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CHALLENGES IN ORBITOLINID IDENTIFICATION AND BIOSTRATIGRAPHIC CONSEQUENCES: AN APTIAN EXAMPLE 眶藻类鉴定的挑战和生物地层学的后果:一个始新世的例子
Pub Date : 2024-03-14 DOI: 10.35463/j.apr.2024.02.02
Felix Schlagintweit
The much-cited importance of Orbitolinidae in biostratigraphy depends on the species and time-interval under consideration. In any case, correct species identification is of primary importance as the basis for drawing conclusions. This task is best achieved in an assemblage where centered sections (axial and transverse) showing all details of the complex megalospheric embryo are available. Difficulties in the correct interpretation of oblique sections is exemplified by an example from the Aptian of Turkey, respectively Praeorbitolina versus Mesorbitolina. Resulting differences in species determination not only has an impact on age interpretation, but also associated geologic-tectonic conclusions, as well as basic taxonomic data (e.g., stratigraphic ranges, statistics on stage diversities).
Orbitolinidae 在生物地层学中的重要性被广泛提及,这取决于所考虑的物种和时间段。在任何情况下,正确的物种鉴定都是得出结论的重要依据。要完成这项任务,最好是能获得显示复杂巨型胚胎所有细节的中心切片(轴向和横向)。正确解释斜切片的困难体现在土耳其始新世的一个例子中,分别是 Praeorbitolina 和 Mesorbitolina。由此导致的物种判定差异不仅会影响年龄解释,还会影响相关的地质构造结论以及基本的分类数据(如地层范围、阶段多样性统计)。
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引用次数: 0
BELOPTERA LONGA NAEF, 1922 (CEPHALOPODA: COLEOIDEA) FROM THE LATE EOCENE (PALAEOGENE) OF UKRAINE Beloptera longa naef, 1922 (cephalopoda: coleoidea) from the late eocene (palaeogene) of ukraine 克拉科夫晚始新世(古近纪)的 Beloptera longa naef, 1922 (cephalopoda: coleoidea)
Pub Date : 2024-03-11 DOI: 10.35463//j.apr.2024.02.01
V. Dernov
Beloptera longa Naef, 1922 and Beloptera aff. longa Naef, 1922 have been described from the Priabonian Mandrykivka Beds of the Rybalsky Quarry Konzentrat-Lagerstätte in addition to previously known Beloptera belemnoidea de Blainville, 1825, B. cf. longa Naef, 1922, Belosaepia blainvillei (Deshayes, 1835) and Vasseuria occidentalis Munier-Chalmas, 1880. The geographical and stratigraphic distribution of the species Beloptera longa, which was previously recorded only in the Bartonian of England, has been expanded. Cephalopod assemblages from the Priabonian of Ukrainian consist of the genera Beloptera, Vasseuria, Belosaepia, Aturia and Eutrephoceras.
