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Systematics, age, paleoecology and paleobiogeography of Middle to Late Jurassic benthic foraminiferal assemblages from Jumara Dome, Kutch, Gujarat, India 印度古吉拉特邦库奇Jumara Dome中晚侏罗世底栖有孔虫组合的系统、年龄、古生态学和古生物地理学
IF 1.5 4区 地球科学 Q3 PALEONTOLOGY Pub Date : 2021-01-01 DOI: 10.47894/mpal.67.1.06
Syed Md Wasim, A. Talib, S. H. Alvi
The studied section of the Patcham-Chari formations of Middle to Upper Jurassic rocks exposed at Jumara Dome, Kutch Basin, Gujarat, India yielded rich foraminiferal assemblages comprising 104 species, of which 49 are reported for the first time from the Indian region. The foraminiferal species are systematically described and illustratedwith the help of SEMimaging. The foraminiferal assemblages are used to interpret the age, paleoecology, and paleobiogeography. Although most of the foraminiferal species are rather long ranging, on the basis of a large number of species characteristic of the Bathonian, Callovian and Oxfordian, globally as well as in the Indian region, a Bathonian to Oxfordian age is assigned to the studied sequence. Based on the dominance of the suborder Lagenina, superfamily Nodosariacea, family Vaginulinidiae, calcareous hyaline forms, the genus Lenticulina, high Fisher’s diversity index, morphogroup J2, shallow infaunal life habit, detrivores, bacterial scavengers, and omnivores feeding strategy in the assemblages, a shallow water, near shore, open marine environment ranging from mid- to outer shelf with normal salinity and normal to well-oxygenated waters with high nutrient influx is interpreted for the studied assemblages. The Middle to Late Jurassic Jumara foraminiferal assemblages are assigned to a separate province of the Antiboreal Realm, the Indo-East African Province, located at the southern margin of the Tethyan Realm and transitional between the Tethyan and the Antiboreal realms.
对印度古吉拉特邦库奇盆地Jumara Dome中上侏罗统patchama - chari组进行了研究,发现了丰富的有孔虫组合,包括104种,其中49种为印度地区首次报道。在半成像的帮助下,系统地描述和说明了有孔虫物种。有孔虫组合被用来解释时代、古生态和古生物地理。虽然大多数有孔虫物种分布范围较长,但根据全球以及印度地区的大量Bathonian, Callovian和oxford的物种特征,将所研究的层序划分为Bathonian - oxford时代。基于Lagenina亚目、Nodosariacea超科、Vaginulinidiae科、calcalhyaline形态、Lenticulina属、高fishericoss多样性指数、形态群J2、浅水动物生活习性、腐食动物、细菌食腐动物和杂食动物的摄食策略,在浅水、近海岸,所研究的组合解释了从中陆架到外陆架的开放海洋环境,盐度正常,正常到富氧,高营养物流入。中至晚侏罗世Jumara有孔虫组合被分配到Antiboreal领域的一个单独的省,印度-东非省,位于特提斯王国的南缘,是特提斯和Antiboreal领域之间的过渡。
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引用次数: 5
Eocene larger foraminiferal biostratigraphy, depositional history and paleogeography of the Sylhet Limestone of the Mikir Hills of Assam, NE India: Implications for an Open Tethys 印度东北部阿萨姆邦Mikir山Sylhet石灰岩始新世大型有孔虫生物地层、沉积历史和古地理:对开放特提斯的启示
IF 1.5 4区 地球科学 Q3 PALEONTOLOGY Pub Date : 2021-01-01 DOI: 10.47894/mpal.67.5.01
Shantajhara Biswal, Kapesa Lokho, U. Shukla, K. Whiso, K. Prakash
This study presents results of integrated larger foraminiferal biostratigraphy, lithofacies and stable carbon isotope (13 C org) analysis carried out in the Sylhet Limestone of Dillai Parbat Limestone Mine succession, Karbi Anglong District, Assam. Larger foraminiferal assemblages assign a middle Eocene age to the studied succession of Sylhet Limestone. The Shallow Benthic Zone (SBZ) 13 and SBZ 16-18 and a barren interval in between is recognized in the studied part of the Sylhet Limestone. The SBZ 13 Zone (early Lutetian) is represented by fossiliferous limestone at the base of the section and is inferred to have been deposited in a subtidal environment. The overlying barren interval lies in the arenaceous interval and was likely deposited in a subtidal channel sand shoaling bars. The upper part of the section ismarked by SBZ16-18 (late Lutetian to Bartonian) and was deposited in subtidal channel sand inner neritic zone environments, followed by intertidal marl to subtidal limestone. The organic carbon stable isotopic data reveals a sharp negative excursion at about 34-35m of the succession, which may be correlated with Middle Eocene Climatic Optimum (MECO) and/or vegetation types. This and earlier studies from India and elsewhere indicate that during the middle Eocene, the Neo-Tethys Sea was open and connected through the East and West coasts of India, Northeast India, western Himalaya, and most of the Middle East and southeast Europe. This vast area produced enormous amount of petroleum.
