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Diminutive Epibionts on Crinoid Stems from the Lower/Middle Devonian Boundary in the Barrandian, Czech Republic 捷克巴兰地地区下/中泥盆世界线海百合茎上的小型表观生物
M. Mergl, Nikola Šmídtová
Abstract Diminutive crinoid holdfasts and cemented tests of the foraminifers Psammosphaera and Tolypammina were observed on coarse bioclasts in weathered limestones of the Daleje-Třebotov Formation. Specimens were obtained in 1984 by washing so called “white beds” at a temporary locality in Praha- Barrandov. A few millimeter sized bioclasts with epibionts were freed from hard limestone beds of the Třebotov Limestone near the Lower/Middle Devonian boundary by long-term weathering. Many of the crinoid holdfasts attached to pluricolumnals provoked a stereomic response of the host crinoid. Also the growth orientation of the crinoid epibiont is not random and indicates some crinoid-epibiont to crinoidhost interaction. Reaction of host stereome and non-random stem orientation offer direct evidence of epibiont larval settlement and subsequent growth on the stem of a living crinoid host. The extensive growth of the host stereome ended by partial to total engulfing of the epibiont holdfast. This indicates advancing and finally successful defence of the host crinoid against the epibiont. The holdfast gives evidence that the small host crinoids offered a somewhat higher tier for even smaller epibiont crinoids. However, other observed holdfasts indicate fixation of larva and growth over loose bioclasts lying on a sea bed. Location of foraminifer test on bioclasts confims that foraminifers cemented and grew on loose echinodermal and brachiopod remains and preferred crevices and similar protected sites with concave profiles. This is clear evidence that diverse bioclasts (brachiopod shells, pelmatozoan ossicles) provided the hard substrate suitable for epibiont life on a sea bed.
摘要/ Abstract摘要:在Daleje-Třebotov组风化灰岩的粗粒生物碎屑中,观察了有孔虫Psammosphaera和Tolypammina的小型海红类固结物和胶结试验。1984年,在普拉哈-巴兰多夫的一个临时地点,通过清洗所谓的“白床”获得了标本。在中、下泥盆统界线附近的Třebotov灰岩中,经长期风化作用,从坚硬的灰岩层中释放出若干毫米大小的生物碎屑。许多附着在多柱上的海百合支架引起了宿主海百合的立体反应。此外,海百合表面生物的生长方向也不是随机的,表明海百合表面生物与海百合寄主存在相互作用。寄主体体的反应和茎的非随机取向提供了幼虫在活的海百合寄主茎上定居和随后生长的直接证据。寄主体体体的广泛生长以部分或全部吞噬表面菌体而结束。这表明寄主海百合最终成功防御了外生体。这一结果表明,小的寄主海百合为更小的外生海百合提供了更高的层次。然而,其他观察到的固结物表明幼虫的固定和生长在海底松散的生物碎屑上。有孔虫在生物碎屑上的定位试验证实,有孔虫在松散的棘皮动物和腕足动物遗骸上以及偏爱的裂缝和类似的凹形保护部位上胶结生长。这是一个明确的证据,表明不同的生物碎屑(腕足动物的壳,贝壳类动物的小骨)提供了适合海底表面生物生活的坚硬基质。
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引用次数: 0
Construction of the Metro Line D in Praha-Pankrác: An Introductory Report About a Unique Opportunity for Study of the Upper Ordovician Fossil Assemblages and Sediments in the Prague Basin (Czech Republic) Praha-Pankrác地铁D线建设:捷克布拉格盆地上奥陶统化石组合与沉积物研究的独特机会
J. Bruthansová, J. Bruthans, P. Kraft
Abstract The construction of the new metro line D in Praha-Pankrác provides a unique opportunity to study different aspects such as lithology, stratigraphy and fossil assemblages from the Upper Ordovician and Silurian of the Prague Basin. Results from the sections in tunnels mined so far allowed detailed information about the succession of fossil assemblages, facies variability and actual thicknesses of the upper part of the Bohdalec Formation, the Králův Dvůr and Kosov formations in this part of Prague Basin. The stratigraphic position of Michle Facies in the Bohdalec Formation was also indicated.
