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Sperm Ultrastructure of the Protobranchia: Comparison with Other Bivalve Mollusks and Potential Taxonomic and Phylogenetic Significance 原鳃动物的精子超微结构:与其他双壳类软体动物的比较及其潜在的分类和系统发育意义
Pub Date : 2017-06-08 DOI: 10.3158/2158-5520-17.11.1
J. Healy, P. Mikkelsen, Gonzalo Giribet, R. Bieler
Abstract Sperm ultrastructure of nine species of protobranch bivalves, representing three of four extant orders (Solemyida, Nuculida, Nuculanida), is discussed. Greatest diversity occurs in Solemyida (acrosomal vesicle low-conical, tall-conical, or very elongate, with radial plates; nucleus rod-shaped, teardrop-shaped, or very elongate; four, five, or six mitochondria) and the least in Nuculida (acrosomal vesicle low- to tall-conical; lacking radial plates; nucleus rod-shaped, five or six mitochondria) followed by Nuculanida (short, conical acrosomal vesicle with radial plates; spheroidal nucleus; four or five mitochondria). The wide variety of shapes in Solemyidae suggests taxonomic potential, especially in resolution and/or recognition of supraspecific taxa, but no diagnostic family characters were identified. Taxonomic potential exists for Nuculida (acrosomal shape) and Nuculanida (mitochondrial number). Protobranch sperm is highly diverse, and no defining character of the whole group was found. Support was found for the Nuculida and Nuculanida as natural groups but not for their close relationship. Nuculanida and Solemyida exhibit radial plates in the acrosomal vesicle but otherwise share no derived characters. The striking similarity of most sperm features of Nuculanida with certain pteriomorphians, especially Pectinoidea, a relationship also suggested by some mitochondrial DNA sequence data, poses interesting questions concerning their relationships and/or shared functional constraints.
摘要本文讨论了现存四目(重节目、Nuculida目、Nuculanida目)中三目9种原分支双壳类动物的精子超微结构。最大的多样性发生在Solemyida(顶体囊泡低圆锥形,高圆锥形,或非常长,具有放射状板;核棒状,泪滴状,或非常拉长;4、5或6个线粒体),核内最少(顶体囊泡低至高圆锥形;缺少径向板的;核棒状,有5或6个线粒体),其次是核胞体(短,圆锥形顶体囊泡,有放射状板;球状核;四个或五个线粒体)。Solemyidae中各种各样的形状提示了分类潜力,特别是在超种分类群的分辨和/或识别方面,但没有发现诊断性的科特征。在顶体形状和线粒体数量上存在分类潜力。原分支精子是高度多样化的,没有发现整个群体的定义特征。支持Nuculida和Nuculanida是自然类群,但不支持它们的亲密关系。Nuculanida和Solemyida在顶体囊泡中显示放射状板,但在其他方面没有相同的衍生特征。Nuculanida的大多数精子特征与某些翼形动物,特别是Pectinoidea惊人的相似,一些线粒体DNA序列数据也表明了这种关系,这提出了关于它们之间关系和/或共享功能限制的有趣问题。
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引用次数: 6
The Eriocaulaceae of Peru: Three taxonomic novelties, and a revised checklist with key 秘鲁仙人掌科:3个新分类,及一份带关键字的修订清单
Pub Date : 2017-05-24 DOI: 10.3158/2158-5520-17.10.1
N. Hensold
Abstract Paepalanthus piscatorum, sp. nov. (P. subsect. Polyactis) is described with two varieties endemic to the eastern Andes in Peru and Ecuador. Paepalanthus dichotomus var. glabrescens Moldenke is raised to species status as Paepalanthus glabrescens comb. et stat. nov. (P. subsect. Dichocladus), endemic to the vicinity of Chachapoyas. Affinities and floral morphology of both species are discussed in detail. The variability in Syngonanthus peruvianus is documented, with an updated species description. Paepalanthus stuebelianus is placed in synonymy of P. intermedius. In addition a revised checklist for Peru is presented, updating the account in the Catalogue of the