本文介绍了阿萨姆邦Karbi Anglong地区Dillai Parbat石灰岩矿演替中Sylhet石灰岩的大型有孔虫生物地层、岩相和稳定碳同位素(13corg)综合分析结果。较大的有孔虫组合确定了所研究的锡尔赫特石灰岩演替的始新世中期。在锡尔赫特灰岩研究区发现了浅底栖带(SBZ) 13和SBZ 16-18及其之间的一段贫瘠带。sbz13带(早鲁特世)以剖面底部的化石灰岩为代表,推测其沉积于潮下环境。上覆秃层位于砂质层中,可能沉积于潮下河道砂滩坝中。剖面上部以SBZ16-18为标志(晚鲁特世—巴尔顿世),沉积于潮下河道砂内浅海带环境,其次为潮间泥灰岩—潮下灰岩。有机碳稳定同位素数据显示,在演替的34 ~ 35m处出现了急剧的负偏移,这可能与中始新世最适气候(MECO)和/或植被类型有关。这项研究以及来自印度和其他地方的早期研究表明,在始新世中期,新特提斯海是开放的,并通过印度的东西海岸、印度东北部、喜马拉雅西部、中东和东南欧的大部分地区连接在一起。这片广阔的地区出产了大量的石油。
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引用次数: 2
Septfontainella carpatobalcanica n. gen., n. sp., a new hauraniid foraminifera from the upper Berriasian of Serbia and Romania 塞尔维亚和罗马尼亚上Berriasian的一种新的hauriid有孔虫
IF 1.5 4区 地球科学 Q3 PALEONTOLOGY Pub Date : 2021-01-01 DOI: 10.47894/mpal.67.6.05
F. Schlagintweit, I. Bucur, George Pleș
Septfontainella (type-species S. carpatobalcanica n. sp.), a new genus of larger benthic foraminifera (family Hauraniidae, subfamily Amijiellinae) is described from the upper Berriasian of the Carpatho-Balkanides, eastern Serbia and Romania (Southern Capathians), both part of the Getic domain. It is characterized by large elongate dimorphic tests with a reduced coiled stage and a rectilinear-uniserial adult part with chambers displaying a coarse and irregular subepidermal network. Triangular-shaped first order radial partitions extend into the chamber lumen attaching to the succeeding septum. Septfontainella gen. nov. is compared with the Lower Jurassic Cymbriaella Fugagnoli and the Lower Cretaceous Bramkampella Redmond. Septfontainella n. gen. further substantiates the occurrence of a Berriasian origination event of larger benthic foraminifera in inner carbonate platform facies of the Neotethysian realm.