Praha-Pankrác新地铁D线的建设为研究布拉格盆地上奥陶统和志留系的岩性、地层学和化石组合等不同方面提供了独特的机会。迄今为止挖掘的隧道剖面的结果提供了关于布拉格盆地这部分Bohdalec组上部、Králův Dvůr组和Kosov组的化石组合演为、相变化和实际厚度的详细信息。指出了Bohdalec组michel相的地层位置。
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引用次数: 1
The Earliest Cyrtina (Brachiopoda, Cyrtinidina) in the Devonian of the Barrandian (Czech Republic) Barrandian(捷克共和国)泥盆纪最早的Cyrtina(腕足目,Cyrtinidina)
M. Mergl
Abstract The earliest known Cyrtina in the Prague Basin has been discovered in the Kotýs Limestone of the Lochkov Formation (Lochkovian) among a rich brachiopod-coral fauna at Branžovy ridge near Bubovice (Beroun District, Czechia). Rare and imperfectly preserved silicified shells are assigned to Cyrtina praecedens Kozłovski, 1929, a species originally described from Podolia, Ukraine. The species is known also in north-eastern Russia (Tajmyr and Sette-Daban Mts) and likely also in New South Wales, Australia. Its distribution provides evidence of the rapid spread of Cyrtina across the shallow shelves of Laurussia, Siberia and Gondwana in the Early Devonian. The Devonian and Carboniferous distribution of Cyrtina is restricted to the agitated, shallow-water carbonate environment in tropical and temperate climatic belts.
布拉格盆地已知最早的Cyrtina是在靠近Bubovice (Beroun地区)的Branžovy山脊的Lochkov组(Lochkovian)的Kotýs石灰岩中发现的。稀有且保存不完整的硅化壳归属于Cyrtina preecedens Kozłovski, 1929,一个最初描述于乌克兰Podolia的物种。该物种在俄罗斯东北部(Tajmyr和Sette-Daban mt)也有发现,可能在澳大利亚的新南威尔士州也有发现。它的分布证明了早泥盆世古陆、西伯利亚和冈瓦纳的浅海陆架上古陆的快速传播。泥盆系和石炭系水草的分布局限于热带和温带气候带的激流、浅水碳酸盐环境。
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引用次数: 0
Seminal Insights Concerning the Variety of Sidertic Structures and Variable Genesis and of Sideritic Occurrences: An Unexplored Source of Palaeontological Information (Sydney Coalfield, Middle Pennsylvanian, Canada) 关于菱铁矿结构多样性、成因变化和产状的开创性见解:一个未开发的古生物信息来源(加拿大中宾夕法尼亚悉尼煤田)
E. Zodrow, J. Pšenička
Abstract Reported are descriptions of twelve samples representing a variety of sideritic structures, including nodules mostly from roof shale of the Middle Pennsylvanian Sydney Coalfield, Canada. The co-occurrence of fossiliferous nodules and compression fossils in the shaley roof rocks at Point Aconi enhance greatly palaeontological information. Newly discovered in a coal seam, and part of the sample, is a 40 mm thick continuous sheet-like layer of siderite with abundant permineralized-like-compressed small to micron-sized structures in a rather evenly-sized sideritic matrix, probably indicating a genetic origin different from that of the nodules. Methods include some thin-section, and two X-Ray analysis. However, large systematic sampling is a prerequisite to explore that situation, which additionally could provide faunal information for Euromerican correlation.
摘要本文报道了12个代表各种菱铁矿结构的样品,其中包括结核,主要来自加拿大中宾夕法尼亚悉尼煤田的顶板页岩。阿科尼角泥质顶板岩中化石结核与压缩化石的共生,极大地增强了古生物学信息。在煤层和部分样品中新发现的是一个40毫米厚的连续片状菱铁矿层,在相当均匀的菱铁矿基质中有丰富的过矿化压缩小到微米大小的结构,可能表明其成因与结核不同。方法包括一些薄切片和两次x射线分析。然而,大规模的系统采样是探索这种情况的先决条件,它还可以为欧美相关性提供动物区系信息。
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引用次数: 0
Heterosporangia in Isoetes Pantii (Isoetaceae, Pteridophyta): Revealing the Beginnings of Heterospory and Recalling Paleozoic Ancestors? 泛水韭(水韭科,蕨类植物)的异孢子体:揭示异孢子的起源并回顾古生代的祖先?
J. Bek, H. Goswami
Abstract The extant plant genus Isoetes (Isoetaceae; lycophyte, quillwort) is important from an evolutionary point of view. Species of this heterosporous genus are small herbs (up to 50 centimeters) and exhibit some morphological, anatomical and embryological features of their Paleozoic arborescent lycopsid ancestors. The species Isoetes pantii produces three kinds of microspores (monolete, alete and trilete) and two types of trilete megaspores in one and the same heterosporangium. We attempt to associate these unusual functional megaspores with various Paleozoic spores described mainly from Devonian barinophytaleans such as Omniastrobus dawsonii, Barinophyton richardsonii, B. citrulliforme and Protobarinophyton pennsylvanicum. These have two kinds of spores in a sporangium and provide the first palynological evidence of heterospory at 405 Ma. The germination of microspores and megaspores and production of gametophytes within the heterosporangia of I. pantii corresponds with that of some of its Paleozoic ancestors. Retention of megaspores within heterosporangia and their germination in situ offers evidence that I. pantii exhibits the probable route of evolution of the seed habit. These observations support the hypothesis that a typical heterosporangium was the cradle for the evolution of heterospory.