除了之前已知的 Beloptera belemnoidea de Blainville, 1825、B. cf. longa Naef, 1922、Belosaepia blainvillei (Deshayes, 1835) 和 Vasseuria occidentalis Munier-Chalmas, 1880 外,还描述了来自 Rybalsky Quarry Konzentrat-Lagerstätte 的 Priabonian Mandrykivka 床的 Beloptera longa Naef, 1922 和 Beloptera aff.Beloptera longa 这一物种以前仅在英格兰的巴顿期有记录,其地理分布和地层分布已得到扩展。乌克兰普里阿本纪的头足类组合包括 Beloptera 属、Vasseuria 属、Belosaepia 属、Aturia 属和 Eutrephoceras 属。
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引用次数: 0
GEOMETRIC MORPHOMETRIC APPROACH TO ESTABLISH PHYLOGENETIC AFFINITIES OF ENIGMATIC PTEROSAUR SPECIMENS FROM THE LOWER CRETACEOUS OF SOUTH KOREA 用几何形态计量学方法确定韩国下白垩统神秘翼龙标本的系统发育亲缘关系
Pub Date : 2024-02-11 DOI: 10.35463/j.apr.2024.01.06
Chan-gyu Yun
Fragmentary but scientifically important pterosaur fossils have been reported in Lower Cretaceous strata such as the Jinju and Hasandong formations of South Korea. Recently, several large teeth and a second wing phalanx found in these strata have been suggested to have affinities with Boreopteridae, a clade in which has so far been reported only in the Yixian Formation of China. Here, the phylogenetic affinities of these proposed boreopterid specimens are reassessed, using two-dimensional geometric morphometric shape analyses. The morphometric analyses found the South Korean pterosaur teeth in different morphospaces from the Yixian Fm. boreopterids, and plot them closer to anhanguerians. An isolated second wing phalanx does plot close to a boreopterid Zhenyuanopterus longirostris in a morphospace defined by first two principal component axes, but taxa from different lineages are also plotted as closely, or even closer. Thus, there seems to be no clear evidence that these specimens belong to Boreopteridae, and the tooth fossils appear to be very different from those of established members of this clade. Therefore, the hypothesis that some South Korean pterosaur fossils can be referred to boreopterids, is here tentatively rejected.
在下白垩统地层(如韩国的晋州地层和哈山东地层)中发现了一些零碎但具有重要科学价值的翼龙化石。最近,在这些地层中发现的几颗大牙齿和第二翼趾骨被认为与蝶龙科(Boreopteridae)有亲缘关系。在此,我们利用二维几何形态分析重新评估了这些拟议的膛翅目标本的系统发育亲缘关系。形态分析发现,南朝鲜翼龙的牙齿与黟县地层的镗翼龙牙齿处于不同的形态空间,并将它们与安汉古龙类划定为更接近的类群。在由前两个主成分轴定义的形态空间中,一个孤立的第二翼趾骨确实与北缘翼龙(Zhenyuanopterus longirostris)接近,但来自不同类群的类群也与之接近,甚至更接近。因此,似乎没有明确的证据表明这些标本属于伯劳蝶科,而且这些牙齿化石似乎与该支系已有成员的牙齿化石非常不同。因此,南韩的一些翼龙化石可归属于膛翅目的假说在此被暂时否定。
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引用次数: 0
GEOMETRIC MORPHOMETRIC APPROACH TO ESTABLISH PHYLOGENETIC AFFINITIES OF ENIGMATIC PTEROSAUR SPECIMENS FROM THE LOWER CRETACEOUS OF SOUTH KOREA 用几何形态计量学方法确定韩国下白垩统神秘翼龙标本的系统发育亲缘关系
Pub Date : 2024-02-11 DOI: 10.35463/j.apr.2024.01.06
Chan-gyu Yun
Fragmentary but scientifically important pterosaur fossils have been reported in Lower Cretaceous strata such as the Jinju and Hasandong formations of South Korea. Recently, several large teeth and a second wing phalanx found in these strata have been suggested to have affinities with Boreopteridae, a clade in which has so far been reported only in the Yixian Formation of China. Here, the phylogenetic affinities of these proposed boreopterid specimens are reassessed, using two-dimensional geometric morphometric shape analyses. The morphometric analyses found the South Korean pterosaur teeth in different morphospaces from the Yixian Fm. boreopterids, and plot them closer to anhanguerians. An isolated second wing phalanx does plot close to a boreopterid Zhenyuanopterus longirostris in a morphospace defined by first two principal component axes, but taxa from different lineages are also plotted as closely, or even closer. Thus, there seems to be no clear evidence that these specimens belong to Boreopteridae, and the tooth fossils appear to be very different from those of established members of this clade. Therefore, the hypothesis that some South Korean pterosaur fossils can be referred to boreopterids, is here tentatively rejected.