Septfontainella(模式种S. carpatobalcanica n. sp.)是一种大型底栖有孔虫(Hauraniidae, Amijiellinae亚科)的新属,描述于喀尔巴阡-巴尔干半岛的上Berriasian,塞尔维亚东部和罗马尼亚(南卡帕西亚),两者都是古域的一部分。它的特征是大的细长二态试验,具有减少的卷曲阶段和直线-单列的成体部分,其腔室显示粗糙和不规则的皮下网络。三角形的一级放射状隔墙延伸到腔内,与后继隔膜相连。将Septfontainella gen. 11与下侏罗世Cymbriaella Fugagnoli和下白垩世Bramkampella Redmond进行了比较。Septfontainella n. gen进一步证实了新特提斯宙内碳酸盐岩台地相大型底栖有孔虫Berriasian起源事件的发生。
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引用次数: 0
Little pieces of mystery from the Silurian of the Dingle Peninsula, Ireland 来自爱尔兰丁格尔半岛志留纪的神秘小片段
IF 1.5 4区 地球科学 Q3 PALEONTOLOGY Pub Date : 2021-01-01 DOI: 10.47894/mpal.67.6.03
A. Ferretti, Lorenzo Serafini
Three-dimensionally preserved fragments of the enigmatic organism Sandvikina, previously reported from the Silurian of Ireland, Scotland and Scandinavia, were recovered from the original Irish material that had revealed the problematic in thin sections. The new specimens derive from conodont residue and document silicified sectors of the distal part of the skeleton. Adifferent pattern characterizes the outer and inner walls, with an external net-like reticulated and an internal continuous and apparently imperforate framework, getting therefore a filter-like feeding system function highly improbable.
先前在爱尔兰、苏格兰和斯堪的纳维亚的志留纪发现的神秘生物Sandvikina的三维保存碎片,从原始的爱尔兰材料中恢复,这些材料在薄切片中揭示了问题。这些新标本来自牙形石残留物,并记录了骨骼远端部分的硅化部分。外墙和内墙有着不同的图案,外部是网状的网状结构,内部是连续的、明显不穿孔的框架,因此像过滤器一样的供料系统的功能是极不可能的。
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引用次数: 0
Nephrolepidina and unispiralled Miogypsinidae from the Oligo-Miocene toe-of-slope succession of Gran Sasso (L'Aquila, Central Apennines - Italy): biometric and evolutionary remarks 新近至中新世Gran Sasso (L'Aquila,中亚平宁-意大利)坡脚演替中的肾鳞目和单尖目Miogypsinidae:生物特征和进化评论
IF 1.5 4区 地球科学 Q3 PALEONTOLOGY Pub Date : 2021-01-01 DOI: 10.47894/mpal.67.5.04
F. Schiavinotto, A. Benedetti
A biometric analysis was performed on twelve Nephrolepidina and ten Miogypsina and Miogypsinoides populations, coming from the Tre Valloni section, an Oligo-Miocene calcarenitic toe-of-slope succession outcropping in the southern sector of Gran Sasso (L'Aquila, Central Apennines - Italy). Miogypsina populations are investigated applying two essential parameters, i.e., the number of the nepionic chambers and the protoconch diameter. Previously detected evidences of reworking in the same material suggested to use some cautions to avoid misleading results. A mixed typological-statistical methodology is used, leading from starting "disordered" assemblages to final "virtual but reliable" populations, cleaned up from reworked specimens, in order to implement the accuracy of the biostratigraphic results. This goal is reached for the lower part of the succession only. According to the biometric limits proposed for the various chronospecies in the recent literature, the resulting populations are referred to Miogypsinoides ex. interc. complanatus-formosensis up to Md. bantamensis, and to Miogypsina basraensis up to M. ex. interc. gunteri-tani. This sequence of taxa, in general, is in agreement with the principle of nepionic acceleration, with a progressive reduction in the average number of nepionic chambers (X), arranged in a single spiral, just peculiar in the more primitive populations. Miogypsinoides bantamensis and Miogypsina basraensis are recognized biometrically for the first time in the Central Apennines. Nephrolepidina populations