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Growth pattern of the siliceous skeletons of living Spumellaria (Radiolaria) from the Kuroshio Current, offshore southwestern Shikoku Island, Japan 日本四国岛西南近海黑潮潮中活刺藻(放射菌属)硅质骨架的生长模式
IF 0.9 Q2 Earth and Planetary Sciences Pub Date : 2021-07-01 DOI: 10.1016/j.revmic.2021.100504
Rie S. Hori , Takenobu Shinki , Akihiro Iwakiri , Atsushi Matsuoka , Noritoshi Suzuki , Kaoru Ogane , Akihiro Tuji

Experimental culture studies of living radiolarians experience many difficulties because radiolarians are too sensitive to be maintained alive in artificial conditions. We here developed an improved experimental protocol in order to detect newly-formed parts of the polycystine radiolarian skeletons after they were labeled with fluorescent markers. Living samples were collected from surface seawater of the Kuroshio Current offshore from Kashiwajima Island, Kochi Prefecture on the island of Shikoku, Japan. The taxonomic composition of the Kashiwajima radiolarian samples varied markedly across the seasons and sampling sites. Spumellaria were usually abundant in samples collected near Kashiwajima, and had high diversity in shell morphology. Here, we performed experimental studies on living spumellarian radiolarians to clarify their siliceous skeletogenesis. Our culture-based research on spumellarians provided the following results.

(A) The pattern of silica deposition on the skeleton of Spongaster tetras tetras varies depending on the growth stage. Three ontogenetic stages were identified: (1) young, (2) progressively-growing and (3) fully-grown. Furthermore, novel varied types of skeletal growth modes were determined as follows: (i) rapid growth on the outermost shell with enlargement of shell size and (ii) internal deposition on skeletal components without change in shell size. From measurements of the growth rate of cultured radiolarians, it is assumed that S. tetras tetras cell has at least a month-long life cycle.

(B) Spherical spumellarians have their own shell-forming strategy that permits additional silicogenesis while maintaining the taxonomically-specific shape of their shell; the meshwork of their shell surface consists of polygonal-shaped pore-frames formed by heterogeneous patterns of added silica.

Our direct observation with fluorescence markers detected a skeletogenesis process of Haliommilla capillacea possessing a fragile spherical shell. We thus confirm the bridge-growth hypothesis for morphogenesis of the polygonal frame as proposed in previous studies. Spongosphaera streptacantha, having extremely long spines, exhibited a special growth pattern of spine elongation in a centrifugal direction, without a change in width. This was achieved by a particular temporal pattern of silicification characterized by an alternating pattern of patchy addition of new silica on the entire part of each blade from the base to the tip. Moreover, a thin layer of new silica was added by a flowing deposition along the edge of the spine. Based on these culture-based studies using living radiolarians, we conclude that Spumellaria control and maintain a particular shell shape by regulating where silicification takes place on the skeleton, particularly depending on the growth stage.

活体放射虫的实验培养研究遇到了许多困难,因为放射虫过于敏感,无法在人工条件下维持生命。我们在此开发了一种改进的实验方案,以便在荧光标记后检测多胱氨酸放射虫骨架的新形成部分。活体样本采集自日本四国岛高知县鹿岛近海黑潮表层海水。柏岛放射虫样品的分类组成在不同季节和采样地点有显著差异。在柏岛附近采集的样品中通常有大量的刺藻,并且在壳形态上具有很高的多样性。在这里,我们对活的刺虫放射虫进行了实验研究,以阐明它们的硅质骨骼形成。我们对棘球菌的培养研究得出以下结果:(A)不同生长阶段,海绵骨架上二氧化硅沉积的模式不同。确定了三个个体发生阶段:(1)幼体,(2)逐渐生长和(3)完全生长。此外,确定了新的不同类型的骨骼生长模式:(i)在最外层外壳上快速生长,外壳尺寸增大;(ii)在骨骼部件上内部沉积,外壳尺寸不变。根据对培养放射虫生长速率的测量,可以假设S. tetras tetras细胞至少有一个月的生命周期。(B)球形spumellarians有自己的壳形成策略,允许额外的硅发生,同时保持其外壳的分类特异性形状;其壳表面的网状结构由添加的非均相二氧化硅形成的多边形孔框组成。我们用荧光标记直接观察毛缕花的骨骼形成过程,发现毛缕花有一个脆弱的球形外壳。因此,我们证实了先前研究中提出的多边形框架形态发生的桥梁生长假说。Spongosphaera streptacantha具有超长的棘,棘呈离心方向延伸的特殊生长模式,而棘的宽度没有变化。这是通过一种特殊的硅化时间模式来实现的,其特征是在每个叶片的整个部分从基部到尖端上交替地添加新的二氧化硅。此外,通过沿脊柱边缘的流动沉积,增加了一层薄薄的新二氧化硅。基于这些利用活放射虫进行的基于培养的研究,我们得出结论,Spumellaria通过调节骨骼上硅化的位置来控制和维持特定的外壳形状,特别是取决于生长阶段。
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引用次数: 1
Middle to Late Paleocene larger benthic foraminifera from the Sirt Basin (Libya) 利比亚锡尔特盆地中晚古新世大型底栖有孔虫
IF 0.9 Q2 Earth and Planetary Sciences Pub Date : 2021-07-01 DOI: 10.1016/j.revmic.2021.100481
Aleš Vršič , Elżbieta Machaniec , Hans-Jürgen Gawlick

