil record. For their identification, we conducted morphological comparisons using multivariate and cladistic analyses comparing them with previously described small theropod teeth from Upper Cretaceous North America Formations. This dental material can be assigned mainly to two families: Dromaeosauridae, Troodontidae; and one subfamily: Saurornitholestinae. Also, some of the teeth were assigned to Richardoestesia and two specimens were indeterminate. Dromaeosaurids represent the most abundant group of theropods in “El Gallo” followed by the Richardoestesia dental Morphotype. This diversity pattern is similar to the one reported in the Aguja Formation of Texas for the same period of time. The use of multivariate statistical techniques and cladistic analyses allowed us to evaluate the similarity between specimens, however, taxonomical assignments are difficult due to the lack of knowledge of intraspecific morphological and/or ontogenetic variation that can lead to misinterpretations. Even so, the sample of 47 specimens yields valuable information that allows suggesting the presence of certain taxa and providing knowledge of the diversity of the continental Cretaceous fauna in Mexico.
{"title":"Isolated teeth of small theropods from the El Gallo Formation, Baja California, Mexico","authors":"Vanessa Alexandra García-Gil , Angélica Torices , Mirella López-Miguel , Marisol Montellano-Ballesteros","doi":"10.1016/j.cretres.2025.106292","DOIUrl":"10.1016/j.cretres.2025.106292","url":null,"abstract":"<div><div>In this work, 47 isolated small theropod teeth from different microsites of “El Gallo” Formation, Baja California, Mexico are studied. Isolated theropod teeth constitute important evidence to analyse theropod diversity, since they are quite common in the fossil record. For their identification, we conducted morphological comparisons using multivariate and cladistic analyses comparing them with previously described small theropod teeth from Upper Cretaceous North America Formations. This dental material can be assigned mainly to two families: Dromaeosauridae, Troodontidae; and one subfamily: Saurornitholestinae. Also, some of the teeth were assigned to <em>Richardoestesia</em> and two specimens were indeterminate. Dromaeosaurids represent the most abundant group of theropods in “El Gallo” followed by the <em>Richardoestesia</em> dental Morphotype. This diversity pattern is similar to the one reported in the Aguja Formation of Texas for the same period of time. The use of multivariate statistical techniques and cladistic analyses allowed us to evaluate the similarity between specimens, however, taxonomical assignments are difficult due to the lack of knowledge of intraspecific morphological and/or ontogenetic variation that can lead to misinterpretations. Even so, the sample of 47 specimens yields valuable information that allows suggesting the presence of certain taxa and providing knowledge of the diversity of the continental Cretaceous fauna in Mexico.</div></div>","PeriodicalId":55207,"journal":{"name":"Cretaceous Research","volume":"181 ","pages":"Article 106292"},"PeriodicalIF":1.7,"publicationDate":"2025-12-12","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"145840982","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
Pub Date : 2025-12-11DOI: 10.1016/j.cretres.2025.106302
Cheng Wu , Lin Chen , Zongsheng Lu , Jiaxin Yan , Kening Lu
The extensive deposition of black shales during the Cretaceous Period formed significant petroleum resources. Sea-level rise and tectonic activity often led to seawater incursions into terrestrial lacustrine basins, a process linked to the formation of high-quality hydrocarbon source rocks. However, the precise influence of these marine transgression events on source rock deposition remains poorly constrained. This study investigates the Qingshankou Formation in the Songliao Basin by integrating paleontological, inorganic, and organic geochemical data. Four discrete marine incursion events were identified, allowing for the reconstruction of paleoclimatic conditions, depositional environments, organic matter provenance, and primary productivity levels. Based on these results, three depositional models are proposed: ‘Burst’, ‘Destroy’, and ‘Slow’. The ‘Burst’ model enhances the formation of high-quality source rocks by increasing primary productivity via marine algae and nutrient influx, which fosters an anoxic environment favorable for organic matter preservation. In contrast, the ‘Destroy’ model inhibits source rock formation, as excessive terrestrial detrital input dilutes organic matter and disrupts anoxic preservation conditions. Finally, the ‘Slow’ model has an indirect influence and typically follows the ‘Destroy’ phase. During this subsequent stage, the lacustrine environment becomes relatively stable, inheriting the abundant nutrient load from preceding phase while developing the persistent anoxia through internal biogeochemical regulation within the lake system, which is required for effective organic matter preservation. Ultimately, the interplay between high productivity and favorable preservation conditions represents the key control on source rock formation in marine-influenced lacustrine systems.
{"title":"The impact of transgression on the development model of lacustrine source rocks: Insights from the Upper Cretaceous Qingshankou Formation source rocks in the Songliao Basin","authors":"Cheng Wu , Lin Chen , Zongsheng Lu , Jiaxin Yan , Kening Lu","doi":"10.1016/j.cretres.2025.106302","DOIUrl":"10.1016/j.cretres.2025.106302","url":null,"abstract":"<div><div>The extensive deposition of black shales during the Cretaceous Period formed significant petroleum resources. Sea-level rise and tectonic activity often led to seawater incursions into terrestrial lacustrine basins, a process linked to the formation of high-quality hydrocarbon source rocks. However, the precise influence of these marine transgression events on source rock deposition remains poorly constrained. This study investigates the Qingshankou Formation in the Songliao Basin by integrating paleontological, inorganic, and organic geochemical data. Four discrete marine incursion events were identified, allowing for the reconstruction of paleoclimatic conditions, depositional environments, organic matter provenance, and primary productivity levels. Based on these results, three depositional models are proposed: ‘Burst’, ‘Destroy’, and ‘Slow’. The ‘Burst’ model enhances the formation of high-quality source rocks by increasing primary productivity via marine algae and nutrient influx, which fosters an anoxic environment favorable for organic matter preservation. In contrast, the ‘Destroy’ model inhibits source rock formation, as excessive terrestrial detrital input dilutes organic matter and disrupts anoxic preservation conditions. Finally, the ‘Slow’ model has an indirect influence and typically follows the ‘Destroy’ phase. During this subsequent stage, the lacustrine environment becomes relatively stable, inheriting the abundant nutrient load from preceding phase while developing the persistent anoxia through internal biogeochemical regulation within the lake system, which is required for effective organic matter preservation. Ultimately, the interplay between high productivity and favorable preservation conditions represents the key control on source rock formation in marine-influenced lacustrine systems.</div></div>","PeriodicalId":55207,"journal":{"name":"Cretaceous Research","volume":"181 ","pages":"Article 106302"},"PeriodicalIF":1.7,"publicationDate":"2025-12-11","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"145840983","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":3,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}