are investigated applying the recently introduced biometry of the fifth stage of the neanic equatorial chamberlets, supplying additional data about parameters and factors that had already demonstrated to have taxonomic-biostratigraphic significance. The biometric analysis of the embryo-nepionic stages was performed as an integration of the data obtained in a previous study, excluding the uppermost samples in which Nephrolepidina is clearly reworked, being characterized by primitive morphology, i.e., almost isolepidine embryo, few adauxiliary chamberlets, and ogival equatorial chamberlets. The neanic data evidence that the "Degree of stolonic distalization" (Factor FD5) and the "Shape Index" (Factor SI5), at the 5th nepionic stage of the population have a very good fitting with the previously evidenced Neanic acceleration, as the overall trend to modify the shape of the equatorial chambers, connected with a gradually more distal position of the radial stolons. Unispiralled Miogypsina, associated with Nephrolepidina at an advanced evolutionary stage, suggest a continuous reworking of primitive miogypsinids, without an involvement of more evolute bispiralled forms. The biostratigraphic assessment of the Venacquaro and Glauconitic Calcarenite formations is improved, assigning the Venacquaro Formation to the Chattian-lowermost Aquitanian (SBZ22B-basal SBZ24) and the base of the Glauconitic Calcatenite to the Aquitanian (SBZ2
对意大利Gran Sasso (L’aquila,中亚平宁)南段渐新世-中新世钙屑岩坡脚演替露头的Tre Valloni剖面的12个Nephrolepidina、10个Miogypsina和Miogypsinoides种群进行了生物特征分析。采用两个基本参数,即海螺腔的数量和原海螺直径,对Miogypsina种群进行了研究。先前在同一材料中发现的返工证据建议使用一些谨慎措施以避免误导性结果。使用混合类型-统计方法,从开始的“无序”组合到最终的“虚拟但可靠”种群,从重新加工的标本中清理出来,以实现生物地层学结果的准确性。这一目标只适用于继承权较低的部分。根据最近文献中提出的各种时间物种的生物特征限制,所得到的种群被称为miogypsinides ex. interc。直到班塔马氏斑胸蛾和直到外interc斑胸蛾的basramiogypsina。gunteri-tani。总的来说,这个分类群的序列与nepionic加速的原理是一致的,nepionic室的平均数量(X)逐渐减少,排列成一个螺旋状,这在更原始的种群中是特有的。bantamensis miogypsinides和basraensis miogypsinensis在亚平宁中部首次被生物识别。本文应用新近引入的新赤道腔室第五期生物计量学方法对肾鳞虫种群进行了研究,提供了关于已经证明具有分类-生物地层学意义的参数和因素的额外数据。胚胎-肾腺阶段的生物特征分析是作为先前研究数据的整合进行的,排除了最上面的样本,其中肾腺被明显改造,其特征是原始形态,即几乎是异肾腺胚胎,少数辅助腔室和卵母赤道腔室。neanic数据表明,种群第5 neanic阶段的“匍匐体远化程度”(FD5因子)和“形状指数”(SI5因子)与先前证明的neanic加速度有很好的拟合,这是赤道室形状改变的总体趋势,与径向匍匐体逐渐向远端移动有关。单螺旋形Miogypsina在进化的高级阶段与肾鳞翅目(Nephrolepidina)有关联,表明原始Miogypsina的不断改造,而没有更进化的双螺旋形Miogypsina的参与。改进了Venacquaro组和海绿石钙砾岩组的生物地层评价,将Venacquaro组划分为夏梯-最下阿基坦期(sbz22b -基底SBZ24),将海绿石钙砾岩的基底划分为阿基坦期(SBZ24)。所研究的沉积序列是沿坡向坡脚沉积的,其特征是由浅水环境迁移的分类群和沿河道侵蚀的经过改造的原始标本混合而成。
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引用次数: 3
Taxonomic reassessment of four larger benthic Foraminifera (soritoids and orbitolinids) described from the Lower to mid-Cretaceous shallow-water carbonates of Iraq 伊拉克下白垩世至中白垩世浅水碳酸盐中描述的四种大型底栖有孔虫(soritoids和orbitolinids)的分类重新评估
IF 1.5 4区 地球科学 Q3 PALEONTOLOGY Pub Date : 2021-01-01 DOI: 10.47894/mpal.67.5.05
L. Consorti, F. Schlagintweit
Four Cretaceous foraminifers belonging to the subfamilies Pseudorhapydionininae, Orbitolininae and Dictyoconinae are here taxonomically revised: Pseudorhipidionina tubaensis, Bicyclina rifaiensis, Orbitolina arcuala, and Simplorbitolina nakhali. It is revealed that they should be considered junior synonyms of several existing species or inaccurately defined for specific identification. The new nominal names introduced to indicate such synonyms are invalidated, thus they are to be considered as nomen nudum.