The Sirt Basin of Libya hosts significant hydrocarbon reservoirs in Paleocene shallow-marine carbonates (Upper Sabil Formation) with abundant larger benthic foraminifera. Benthic foraminifera proved to be an essential biostratigraphic tool for precise biostratigraphic dating of these carbonate platform/ramp deposits. A detailed systematic description of the Middle to Late Paleocene larger benthic foraminifera from the Sirt Basin is presented, which refines the biostratigraphic age of the Paleogene shallow-marine carbonates. Eight taxa are identified at a species level. Among them, Ranikothalia solimani is the most important taxon for the correlation of the Upper Sabil Formation. The taxa Miscellanea yvettae, Miscellanites primitivus, and Glomalveolina primaeva with co-occurrence of orthophragminid foraminifera constrain the age of the Upper Sabil Formation as Middle to Late Paleocene (Selandian-Early Thanetian).

利比亚Sirt盆地具有重要的古新世浅海碳酸盐岩储集层(上Sabil组),具有丰富的大型底栖有孔虫。底栖有孔虫被证明是对这些碳酸盐岩台地/斜坡沉积进行精确生物地层测年的重要生物地层工具。对锡尔特盆地中-晚古新世大型底栖有孔虫进行了系统详细的描述,从而细化了古近系浅海碳酸盐的生物地层年龄。在物种水平上鉴定出8个分类群。其中,Ranikothalia solimani是上萨比尔组对比最重要的分类群。杂集亚杂集亚(anea yvettae)、杂集亚(anites primitivus)、杂集亚(Glomalveolina primaeva)类群与共现的orthophragminid有孔虫类群限定上萨比勒组的时代为古新世中晚期(selantian—早Thanetian)。
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引用次数: 4
Reconstruction of oceanographic and environmental conditions in the eastern Mediterranean (Kottafi Hill section, Cyprus Island) during the middle Miocene Climate Transition 中新世中期气候过渡时期东地中海(塞浦路斯岛Kottafi Hill段)海洋环境条件的重建
IF 0.9 Q2 Earth and Planetary Sciences Pub Date : 2021-04-01 DOI: 10.1016/j.revmic.2020.100480
M. Athanasiou , M.V. Triantaphyllou , M.D. Dimiza , A. Gogou , I. Panagiotopoulos , A. Arabas , E. Skampa , K. Kouli , M. Hatzaki , E. Tsiolakis

The multi-proxy investigation of the deep-marine Kottafi Hill section (KHS), a part of the carbonate system of the Miocene Pakhna Formation, Cyprus, involved such proxies as calcareous nannofossil analysis, measurements of the oxygen and carbon isotope composition of the planktonic foraminifer Orbulina universa, and determination of the pollen and palynomorph contents, revealed the importance of these sedimentary sequences in the assessment of the impact of major global events during the middle Miocene on the regional scale. The KHS spans the 20.89–11.6 Ma time interval, during which eighteen OC-rich siltstone intercalated laminae have been deposited under warm and humid climate at 15.5–11.6 Ma. These layers can be possibly considered as the precursors of sapropelic layers mostly developed in the eastern Mediterranean Basin during Pliocene–Holocene. The global glacial events Mi3a–Mi5, traced by δ18O planktonic foraminifera records in the KHS, represent the stepwise cooling phase during the middle Miocene Climate Transition.