Flowering Plants and Gymnosperms of Peru (1993), and providing global habitat and distribution information, general comments on phytogeographic patterns in the family, and a dichotomous key to Peruvian species. Images of representative specimens are provided in a supplementary online resource (http://fieldmuseum.org/fieldiana-hensold). The new total of 31 species reflects 13 new taxon records, three deleted species records, three name changes due to synonymy, and several changes in distributional records. Ten endemic taxa are reported, for which conservation status is provisionally assessed for eight, according to the International Union for Conservation of Nature guidelines. Four taxa were found to be widely disjunct from white sand areas of northern South America. Resumen Paepalanthus piscatorum sp. nov. (P. subsect. Polyactis) se describe con dos variedades, endémicas de la parte de los Andes orientales en el Perú y Ecuador. Paepalanthus dichotomus var. glabrescens Moldenke se eleva a nivel de especie como P. glabrescens, comb. et stat. nov. (P. subsect. Dichocladus), endémico a la zona de Chachapoyas. Se discuten las afinidades y la morfología floral de ambas especies en detalle. Se documenta la variabilidad en Syngonanthus peruvianus, con una descripción de la especie actualizada. Paepalanthus stuebelianus está colocado en la sinonimia de P. intermedius. Además se presenta una lista revisada de las eriocauláceas del Perú, actualizando el Catálogo de las Angiospermas y Gimnospermas del Perú (1993), incluyendo la información del hábitat y distribución global, observaciones generales sobre los patrones fitogeográficas en la familia, y una clave dicotómica para especies peruanas. Imágenes de especímenes representativos se proporcionan en un recurso suplementario disponible en línea (http://fieldmuseum.org/fieldiana-hensold). El nuevo total de 31 especies refleja 13 nuevos registros de taxones, tres registros de especies eliminadas, tres cambios de nombre debido a la sinonimia, y varios cambios en registros de distribución. Se reportan diez taxones endémicos, y el estado de conservación se evalúa provisionalmente para ocho, de acuerdo con las directrices de la UICN. Se encontraron cuatro taxones que están ampliamente disjuntos de las zonas de arena blanca de la parte norte de Sudam
摘要:鱼纹花(Paepalanthus piscatorum, sp. 11 .)Polyactis)是秘鲁和厄瓜多尔安第斯山脉东部特有的两个品种。将拟南花(Paepalanthus dichotomus var. glabrescens Moldenke)提升为梳状拟南花(Paepalanthus glabrescens comb)种。[译]〔11〕恰恰波亚附近特有的双枝属植物。详细讨论了两种植物的亲缘关系和花形态。在秘鲁Syngonanthus pervianus变异被记录,与更新的物种描述。拟南花是拟南花的同义植物。此外,还提出了秘鲁的订正清单,更新了《秘鲁开花植物和裸子植物目录》(1993年)中的记述,并提供了全球生境和分布信息,对该科植物地理格局的一般性评论,以及秘鲁物种的二分法索引。代表性标本的图像在补充在线资源(http://fieldmuseum.org/fieldiana-hensold)中提供。新发现的31种包括13个新的分类单元记录,3个被删除的物种记录,3个因同义而发生的名称变化,以及一些分布记录的变化。根据国际自然保护联盟的指导方针,报告了10个地方性分类群,其中8个被暂时评估为保护状态。在南美洲北部的白沙地区发现了四个分类群。草甸花属11 . (P.亚组)Polyactis) se描述了condos variedades, endsammicas de los Andes orientales和Perú在厄瓜多尔。无毛的毛茛,梳状。[译]〔11〕Dichocladus), endsamico a la zona de Chachapoyas。我们讨论了一些关于植物的细节,如morfología。关于秘鲁制度可变性的文件,请参阅descripción关于特别实现性的文件。张玉兰,张玉兰,张玉兰,张玉兰。此外,提供了una lista revisada de las eriocaulaceas del秘鲁,actualizando el专业de las Angiospermas y Gimnospermas del秘鲁(1993),包括可以看看德尔栖息地抽烟distribucion全球observaciones发表尤其洛杉矶赞助人fitogeograficas en la familia,, y una劈开dicotomica对位especies peruanas。Imágenes de especímenes代表们在línea (http://fieldmuseum.org/fieldiana-hensold)中看到比例和非递归补充。新物种总数为31个,反映了13个新物种登记,树木登记,树木cambios de nombre debido a la sinoniia,树木cambios en registros de distribución。这一报告是由conservación Se evalúa临时行政主管部门、联合国行政主管部门和联合国行政主管部门共同编写的。在与苏丹北部地区的白色区域的冲突中,有两个例外情况:están。