在下白垩统地层(如韩国的晋州地层和哈山东地层)中发现了一些零碎但具有重要科学价值的翼龙化石。最近,在这些地层中发现的几颗大牙齿和第二翼趾骨被认为与蝶龙科(Boreopteridae)有亲缘关系。在此,我们利用二维几何形态分析重新评估了这些拟议的膛翅目标本的系统发育亲缘关系。形态分析发现,南朝鲜翼龙的牙齿与黟县地层的镗翼龙牙齿处于不同的形态空间,并将它们与安汉古龙类划定为更接近的类群。在由前两个主成分轴定义的形态空间中,一个孤立的第二翼趾骨确实与北缘翼龙(Zhenyuanopterus longirostris)接近,但来自不同类群的类群也与之接近,甚至更接近。因此,似乎没有明确的证据表明这些标本属于伯劳蝶科,而且这些牙齿化石似乎与该支系已有成员的牙齿化石非常不同。因此,南韩的一些翼龙化石可归属于膛翅目的假说在此被暂时否定。
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引用次数: 0
NEW MIDDLE TRIASSIC BELL-SHAPED NASSELLARIAN RADIOLARIA FROM ALPINE AND CARPATHIAN AREAS 高山和阡陌地区新发现的中三叠世钟状纳塞尔放射虫
Pub Date : 2024-02-11 DOI: 10.35463/j.apr.2024.01.05
P. Dumitrică
This paper is a continuation of a previous one in which I described and revised different Middle Triassic nassellarian Radiolaria from the Alps and Carpathians illustrated with drawings. On the present paper, are described two new groups of Middle Triassic dicyrtid Nassellaria with bell-shaped shell from the same area to enrich the inventory of these microfossils. Two new families (Gorispelidae and Humerocyrtiidae), five new genera (Silicotintinnabulum, Colpotrelus, Gorispela, Pozsvartia and Humerocyrtis) and 30 new species are described from these genera and illustrated with drawings.
这篇论文是前一篇论文的延续,在前一篇论文中,我描述并修订了阿尔卑斯山和喀尔巴阡山脉的不同中三叠世钟形放射虫,并附图说明。本文描述了同一地区出土的两组新的中三叠世钟形壳双壳类纳西利亚虫,以丰富这些微化石的清单。本文描述了这些属中的两个新科(Gorispelidae 和 Humerocyrtiidae)、五个新属(Silicotintinnabulum、Colpotrelus、Gorispela、Pozsvartia 和 Humerocyrtis)和 30 个新种,并附有插图。
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引用次数: 0
PALYNOSTRATIGRAPHIC AND PALAEOENVIRONMENTAL INVESTIGATIONS OF THE MAASTRICHTIAN FROM OARDA DE JOS (SOUTHWESTERN TRANSYLVANIAN BASIN) 奥尔达-德约斯(西南特兰西瓦尼亚盆地)马斯特里赫特古地层的古地层学和古环境研究
Pub Date : 2024-02-11 DOI: 10.35463/j.apr.2024.01.07
D. Țabără, Z. Csiki-Sava
A palynological, palynofacies and organic geochemical investigation of the Upper Cretaceous continental vertebrate site of Oarda de Jos allowed us to constrain its age and to make more detailed inferences about its depositional environment. The recovered palynomorph assemblage is represented mainly by angiosperm pollen and fern spores, alongside rare taxa of gymnosperms. Biostratigraphically significant taxa identified in the assemblage restrict the age of the studies deposits to the later early Maastrichtian. The palynofloral evidence indicates plant communities that mainly preferred higher-altitude areas and cooler-wetter conditions. However, a spore and pollen assemblage which derives from vegetation typical of lacustrine areas or riverbanks also occurs in the studied deposits. The studied palynoflora shows quite significant differences compared to previously published palynological assemblages from the same geological unit (the Sebeș Formation), although from deposits located slightly to the north, at Pâclișa. Organic geochemical data show that certain organic biomarkers such as n-alkanes n-C16 to n-C18 reach the highest values, and indicate that they may derive from freshwater colonial green algae such as Scenedesmus. The palynofacies constituents used to reconstruct the depositional environment suggest an exclusive terrestrial organic matter deposited in a stagnant-water fluvial/lacustrine environment.