对塞浦路斯中新世Pakhna组碳酸盐岩体系的一部分深海Kottafi Hill剖面(KHS)进行了多代理研究,包括钙质纳米化石分析、浮游有孔虫Orbulina universa氧碳同位素组成测量、花粉和孢粉含量测定等。揭示了这些沉积序列在评估中新世中期重大全球事件在区域尺度上的影响方面的重要性。KHS跨度为20.89 ~ 11.6 Ma,在15.5 ~ 11.6 Ma温暖湿润的气候条件下,沉积了18条富含oc的粉砂岩夹层。这些层可能是上新世-全新世期间主要发育于东地中海盆地的腐泥层的前身。根据KHS的δ18O浮游有孔虫记录,全球冰川事件mi3 - mi5代表了中新世中期气候转变的逐步冷却阶段。
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引用次数: 4
Benthic foraminiferal assemblages and biostratigraphy of the Antalo Limestone, Mekelle Basin, Northern Ethiopia 埃塞俄比亚北部Mekelle盆地Antalo灰岩底栖有孔虫组合与生物地层学
IF 0.9 Q2 Earth and Planetary Sciences Pub Date : 2021-04-01 DOI: 10.1016/j.revmic.2020.100479
Damenu Adefris , M.E. Nton , O.A. Boboye , Balemwal Atnafu

This study focused on the identification, detailed systematic descriptions and stratigraphical distributions of different foraminifera and other species to make the first systematic paleontological analysis, interpret paleoenvironment and determine the age of Antalo Limestone of the Mekelle Basin. Fieldwork and section logging, with sample collection at seven outcrop locations, followed by petrography and standard micropaleontological analysis were carried out. Abundant foraminifera fossils of 17 species were identified from 42 thin sections and 5 washed samples of marl and shale. The faunal assemblage also comprised dasyclad green algae, calpionellid, brachiopods and stromatoporoids (Cladocoropsis mirabilis). The benthic foraminiferal assemblage is represented by 3 species of the order Lituolida (Freixialina planispiralis, Nautiloculina circularis and N. oolithica), 11 species of the order Loftusiida (Alveosepta jaccardi, Choffatella tingitana, Conicokurnubia orbitoliniformis, Everticyclammina virguliana, Kurnubia jurassica, K. morrisi, K. palastiniensis, K. wellingsi, Praekurnubia crusei, Pseudocyclammina lituus and Siphovalvulina variabilis), 2 species of the order Textulariida (Redmondoides lugeoni and Valvulina lugeoni) and 1 species of the order Miliolida (Miliolinella quinquangula). The green algae comprised 3 species of the order Dasycladales (Clypeina jurassica, Salpingoporella annulata and Salpingoporella gr. pygmaea). The calpionellid is represented by 1 species of the order Calpionellidae (Calpionella alpina). The brachiopods comprised 2 species of the orders Rhynchonellida (Somalirhynchia africana) and Terebratulida (Somalithyris bihendulensis). Based on the stratigraphical distributions of these benthic foraminifera, dasyclad green algae, calpionellid and brachiopods, three zones namely; Kurnubia palastiniensis, Somalirhynchia africana/Somalithyris bihendulensis and Alveosepta jaccardi/Pseudocyclammina lituus have been established, signifying a Callovian – Oxfordian, a Callovian – Early Kimmeridgian and a Kimmeridgian – Tithonian ages respectively. Therefore, a Callovian – Tithonian age has been assigned to the Antalo Limestone. This faunal assemblage indicated a shallow – deep marine environment with normal marine salinity which has a strong affinity with the faunas of the Middle – Upper Jurassic Peri-Tethyan carbonates, indicating closely related geological evolution. Six benthic foraminiferal morphogroups have been identified, indicating epifaunal – infaunal microhabitats and active deposit-fe