摘要:现存的水韭属植物;从进化的角度来看,石松(冬虫夏草)是重要的。这个异孢子属的物种是小型草本植物(可达50厘米),并表现出其古生代树栖石松祖先的一些形态、解剖和胚胎学特征。在一个异孢囊中产生三种小孢子(单孢子、双孢子和三孢子)和两种三孢子大孢子。我们试图将这些不寻常的功能大孢子与各种主要来自泥盆纪钡生植物的古生代孢子联系起来,如Omniastrobus dawsonii, Barinophyton richardsonii, B. citrulliforme和Protobarinophyton pennsylvania。它们在一个孢子囊中有两种孢子,提供了405 Ma异孢子的第一个孢粉学证据。异孢子囊内小孢子、大孢子的萌发和配子体的产生与其古生代的一些祖先相一致。异孢子囊内大孢子的保留及其原位萌发提供了证据,表明大孢子具有种子习性的可能进化途径。这些观察结果支持了一个典型异孢子囊是异孢子进化的摇篮的假设。
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引用次数: 0
The Rare Trilobites Exallaspis (Odontopleuridae) and Kosovoproetus? (Tropidocoryphidae) in the Homerian (Silurian) Of The Prague Basin, Bohemian Massif 稀有三叶虫(齿胸虫科)和科索沃三叶虫?波西米亚山脉布拉格盆地荷马纪(志留纪)中的(棘球绦虫科)
František Hartl, V. Vokáč, M. Pavlovic, T. Mrázek, Martin Šafka
Abstract The deep-water Aulacopleura koninckii Assemblage in the lower Homerian (T. testis Sub-Biozone) “Aulacopleura shales“ strata at the classical ‘Barrande’s pits’ locality on the Černidla hillside at Loděnice in the Prague Basin is supplemented by the addition of two new trilobite taxa, viz. Exallaspis? perunicana sp. n. and Kosovoproetus? aff. praecursor (Přibyl & Vaněk 1987). The palaeogeographic distribution of Exallapis Ramsköld & Chatterton, 1991 is extended by the occurrence of Exallaspis? perunicana sp. n. and the younger Exallaspis sp. in the upper Homerian (the P. parvus – G. nassa Biozone) of the Prague Basin, reflecting the faunal migration between the southern shelf of the Baltica palaeocontinent and the Perunica microcontinent accross the Rheic Ocean. The small dimensions of the K.? aff. praecursor exoskeletons compared to those of K.? praecursor in the bordering shallow-water Liolalax–Sphaerexochus– Cheirurus Assemblage represent another example of adaptive nanism in trilobites of the Aulacopleura-Raphiophorus Biofacies.
在布拉格盆地lod尼斯Černidla山坡上经典的“Barrande’s pits”地区的“Aulacopleura shale”地层中,深水Aulacopleura koninckii组合被两个新的三叶虫分类群Exallaspis?科索沃共和国和科索沃共和国?事件前兆(Přibyl & vank 1987)。Exallaspis Ramsköld & Chatterton, 1991的古地理分布因Exallaspis?布拉格盆地上Homerian (P. parvus - G. nassa生物带)中的perunicana sp.和更年轻的Exallaspis sp.,反映了波罗的海古大陆南部陆架和perunicica微大陆之间跨越里海的动物迁徙。k的小尺寸?前驱外骨骼与k ?浅水边缘Liolalax-Sphaerexochus - Cheirurus组合的前体是aulacopleura - raphiphohorus生物相三叶虫适应性nanism的另一个例子。
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引用次数: 0
Source of K-Feldspars and Kaolinitization in Arkoses of West and Central Bohemian Continental Permo–Carboniferous 西、中波西米亚大陆二叠纪-石炭系岩溶中钾长石的来源及高岭石成矿作用
Jiří Jiránek, J. Klomínský, Jana Jiránková
Abstract The examined K-feldspars in arkoses of the continental Upper Carboniferous – Lower Permian of the West and Central Bohemian basins originated in the whole bed sequence (Duckmantian to Autunian; 314.2–297.1 Ma) from one source, which were so-called mountain granites of the older intrusive complex (OIC) of the Krušné hory Mts, as confirmed by the X-ray and geochemical analyses. The presence of feldspar clasts from other granitoids (Merklín and Louny massifs) was found only in the straight transgreding basal Carboniferous rocks. While other arkoses underwent synsedimentary and/or post-sedimentary kaolinization, in the deposits of Kaznějov and Horní Bříza (Nýřany Member of the Kladno Formation) pre-sedimentary kaolinization (sedimentation of sandstones and conglomerates with kaolinitic cement) occurred.