通过对上白垩世 Oarda de Jos 大陆脊椎动物遗址的古生物学、古地貌学和有机地球化学调查,我们得以确定该遗址的年龄,并对其沉积环境做出更详细的推断。所发现的古动物群主要是被子植物花粉和蕨类植物孢子,以及罕见的裸子植物类群。从生物地层学的角度来看,在该组合中发现的重要类群将研究沉积的年代限定在晚马斯特里赫特早期。古植物证据表明,植物群落主要喜欢海拔较高的地区和凉爽潮湿的环境。不过,在所研究的沉积物中也出现了源自湖沼地区或河岸典型植被的孢子和花粉组合。所研究的古植物群落与之前公布的同一地质单元(塞贝雅地层)的古植物群落相比,差异非常明显,尽管这些古植物群落来自稍靠北的帕克里雅沉积层。有机地球化学数据显示,某些有机生物标志物(如 n-C16 至 n-C18 烷烃)达到了最高值,表明它们可能来自淡水殖民绿藻(如 Scenedesmus)。用于重建沉积环境的古乐彩网成分表明,只有陆地有机物沉积在死水河/湖泊环境中。
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引用次数: 0
PALYNOSTRATIGRAPHIC AND PALAEOENVIRONMENTAL INVESTIGATIONS OF THE MAASTRICHTIAN FROM OARDA DE JOS (SOUTHWESTERN TRANSYLVANIAN BASIN) 奥尔达-德约斯(西南特兰西瓦尼亚盆地)马斯特里赫特古地层的古地层学和古环境研究
Pub Date : 2024-02-11 DOI: 10.35463/j.apr.2024.01.07
D. Țabără, Z. Csiki-Sava
A palynological, palynofacies and organic geochemical investigation of the Upper Cretaceous continental vertebrate site of Oarda de Jos allowed us to constrain its age and to make more detailed inferences about its depositional environment. The recovered palynomorph assemblage is represented mainly by angiosperm pollen and fern spores, alongside rare taxa of gymnosperms. Biostratigraphically significant taxa identified in the assemblage restrict the age of the studies deposits to the later early Maastrichtian. The palynofloral evidence indicates plant communities that mainly preferred higher-altitude areas and cooler-wetter conditions. However, a spore and pollen assemblage which derives from vegetation typical of lacustrine areas or riverbanks also occurs in the studied deposits. The studied palynoflora shows quite significant differences compared to previously published palynological assemblages from the same geological unit (the Sebeș Formation), although from deposits located slightly to the north, at Pâclișa. Organic geochemical data show that certain organic biomarkers such as n-alkanes n-C16 to n-C18 reach the highest values, and indicate that they may derive from freshwater colonial green algae such as Scenedesmus. The palynofacies constituents used to reconstruct the depositional environment suggest an exclusive terrestrial organic matter deposited in a stagnant-water fluvial/lacustrine environment.