本研究重点对不同有孔虫及其他物种进行了鉴定、详细系统描述和地层分布,首次进行了系统的古生物分析,解释了古环境,确定了Mekelle盆地Antalo灰岩的年龄。野外工作和剖面测井,在7个露头位置采集样本,然后进行岩石学和标准微古生物学分析。从泥灰岩和页岩的42个薄片和5个水洗样品中鉴定出了17种丰富的有孔虫化石。动物群中还包括绿甲藻、卡贝类、腕足类和层孔类(神奇枝孢虫)。底栖有孔虫群落中有3种为Lituolida目(Freixialina planispiralis、Nautiloculina circularis和N. oolithica), 11种为Loftusiida目(Alveosepta jaccardi、Choffatella tingitana、Conicokurnubia orbitoliniformis、Everticyclammina virguliana、Kurnubia jurassica、K. morrisi、K. palastiniensis、K. wellingsi、Praekurnubia crusei、pseudoyclammina lituus和Siphovalvulina variabilis);毛茛目2种(红毛茛属和花毛茛属)和千毛蕨目1种(千毛蕨属)。绿藻由3种绿藻类组成,分别为:朱绿藻、环绿藻和小绿藻。calpionellides (calpionellae alpina)以calpionellae科1种为代表。腕足动物包括非洲索马利亚目(somaliirhynchia africana)和有翅目(Somalithyris bihendulensis) 2种。根据底栖有孔虫、底栖绿藻、底栖甲壳类和腕足类的地层分布,划分为三个带:已经建立了Kurnubia palastiniensis、Somalirhynchia africana/Somalithyris bihendulensis和Alveosepta jaccardi/Pseudocyclammina lituus,分别代表了Callovian - Oxfordian、Callovian - Early kimmeridian和kimmeridian - Tithonian时代。因此,安塔洛灰岩被确定为卡洛—提通尼亚时代。该动物组合显示了一个正常盐度的浅海-深海环境,与中-上侏罗统近特提斯碳酸盐岩动物群有很强的亲缘关系,表明地质演化密切相关。已经确定了六个底栖有孔虫形态群,表明了底栖-底栖微生境和活跃的沉积物摄食、营养性、草食性、杂食性和细菌清食的摄食策略。
{"title":"Benthic foraminiferal assemblages and biostratigraphy of the Antalo Limestone, Mekelle Basin, Northern Ethiopia","authors":"Damenu Adefris ,&nbsp;M.E. Nton ,&nbsp;O.A. Boboye ,&nbsp;Balemwal Atnafu","doi":"10.1016/j.revmic.2020.100479","DOIUrl":"https://doi.org/10.1016/j.revmic.2020.100479","url":null,"abstract":"<div><p><span>This study focused on the identification, detailed systematic descriptions and stratigraphical distributions of different foraminifera and other species to make the first systematic paleontological analysis, interpret paleoenvironment<span><span> and determine the age of Antalo Limestone of the Mekelle Basin. Fieldwork and section logging, with sample collection at seven outcrop locations, followed by </span>petrography<span> and standard micropaleontological analysis were carried out. Abundant foraminifera fossils<span><span> of 17 species were identified from 42 thin sections and 5 washed samples of marl and shale. The faunal assemblage also comprised dasyclad green algae, </span>calpionellid, brachiopods and stromatoporoids (</span></span></span></span><em>Cladocoropsis mirabilis</em>). The benthic foraminiferal assemblage is represented by 3 species of the order Lituolida (<em>Freixialina planispiralis</em>, <em>Nautiloculina circularis</em> and <em>N</em>. <em>oolithica</em>), 11 species of the order Loftusiida (<em>Alveosepta jaccardi</em>, <em>Choffatella tingitana</em>, <em>Conicokurnubia orbitoliniformis</em>, <em>Everticyclammina virguliana</em>, <em>Kurnubia jurassica</em>, <em>K</em>. <em>morrisi, K</em>. <em>palastiniensis, K</em>. <em>wellingsi</em>, <em>Praekurnubia crusei</em>, <em>Pseudocyclammina lituus</em> and <em>Siphovalvulina variabilis</em>), 2 species of the order Textulariida (<em>Redmondoides lugeoni</em> and <em>Valvulina lugeoni</em>) and 1 species of the order Miliolida (<em>Miliolinella quinquangula</em>). The green algae comprised 3 species of the order Dasycladales (<em>Clypeina jurassica</em>, <em>Salpingoporella annulata</em> and <em>Salpingoporella</em> gr. <em>pygmaea</em>). The calpionellid is represented by 1 species of the order Calpionellidae (<em>Calpionella alpina</em>). The brachiopods comprised 2 species of the orders Rhynchonellida (<em>Somalirhynchia africana</em>) and Terebratulida (<em>Somalithyris bihendulensis</em><span>). Based on the stratigraphical distributions of these benthic foraminifera, dasyclad green algae, calpionellid and brachiopods, three zones namely; </span><em>Kurnubia palastiniensis</em>, <em>Somalirhynchia africana</em>/<em>Somalithyris bihendulensis</em> and <em>Alveosepta jaccardi</em>/<em>Pseudocyclammina lituus</em><span><span><span> have been established, signifying a Callovian<span> – Oxfordian, a Callovian – Early Kimmeridgian<span> and a Kimmeridgian – Tithonian ages respectively. Therefore, a Callovian – Tithonian age has been assigned to the Antalo Limestone. This faunal assemblage indicated a shallow – deep marine environment with normal marine </span></span></span>salinity which has a strong affinity with the faunas of the Middle – Upper Jurassic Peri-Tethyan carbonates, indicating closely related geological evolution. Six benthic foraminiferal morphogroups have been identified, indicating epifaunal – infaunal </span>microhabitats and active deposit-fe","PeriodicalId":45442,"journal":{"name":"REVUE DE MICROPALEONTOLOGIE","volume":null,"pages":null},"PeriodicalIF":0.9,"publicationDate":"2021-04-01","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"https://sci-hub-pdf.com/10.1016/j.revmic.2020.100479","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"91618023","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":0,"RegionCategory":"","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
引用次数: 1
VSI “Updating Micropaleontology and Stratigraphy in Greece and Eastern Mediterranean” of the Hellenic Committee for Paleontology and Stratigraphy in the honor of Professor Emeritus Michael D. Dermitzakis 希腊古生物学和地层学委员会的VSI“更新希腊和东地中海的微古生物学和地层学”,以纪念名誉教授Michael D. Dermitzakis
IF 0.9 Q2 Earth and Planetary Sciences Pub Date : 2021-04-01 DOI: 10.1016/j.revmic.2021.100484
Maria Triantaphyllou, Margarita Dimiza