西、中波希米亚盆地上石炭统—下二叠统陆相长石中的钾长石产于整个层序(Duckmantian—autuniian;314.2 ~ 297.1 Ma),经x射线和地球化学分析证实,它们是Krušné霍利山较老侵入杂岩(OIC)的山地花岗岩。其他花岗岩类(Merklín和Louny地块)的长石碎屑仅存在于直越界的石炭系基底岩中。虽然其他砂岩经历了同沉积和/或沉积后的高岭土化,但在kaznuzov和Horní Bříza (Nýřany属于Kladno组)的沉积中,发生了沉积前的高岭土化(砂岩和砾岩的高岭土胶结物沉积)。
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引用次数: 0
First Record of Podichnus in Byronid Shell from the Lower Devonian (Pragian) of the Prague Basin, Czechia 捷克布拉格盆地下泥盆世(布拉格语)贝龙壳中Podichnus的首次记录
M. Mergl
Abstract Brachiopod etching trace Podichnus, typically with centrifugally arranged clusters of holes or slits, has hitherto been known only in calcium carbonate substrates. The similar etching trace is newly described on calcium phosphate substrate. The trace Podichnus isp. in a wall of a phosphatic byronid test was observed in the lower Devonian Vinařice Limestone (Pragian) in the Koněprusy area of the Prague Basin in the Barrandian area, the Czech Republic. The trace is smaller that majority of described species of this fixichnia, and displays a smooth central disc surrounded by two circlets of holes or pits. Some holes penetrate through wall of byronid test without any biotic response of a byronid. The maker of Podichnus isp. is uknown but among the associated fossils are eligible candidates including rhynchonellids, orthids and terebratulids. It is rare direct evidence of etching activity of the pedicle in the Lower Palaeozoic and the first finding of Podichnus in the Devonian in the Prague Basin. The emended diagnosis of the ichnogenus is presented herein.
腕足动物蚀刻痕迹Podichnus,通常具有离心排列的孔洞或狭缝簇,迄今为止仅在碳酸钙基质中发现。在磷酸钙衬底上也发现了类似的蚀刻痕迹。波迪克努斯的踪迹。在捷克共和国巴兰地地区布拉格盆地konkonuprusy地区的下泥盆纪Vinařice石灰石(布拉格)中观察到一堵墙内的磷酸盐拜伦试验。这一痕迹比大多数已描述的物种都要小,并显示出一个光滑的中央圆盘,周围环绕着两个孔或坑。有些孔洞可以穿透体液壁而没有任何体液的生物反应。Podichnus isp的制造商。是未知的,但在相关的化石中有合格的候选者,包括舌足动物,矫形动物和terebratulids。这是下古生代罕见的弓根刻蚀活动的直接证据,也是布拉格盆地泥盆纪首次发现的Podichnus。本文提出了鱼属的修正诊断。
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引用次数: 1
3D Chemometric Model Simulating the Acitheca Polymorpha Frond: Implications for Reconstructing Carboniferous Ferns (Marattiales, Canada) 模拟Acitheca Polymorpha叶片的三维化学计量学模型:对石炭纪蕨类植物重建的启示
J. D'Angelo, E. Zodrow, J. Pšenička
Abstract Reported are results of an initial approximate imitation of a Carboniferous fern frond, i.e., marattialean Acitheca polymorpha (Schimper), Middle Pennsylvanian Sydney Coalfield, Canada. The simulation experiment is based on the analysis of 14 infrared spectra obtained by means of Fourier Transform Infrared spectroscopy from four detached fragments of sterile polymorphic penultimate-pinna compressions. The calculated relative, semi-quantitative, chemical data from the infrared spectra are the input for principal component analysis deriving a 3D (three-dimensional) chemometric model. To interpret it, the four specimens are placed in hypothetical-frond positions simulating a tripinnate frond, based on diminishing penultimate-rachial widths from 1-mm (distal) to 10-mm (proximal). Hypothetical conclusions include position-dependent chemistries, specifically that of opposing trends of aromaticity vs. aliphaticity in pinnules-rachises. This, in turn, would suggest potential for (i) fern-frond reconstruction, and (ii) for determination of a most likely frond position of fragmentary specimens by “chemical classification”; the predictive aspect. However, further experimental refinement is necessary particularly based on larger frond segments to confirm or disconfirm the overall hypothetical results.