通过对上白垩世 Oarda de Jos 大陆脊椎动物遗址的古生物学、古地貌学和有机地球化学调查,我们得以确定该遗址的年龄,并对其沉积环境做出更详细的推断。所发现的古动物群主要是被子植物花粉和蕨类植物孢子,以及罕见的裸子植物类群。从生物地层学的角度来看,在该组合中发现的重要类群将研究沉积的年代限定在晚马斯特里赫特早期。古植物证据表明,植物群落主要喜欢海拔较高的地区和凉爽潮湿的环境。不过,在所研究的沉积物中也出现了源自湖沼地区或河岸典型植被的孢子和花粉组合。所研究的古植物群落与之前公布的同一地质单元(塞贝雅地层)的古植物群落相比,差异非常明显,尽管这些古植物群落来自稍靠北的帕克里雅沉积层。有机地球化学数据显示,某些有机生物标志物(如 n-C16 至 n-C18 烷烃)达到了最高值,表明它们可能来自淡水殖民绿藻(如 Scenedesmus)。用于重建沉积环境的古乐彩网成分表明,只有陆地有机物沉积在死水河/湖泊环境中。
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引用次数: 0
NEW MIDDLE TRIASSIC BELL-SHAPED NASSELLARIAN RADIOLARIA FROM ALPINE AND CARPATHIAN AREAS 高山和阡陌地区新发现的中三叠世钟状纳塞尔放射虫
Pub Date : 2024-02-11 DOI: 10.35463/j.apr.2024.01.05
P. Dumitrică
This paper is a continuation of a previous one in which I described and revised different Middle Triassic nassellarian Radiolaria from the Alps and Carpathians illustrated with drawings. On the present paper, are described two new groups of Middle Triassic dicyrtid Nassellaria with bell-shaped shell from the same area to enrich the inventory of these microfossils. Two new families (Gorispelidae and Humerocyrtiidae), five new genera (Silicotintinnabulum, Colpotrelus, Gorispela, Pozsvartia and Humerocyrtis) and 30 new species are described from these genera and illustrated with drawings.
这篇论文是前一篇论文的延续,在前一篇论文中,我描述并修订了阿尔卑斯山和喀尔巴阡山脉的不同中三叠世钟形放射虫,并附图说明。本文描述了同一地区出土的两组新的中三叠世钟形壳双壳类纳西利亚虫,以丰富这些微化石的清单。本文描述了这些属中的两个新科(Gorispelidae 和 Humerocyrtiidae)、五个新属(Silicotintinnabulum、Colpotrelus、Gorispela、Pozsvartia 和 Humerocyrtis)和 30 个新种,并附有插图。
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引用次数: 0
PARURGONINA VALANGINIANA N. SP. FROM THE VALANGINIAN OF SW IRAN (ZAGROS ZONE): THE FIRST CRETACEOUS RECORD OF THE GENUS 来自伊朗西南部(扎格罗斯带)的瓦兰吉尼亚Parurgonina valanginiana n. spp:该属的第一个白垩纪记录
Pub Date : 2023-10-18 DOI: 10.35463/j.apr.2024.01.03
FELIX SCHLAGINTWEIT, MOHSEN YAZDI-MOGHADAM, BRUNO GRANIER, ROHOLLAH HOSSEINZADEH
The new larger benthic foraminifer Parurgonina valanginiana is described from the Valanginian deposits of the Fahliyan Formation of southwestern Iran (Zagros Zone). It is separated from the uppermost Oxfordian-lowermost Tithonian generotype P. caelinensis (Cuvillier et al.) by a distinct time gap. The Lower Cretaceous specimens exhibit a larger embryonic chamber whereas the internal structural and other biometric data are generally in conformity with the Upper Jurassic morphotypes. Besides the Middle Jurassic P. primaeva Kamoun & Peybernès and the Upper Jurassic P. caelinensis, P. valanginiana sp. nov. represents the third species of the genus described and possibly a Valanginian marker endemic for the Arabian Plate. In the Fahliyan Formation, P. valanginiana sp. nov. occurs in wackestones associated with Pseudocyclammina lituus (Yokoyama). Additional data for the micropalaeontological assemblages (benthic foraminifera, calcareous algae) of the shallow-water carbonates of the Fahliyan Formation are provided. As it is also the case in the United Arab Emirates (Granier, 2008), it is proposed that the Tithonian/Berriasian boundary does not, as generally indicated in the literature, coincide with the boundary of the Hith Formation and the Fahliyan Formation, but instead occurs within the latter. The new finds provide further evidence that the Tithonian/Berriasian boundary did not represent a major rupture or extinction for larger benthic foraminifera. On the contrary, the Berriasian/Valanginian boundary corresponds to a natural discontinuity (both sedimentological and biological) possibly enhanced by a hiatus of the upper Berriasian strata. The foraminiferal diversity remains quite high during early Valanginian times then falls drastically, which suggests that the Berriasian/Valanginian crisis is a double barelled crisis (first base Valanginian, second intra-Valanginian).