The current Special Issue of Revue de Micropaleontologie, entitled “Updating Micropaleontology and Stratigraphy in Greece and Eastern Mediterranean” includes contributions from the T1S1 special Session [conveners Maria Triantaphyllou (NKUA), Dimitris Kostopoulos (AUTH), George Iliopoulos (UPAT)] organized by the Hellenic Committee for Paleontology and Stratigraphy at the 15th Conference of the Geological Society of Greece (May 2019, Athens, Greece).

本期《微古生物学评论》特刊题为“更新希腊和东地中海的微古生物学和地层学”,其中包括希腊古生物学和地层学委员会在希腊地质学会第15届会议(2019年5月,希腊雅典)上组织的T1S1特别会议[召集人Maria Triantaphyllou (NKUA), Dimitris Kostopoulos (AUTH), George Iliopoulos (UPAT)]的贡献。
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引用次数: 0
Micropalaeontological analysis and palaeoenvironmental interpretation of the upper sedimentary sequence of Corinth Marl (Corinth Isthmus, Greece) 希腊科林斯地峡科林斯泥沼上部沉积层序的微体古生物学分析及古环境解释
IF 0.9 Q2 Earth and Planetary Sciences Pub Date : 2021-04-01 DOI: 10.1016/j.revmic.2020.100466
Theodora Tsourou , Maria V. Triantaphyllou , Sophia Cheilaris , Eugenia G. Fatourou , Ioannis Michailidis , Anastasios P. Nikitas , Maria A. Tzortzopoulou , Margarita Dimiza , Elizabeth Stathopoulou

The current study performs a detailed micropalaeontological analysis, in terms of ostracods, benthic foraminifera and calcareous nannoplankton, in order to reconstruct the palaeoenvironmental history of the sedimentary sequence exposed in the central part of Corinth Isthmus. Twenty one samples from two lateral sections were studied, on both sides of a westwards dipping normal fault at the west end of the central horst, which are part of the same stratigraphical sequence. The calcareous nannofossil analysis provided for the first time dating evidence for the upper Corinth Marl sequence, revealing that Section A (at the footwall block of the fault/the upper part of the Corinth Marl) is attributed to the Early Pleistocene /Calabrian (nannofossil biozone MNN19f) and that Section B (at the hanging wall of the fault) is assigned to the Middle Pleistocene /Chibanian (MNN20). Both parts of the studied sequence not only differentiate in age but bear notably separated species composition of the main ostracod assemblages as well. The combined study of ostracod and benthic foraminiferal microfaunas indicated that at least since Early Pleistocene the Corinth Marl deposits at the central part of Corinth Isthmus corresponded to a shallow, restricted, oligohaline lagoon with salinity alternations expressed as layers enriched in nannofossil content. Towards Middle Pleistocene the lagoon presented increased salinity and signs of shoaling. The studied sequence closes upwards with a bed of Middle Pleistocene age, rich in marine molluscs and corals indicating a radical environmental change: the opening of the lagoon and the establishment of a shallow marine coastal environment.

本研究从介形类、底栖有孔虫和钙质纳米浮游生物等方面进行了详细的微体古生物学分析,以重建科林斯地峡中部暴露的沉积序列的古环境历史。在同一层序中,研究了位于中央断层西端向西倾斜的正断层两侧的两个侧向剖面的21个样品。钙质纳米化石分析首次为上哥林泥沼层序提供了定年证据,表明A段(断层下盘/哥林泥沼上部)属于早更新世/卡拉布良(纳米化石生物带MNN19f), B段(断层上盘)属于中更新世/Chibanian (MNN20)。研究序列的两个部分不仅在年龄上存在差异,而且主要介形虫组合的物种组成也明显不同。对介形虫和底栖有孔虫微动物群的综合研究表明,至少自早更新世以来,科林斯地峡中部的科林斯泥灰岩沉积物对应于一个浅的、受限制的低盐泻湖,其盐度变化表现为富含纳米化石含量的层。到中更新世时,泻湖呈现盐度增加和浅滩化迹象。所研究的层序向上闭合于中更新世地层,富含海洋软体动物和珊瑚,表明环境发生了彻底的变化:泻湖的开放和浅海海岸环境的建立。
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引用次数: 0
On the status of the radiolarian genus Spongoliva Haeckel, 1887 and the description of the genus Spongolivella n. gen 海克尔(Haeckel, 1887)关于海绵虫属的现状和海绵虫属的描述
IF 0.9 Q2 Earth and Planetary Sciences Pub Date : 2021-04-01 DOI: 10.1016/j.revmic.2020.100477
Paulian Dumitrică

The author erects a new genus for the Quaternary artiscin radiolarian species Spongoliva ellipsoides Popofsky 1912. The genus Spongoliva Haeckel 1887 is a nomen dubium because none of its six species has been illustrated and, above all, its type species is Eocene in age, an epoch when the artiscins had not yet appeared.