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引用次数: 4
Assembling a Taxonomic Monograph of Tribe Wirthiotremateae (Lichenized Ascomycota: Ostropales: Graphidaceae) Wirthiotremateae族分类专著汇编(地衣子囊菌纲:Ostropales:石墨科)
Pub Date : 2017-04-28 DOI: 10.3158/2158-5520-16.9.1
Ian D. Medeiros, Ekaphan Kraichak, R. Lücking, A. Mangold, H. Lumbsch
Abstract Phylogenetic studies of the lichenized fungal family Graphidaceae necessitate various nomenclatural changes. Tribe Wirthiotremateae is more narrowly circumscribed on the basis of new molecular data and phenotypical differences towards allied clades. The new genus Austrotrema is described to accommodate the three species of the Thelotrema bicinctulum group, Asteristion is resurrected for the seven species of the Chapsa platycarpa group, and Nadvornikia is expanded to include two non-mazaediate species in addition to its two mazaediate species. Asteristion australianum is newly described for Australian material previously identified as T. albo-olivaceum. Eleven new combinations are made: Asteristion alboannuliforme (Bas.: Thelotrema alboannuliforme), As. albo-olivaceum (Bas.: T. albo-olivaceum), As. cupulare (Bas.: T. cupulare), As. leucophthalmum (Bas.: T. leucophthalmum), As. platycarpoides (Bas.: T. platycarpoides), As. platycarpum (Bas.: T. platycarpum), Austrotrema bicinctulum (Bas.: T. bicinctulum), Au. myriocarpum (Bas.: T. myriocarpum), Au. terebrans (Bas.: T. terebrans), Nadvornikia expallescens (Bas.: Leucodecton expallescens), and N. peninsulae (Bas.: Myriotrema peninsulae). Descriptions and discussions of Wirthiotremateae species are provided, along with keys to the species of the tribe. Phylogenetic analyses were based on six markers: the mitochondrial small subunit (mtSSU) and nuclear large subunit (nuLSU) rDNA, RNA polymerase II largest (RPB1) and second largest (RPB2) subunit, elongation factor 1 alpha (EF1-α), and internal transcribed spacer (ITS).
地衣真菌石墨科的系统发育研究需要进行各种命名变化。Wirthiotremateae部落在新的分子数据和与亲缘分支的表型差异的基础上被更狭窄地限定。新属Austrotrema被描述为容纳theelotrema bicinctulum组的三个物种,Asteristion被复活为Chapsa platycarpa组的七个物种,Nadvornikia被扩展为除了两个马扎虫物种之外还包括两个非马扎虫物种。澳大利亚Asteristion australium是新发现的澳大利亚材料,以前被鉴定为T. albo-olivaceum。11种新组合:Asteristion alboannulformme (Bas;;;;;;albo-olivaceum (Bas)。;;;;cupulare (Bas)。(T. cupulare);leucophthalmum (Bas)。;;;;platycarpoides (Bas)。:鸭嘴兽);platycarpum (Bas)。:桔梗(T. platycarpum);: T. bicinctulum);myriocarpum (Bas)。:肉豆蔻);terebrans (Bas)。: T. terebrans), Nadvornikia expallescens (Bas。: Leucodecton expallescens)和N. peninsula (Bas。:密里奥特里马半岛)。提供了Wirthiotremateae物种的描述和讨论,以及该部落物种的关键。系统发育分析基于6个标记:线粒体小亚基(mtSSU)和核大亚基(nuLSU) rDNA、RNA聚合酶II最大亚基(RPB1)和第二大亚基(RPB2)、延伸因子1 α (EF1-α)和内部转录间隔段(ITS)。
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引用次数: 20
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Fieldiana Life and Earth Sciences
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