Urban biodiversity is affected by human-designed features of public squares

Cities are designed primarily for the benefit of humans but also provide habitat for other species. However, understanding how different components of urban vegetation and other features of urban spaces enable different species or species groups to live in the city remains limited. Here we show that, for the City of Munich, designed features of public urban squares strongly determine the occurrence of different species groups. While taxon richness and abundance increased with increasing ‘greenness’ of the square, different taxa responded to different square features, such as the proportion of lawn, the volume of shrubs and the density of trees, as well as the number of people or pets on these squares. Our results highlight that urban design for human needs affects other species that may cohabit these spaces. Consequently, planning strategies for biodiverse cities that aim to enhance human–nature interactions need to be multifaceted, considering the needs of humans and other taxa to create diverse living cities. The iconic image of ‘the public square’ typifies many cities in Europe and elsewhere, but they transcend spaces for socializing and public deliberation. Focusing on Munich, this study analyzes whether and how design features of public urban squares affect the broader biodiversity living there, finding that greenness matters but that different taxa respond differently to design elements.