作者为第四纪人工放射虫种海绵虫(sponoliva ellipsoides)建立了一个新属(Popofsky, 1912)。海克尔1887海绵藻属是一种未发现的dubium,因为它的六个物种中没有一个被描述过,最重要的是,它的模式物种是始新世,一个艺术品尚未出现的时代。
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引用次数: 1
A preliminary investigation of Eocene larger benthic foraminifera assemblages from Alpine and molasse-type deposits of the Hellenic peninsula (Greece) 希腊半岛阿尔卑斯和糖蜜型沉积物始新世大型底栖有孔虫组合的初步调查
IF 0.9 Q2 Earth and Planetary Sciences Pub Date : 2021-04-01 DOI: 10.1016/j.revmic.2020.100468
Vasiliki-Grigoria Dimou , Olga Koukousioura , Margarita D. Dimiza , Maria V. Triantaphyllou , György Less , Fotini Pomoni-Papaioannou , Georgios Syrides

Larger Benthic Foraminifera (LBF) from five localities in Greece were studied in order to define the morphological features of various nummulitid forms and to describe the foraminiferal assemblages in different geotectonic environments. The selected sites belong to molasse-type formations in Kirki/Thrace Basin and Vasiliki Trikala/Mesohellenic Trough, and carbonate deposits in Alpine Pindos, Ionian and Tripolis geotectonic units.

Taxonomic studies combined with biometric data, when possible, enabled us to identify different morphotypes. In the molasse-type deposits of Thrace Basin (Kirki Thrace), N. perforatus was found along with N. maximus and Assilina exponens suggesting correlation with biozone SBZ 16/17, while in the Mesohellenic Trough deposits (Vasiliki Trikala) the association included N. perforatus group, Operculina gomezi group and Silvestriella tetraedra, indicating a Bartonian age, SBZ 17-18A. In outcrops found within Pindos unit (Perivoli Grevena) three different assemblages have been observed in the microbreccious olistholiths, suggesting three distinguished deposits forming quasi-continuous succession spanning from early to middle and up to late Eocene. The Ionian unit (Manoliasa Ioannina) presented a mixed redeposited fauna including late Eocene, early Paleogene and planktonic foraminifera clasts. Finally, the Tripolis unit (Dervenakia and Tripolis) association was interpreted as Bartonian in age, defined by the presence of Operculina gomezi group and N. perforatus group.