在伊朗西南部(Zagros带)Fahliyan组Valanginian沉积中发现了一种新的大型底栖有孔虫Parurgonina valanginiana。它与最上层的牛津-最下层的梯统属P. caelinensis (Cuvillier et al.)有明显的时间间隔。下白垩统标本显示出较大的胚室,而内部结构和其他生物特征数据与上侏罗纪的形态类型基本一致。除了中侏罗世P. primaeva Kamoun &;P. valanginiana sp. 11 .代表了所描述的该属的第三种,可能是阿拉伯板块特有的Valanginian标志。在Fahliyan组,p.v alanginiana sp. 11 .出现在与pseudoyclammina lituus (Yokoyama)相关的砾石中。提供了法里岩组浅水碳酸盐微体古生物组合(底栖有孔虫、钙质藻类)的附加数据。正如在阿拉伯联合酋长国的情况一样(Granier, 2008),有人提出Tithonian/Berriasian边界并不像文献中普遍指出的那样与Hith组和Fahliyan组的边界一致,而是发生在后者内部。新的发现提供了进一步的证据,表明泰坦纪/贝里亚纪边界并不代表大型底栖有孔虫的重大断裂或灭绝。相反,贝里亚/瓦兰吉尼亚边界对应于自然不连续(沉积学和生物学),可能由于贝里亚上部地层的断裂而增强。有孔虫多样性在早期Valanginian时期保持相当高的水平,然后急剧下降,这表明Berriasian/Valanginian危机是一个双管危机(第一基Valanginian,第二内Valanginian)。
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引用次数: 0
SPINOSAURS AS PHYTOSAUR MIMICS: A CASE OF CONVERGENT EVOLUTION BETWEEN TWO EXTINCT ARCHOSAURIFORM CLADES 棘龙作为植物龙的模仿者:两个已灭绝的始祖进化分支趋同进化的一个案例
Pub Date : 2023-09-09 DOI: 10.35463/j.apr.2024.01.02
CHAN-GYU YUN
Spinosaurids are a highly morphologically specialized clade of large tetanuran theropod dinosaurs, characterized by their proportionally long, narrow snouts and conical teeth with fluted surfaces. The unusual morphology of spinosaurids has been usually compared with modern crocodilians, but distinct differences between them have been reported as well. Here, numerous craniodental characters that are shared by spinosaurids and large, robust phytosaurs often termed as “brachyrostral” forms are reviewed. Phytosaurs are a clade of carnivorous archosauriforms that resemble crocodilians in overall morphology, and as such a similar lifestyle has been inferred for them as well. Although based on preliminary observations, the shared craniodental characters of spinosaurids and “brachyrostral” phytosaurs, including those not shared with crocodilians, raise the possibility that at least some aspects of the lifestyle of these extinct archosauriform clades were more similar to each other rather than to that of crocodilians.
棘龙是大型破伤风目兽脚亚目恐龙的一个形态高度特化的分支,其特征是按比例排列的长而窄的口鼻和表面有凹槽的圆锥形牙齿。棘龙的不同寻常的形态通常与现代鳄鱼进行比较,但它们之间的明显差异也被报道过。本文回顾了棘龙类和大型健壮的植物龙所共有的许多颅齿特征,这些特征通常被称为“短吻”形式。植物龙是食肉始祖龙的一个分支,在整体形态上与鳄鱼相似,因此它们也有类似的生活方式。尽管基于初步的观察,棘龙和“短吻”植物龙(包括那些与鳄鱼不相同的)共有的颅齿特征提出了这样一种可能性,即至少在某些方面,这些已灭绝的始祖龙进化枝彼此之间的生活方式更相似,而不是与鳄鱼的生活方式更相似。
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引用次数: 0
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Acta Palaeontologica Romaniae
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