摘要本文报道了一种石炭系蕨类植物叶片的初步近似模拟结果,即,maratialean Acitheca polymorpha (Schimper), mid pennsylvania Sydney Coalfield, Canada。利用傅里叶变换红外光谱对4个分离的不育多态次耳廓压缩片段进行了14个红外光谱分析,并进行了仿真实验。从红外光谱中计算出的相对、半定量的化学数据是主成分分析的输入,推导出三维化学计量模型。为了解释这一点,根据倒数第二臂宽度从1毫米(远端)减少到10毫米(近端),将四个标本放置在模拟三棱叶的假设前肢位置。假设的结论包括位置依赖的化学反应,特别是芳香性和脂肪性的相反趋势。反过来,这将表明:(1)蕨类植物叶片重建的潜力,以及(2)通过“化学分类”确定碎片标本最可能的叶片位置的潜力;预测方面。然而,进一步的实验改进是必要的,特别是基于更大的前段,以证实或否定整体假设结果。
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引用次数: 1
The Cuticles of (?) Thylacocephalan Arthropod from the Basal Choteč Event (Choteč Formation, Eifelian; Barrandian Area, Czech Republic) 角质层(?)Eifelian基choteik组的囊头类节肢动物巴兰地地区,捷克共和国)
M. Mergl
Abstract Small fragments of phosphatic cuticle have been observed in dark limestone of the early Eifelian age (Choteč Formation) in the interval of the Basal Choteč Event. The cuticle is two-layered, primarily folded, with a chamber between outer and inner walls. Fragments likely represent small cuticle pieces from the margins of the carapace. The exterior of the cuticle is nearly smooth bearing irregular network of wrinkled polygons or shallow pits. Low conical mound-like to high thorn-like spines with annular structure extend from both outer and inner surface of cuticle. Wrinkled and folded bases of these spines indicate moderate flexibility of cuticle. Spines are hollow, the higher ones often with apical opening. The inner surface of carapace carries smaller spines or is nearly smooth. Chamber walls inside the cara-pace are with folds and other structures supporting stiffness of the cuticle. The internal walls of the cuticle are covered by polygonal bumps. These uniformly sized and shaped bumps are about 1 μm sized and likely represents imprints of the epithelial cells adjoined to the basal membranous layer of endocuticle. Biological affinity of cuticle fragments is unclear. They surely represent pieces of the arthropod cara-pace, the most probably a thylacocephalan. Associated fossils indicate a deeper marine environment. Bloom of prasinophytes, abundance of dacryoconarids and organophosphatic brachiopods, and striking rarity and diminutive size of other fauna indicate eutrophic conditions in a neritic sea, likely with hypoxic bottom water. Nectonic mode of life in open sea can be suggested for an animal bearing this cuticle.
摘要:在基底乔特基事件期间,在Eifelian早期(乔特基组)的深色石灰岩中观察到细小的磷化角质层碎片。角质层是两层的,主要是折叠的,在内外壁之间有一个腔室。碎片可能是甲壳边缘的小角质层碎片。角质层的外部几乎光滑,有不规则的皱巴巴的多边形网或浅坑。低圆锥形丘状到高刺状,具环状结构的刺从角质层的外表面和内表面伸出。这些刺的褶皱和折叠表明角质层有适度的柔韧性。棘是中空的,较高的棘通常有顶端开口。甲壳的内表面带有较小的刺或几乎光滑。囊腔壁上有褶皱和其他结构支撑角质层的刚度。角质层内壁被多边形突起覆盖。这些大小均匀、形状均匀的肿块约1 μm大小,可能是附着在胚轴内基膜层的上皮细胞的印迹。角质层碎片的生物学亲和性尚不清楚。它们肯定是节肢动物甲壳纲动物的碎片,最有可能是囊头纲动物。相关的化石表明一个更深的海洋环境。水生植物的大量繁殖、丰富的贝壳类动物和有机磷类腕足动物,以及其他动物惊人的稀有和微小的体型,表明浅海的富营养化状况,可能是低氧的底水。具有这种角质层的动物可能是在公海中生存的。
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引用次数: 2
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Folia Musei rerum naturalium Bohemiae occidentalis. Geologica et Paleobiologica
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