本文对希腊5个地区的大型底栖有孔虫(large Benthic Foraminifera, LBF)进行了研究,以确定其各种数量多的形态特征,并描述其在不同大地构造环境下的有孔虫组合。选择的地点属于Kirki/Thrace盆地和Vasiliki Trikala/中希腊海槽的糖蜜型地层,以及Alpine Pindos、Ionian和Tripolis大地构造单元的碳酸盐矿床。在可能的情况下,分类学研究结合生物特征数据使我们能够识别不同的形态。在色雷斯盆地(Kirki Thrace)的蜜蜜型矿床中,N. perforatus与N. maximus和Assilina指数相结合,表明其与SBZ 16/17生物带有关;在中希腊海槽(Vasiliki Trikala)矿床中,N. perforatus群、Operculina gomez群和Silvestriella tetraedra相结合,表明其属于巴氏期(SBZ 17-18A)。在Pindos单元(Perivoli Grevena)的露头中,在微角砾岩中观察到三种不同的组合,表明三个不同的矿床形成了早至中、晚始新世的准连续演替。伊奥尼亚单元(Manoliasa Ioannina)是一个由始新世晚期、古近世早期和浮游有孔虫碎屑组成的混合再沉积动物群。最后,Tripolis组(Dervenakia和Tripolis)组合在年龄上被解释为Bartonian,由Operculina gomzi组和N. perforatus组的存在来定义。
{"title":"A preliminary investigation of Eocene larger benthic foraminifera assemblages from Alpine and molasse-type deposits of the Hellenic peninsula (Greece)","authors":"Vasiliki-Grigoria Dimou ,&nbsp;Olga Koukousioura ,&nbsp;Margarita D. Dimiza ,&nbsp;Maria V. Triantaphyllou ,&nbsp;György Less ,&nbsp;Fotini Pomoni-Papaioannou ,&nbsp;Georgios Syrides","doi":"10.1016/j.revmic.2020.100468","DOIUrl":"https://doi.org/10.1016/j.revmic.2020.100468","url":null,"abstract":"<div><p>Larger Benthic Foraminifera (LBF) from five localities in Greece were studied in order to define the morphological features of various nummulitid forms and to describe the foraminiferal assemblages in different geotectonic environments. The selected sites belong to molasse-type formations in Kirki/Thrace Basin and Vasiliki Trikala/Mesohellenic Trough, and carbonate deposits in Alpine Pindos, Ionian and Tripolis geotectonic units.</p><p><span>Taxonomic studies combined with biometric data, when possible, enabled us to identify different morphotypes. In the molasse-type deposits of Thrace Basin (Kirki Thrace), </span><em>N. perforatus</em> was found along with <em>N. maximus</em> and <em>Assilina exponens</em> suggesting correlation with biozone SBZ 16/17, while in the Mesohellenic Trough deposits (Vasiliki Trikala) the association included <em>N. perforatus</em> group, <em>Operculina gomezi</em> group and <em>Silvestriella tetraedra</em><span><span>, indicating a Bartonian age, SBZ 17-18A. In outcrops found within Pindos unit (Perivoli Grevena) three different assemblages have been observed in the microbreccious olistholiths, suggesting three distinguished deposits forming quasi-continuous succession spanning from early to middle and up to late Eocene. The Ionian unit (Manoliasa Ioannina) presented a mixed redeposited fauna including late Eocene, early </span>Paleogene and planktonic foraminifera clasts. Finally, the Tripolis unit (Dervenakia and Tripolis) association was interpreted as Bartonian in age, defined by the presence of </span><em>Operculina gomezi</em> group and <em>N. perforatus</em> group.</p></div>","PeriodicalId":45442,"journal":{"name":"REVUE DE MICROPALEONTOLOGIE","volume":null,"pages":null},"PeriodicalIF":0.9,"publicationDate":"2021-04-01","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"https://sci-hub-pdf.com/10.1016/j.revmic.2020.100468","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"89997603","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":0,"RegionCategory":"","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
引用次数: 2
The Cenomanian–Turonian boundary event in Egypt: Foraminiferal turnover and carbon isotopic change 埃及的塞诺曼尼亚-土鲁尼亚边界事件:有孔虫转换和碳同位素变化
IF 0.9 Q2 Earth and Planetary Sciences Pub Date : 2020-12-01 DOI: 10.1016/j.revmic.2020.100463
Sherif M. El Baz , Ahmed A. Kassem

This study deals with the integration between geochemical data (carbon isotope and, total organic carbon) and foraminiferal turnover to track the Oceanic Anoxic Event 2 (OAE2) from the OCT-A8st1 well, which located in October field, central part of Gulf of Suez, Egypt. The studied well is subdivided into two formations, namely the Galala (Early–Late Cenomanian) and Abu Qada (Late Cenomanian–Early Turonian). Concerning the foraminiferal assemblage, agglutinated foraminifera dominate the Galala Formation, while planktonic taxa only occur within a few levels in the Abu Qada Formation. Due to the absence of keeled planktonic forms, the biostratigraphic framework is established on the integration between benthonic and planktonic foraminifera. Three biozones are recognized, including Thomasinella aegyptia–Merlingina cretacea Interval Zone (Early–Middle Cenomanian), Thomasinella fragmentaria–Praebulimina aspera Interval Zone (Middle–Late Cenomanian), and Whiteinella archaeocretacea Taxon Range Zone (Late Cenomanian–Early Turonian). Therefore, the Cenomanian–Turonian boundary interval (CTBI) is placed within the lower part of the Abu Qada Formation, which is characterized by the presence of black shales and low richness of both foraminifera and ostracods. A remarkable increase in TOC values is observed close to the CTBI. Also, the carbon isotope curve shows a distinctive positive δ13C anomaly at the same level of the maximum TOC. The comparison between the recorded δ13C data and those obtained from other sections worldwide, indicates that the base of the Turonian is placed directly above the δ13C excursion. Furthermore, the recorded foraminiferal community shows evidence for the deepening of the environment and extension of the oxygen minimum zone during the CTBI. This conclusion is based on a sequence of bioevents starting with the disappearance of some agglutinated benthic foraminifera, followed by the dominance of opportunistic groups including buliminids, whiteinellids and heterohelicids. Consequently, the studied well may be considered as a local reference section for OAE2.

本文利用地球化学数据(碳同位素和总有机碳)与有孔虫周转量相结合的方法,对埃及苏伊西湾中部十月油田OCT-A8st1井的海洋缺氧事件2 (OAE2)进行了跟踪研究。研究井分为两个组,即Galala(早-晚Cenomanian)和Abu Qada(晚Cenomanian -早Turonian)。在有孔虫组合中,Galala组以凝集有孔虫为主,而Abu Qada组的浮游类群仅在少数层位内出现。由于缺乏龙骨浮游生物形式,生物地层格架建立在底栖和浮游有孔虫之间的整合上。确定了3个生物带,分别为:埃及Thomasinella aegyptia-Merlingina白垩纪间隔带(早-中塞诺曼世)、破碎Thomasinella fragmentaria-Praebulimina aspera间隔带(中晚期塞诺曼世)和怀特inella古白垩纪分类群区间带(晚塞诺曼世-早Turonian)。因此,塞诺曼—turonian边界段位于Abu Qada组下部,其特征是黑色页岩的存在,有孔虫和介形虫的丰富度都很低。TOC值在CTBI附近显著增加。碳同位素曲线在最大TOC的同一水平上也表现出明显的正δ13C异常。对比记录的δ13C值与世界上其他剖面的δ13C值,发现Turonian的基底位于δ13C偏移的正上方。此外,有孔虫群落的记录显示了在CTBI期间环境的加深和氧最小带的扩展。这一结论是基于一系列的生物事件,从一些凝集的底栖有孔虫消失开始,然后是机会主义群体的优势,包括buliminids, whiteinellids和heterohelicids。因此,该井可作为OAE2的局部参考剖面。
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引用次数: 7
Seasonal living coccolithophore distribution in the enclosed coastal environments of the Thessaloniki Bay (Thermaikos Gulf, NW Aegean Sea) 塞萨洛尼基湾(Thermaikos Gulf, NW爱琴海)封闭海岸环境中季节性活球石藻的分布
IF 0.9 Q2 Earth and Planetary Sciences Pub Date : 2020-12-01 DOI: 10.1016/j.revmic.2020.100449
Dimiza M.D. , Koukousioura O. , Michailidis I. , Dimou V.-G. , Navrozidou V. , Aligizaki K. , Seferlis M.

Living coccolithophores collected from a yearly time interval, January to December 2016, at the SP1 station of the Thessaloniki Bay, an enclosed coastal environment located in the inner part of the Thermaikos Gulf (Northwestern Aegean Sea) were analyzed to determine their seasonal variation and evaluate the effects of environmental factors on their density, species composition and distribution. Our results showed that coccolithophores constituted a significant component of the phytoplankton community and were characterized by the dominance of few opportunistic species, as an impact of eutrophic environmental conditions. Temperature seasonal cycle and the variations in nutrient inputs from riverine and/or anthropogenic sources seem to control the seasonal pattern of coccolithophores. Emiliania huxleyi with densities as high as ∼4.0 × 105 cells l−1 predominated during the late spring–early summer, whereas a strong dominance of Gephyrocapsa oceanica with densities ∼2.0 × 105 cells l−1 was observed during the late summer and the autumn. Emiliania huxleyi appeared to dominate the coccolithophore assemblages once more during the winter; while Helicosphaera carteri was continuously present during the wintertime (exceeding ∼2.7 × 103 cells l−1). Emiliania huxleyi was entirely represented by morphotype type A in the studied samples. Morphometric analysis of E. huxleyi coccoliths indicated that the population is dominated by lightly calcified forms, although an increase of more heavily calcified coccoliths was observed during late autumn-early spring in line with the well-documented seasonal succession pattern of E. huxleyi type A forms in the Aegean Sea.

对2016年1 - 12月在爱琴海西北部塞萨洛尼基湾(Thessaloniki Bay) SP1站采集的活粒石藻进行了季节变化分析,并评价了环境因子对其密度、种类组成和分布的影响。结果表明,在富营养化环境条件的影响下,球石藻是浮游植物群落的重要组成部分,具有少数机会性物种占优势的特点。温度季节循环和来自河流和/或人为来源的养分输入的变化似乎控制着球石藻的季节模式。在春末夏初,密度高达~ 4.0 × 105 cells l - 1的赫胥黎Emiliania huxleyi占优势,而在夏末和秋季,密度为~ 2.0 × 105 cells l - 1的Gephyrocapsa oceanica占优势。在冬季,赫胥黎Emiliania huxleyi似乎再次主导了球石藻组合;而卡特螺旋藻(Helicosphaera carteri)在冬季持续存在(超过2.7 × 103细胞l−1)。在所研究的样品中,赫胥黎菌完全以A型形态型为代表。对huxleyi型球粒岩的形态分析表明,该种群以轻度钙化形式为主,尽管在深秋至早春期间观察到重度钙化球粒岩的增加,这与爱琴海中huxleyi型A型球粒岩的季节性演替模式一致。
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引用次数: 6
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