本文对白垩纪有孔虫pseudorhapydioninae亚科、orbitolinae亚科和Dictyoconinae亚科的4种有孔虫进行了分类修正:Pseudorhipidionina tubaensis、Bicyclina rifaiensis、Orbitolina arcuala和Simplorbitolina nakhali。结果表明,它们应该被认为是几个现存物种的低级同义,或者是为了特定的识别而不准确地定义。为表示这类同义词而引入的新的名义名称是无效的,因此它们被认为是nomen nudum。
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引用次数: 3
Transfer of type specimens of Caribbean Foraminifera and Ostracoda to the European Micropalaeontological Reference Centre 加勒比海有孔虫和介形虫的模式标本转移到欧洲微体古生物学参考中心
IF 1.5 4区 地球科学 Q3 PALEONTOLOGY Pub Date : 2021-01-01 DOI: 10.47894/mpal.67.1.07
B. Wilson
While working on Trinidad and Tobago and the western tropical Atlantic Ocean, the author published four papers (Wilson 2006a, 2006b, 2006c;Wilson et al. 2011) that included descriptions of ten new species of foraminifera and ostracoda from the Neogene of the SE Caribbean Sea. These holotypes and associated paratypes have now been transferred to the European Micropalaeontological Reference Centre, AGH, University of Science and Technology, Krakow, Poland, for safekeeping. Among the foraminifera, these are: Amphistegina martybuzasi Wilson, Ramkissoon and McLean 2011; Bolivina jiattongi Wilson 2006; Textularia carrbrowni Wilson 2006; Textularia framptoni Wilson 2006; and Textularia sawhi Wilson 2006. Regarding ostracodes, they are: Bythoceratina coimbrai Wilson 2006; Cytheropteron bremani Wilson 2006; Polycope swietinia Wilson 2006; Polycope trinitatensis Wilson 2066; and Pterygocythereis fithiani Wilson 2006. The specimen slides have been curated in Cabinet 7, Drawer 27.
在研究特立尼达和多巴哥和热带大西洋西部时,作者发表了四篇论文(Wilson 2006a, 2006b, 2006c;Wilson et al. 2011),其中包括对加勒比海东南部新近纪有孔虫和介形虫的十种新种的描述。这些全型和相关的副型现已转移到波兰克拉科夫科技大学AGH欧洲微体古生物学参考中心保存。在有孔虫中,有:Amphistegina martybuzasi Wilson, Ramkissoon and McLean 2011;玻利维亚贾通吉威尔逊2006;Textularia carrbrowni Wilson 2006;浙江本草,2006;和texularia sawhi Wilson 2006。关于介形类,它们是:Bythoceratina coimbrai Wilson 2006;刘志强,2006;林志刚,2006;Polycope trinitatensis Wilson 2066;菲蒂亚尼·威尔逊,2006。标本幻灯片被收藏在7号柜27号抽屉里。
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引用次数: 0
Moncharmontia De Castro 1967, benthic foraminifera from the middle-upper Cenomanian of the Sarvak Formation of SW Iran (Zagros Zone): a CTB survivor taxon Moncharmontia De Castro 1967,伊朗西南部(Zagros区)Sarvak组Cenomanian中上段底栖有孔虫:一个CTB幸存分类群
IF 1.5 4区 地球科学 Q3 PALEONTOLOGY Pub Date : 2021-01-01 DOI: 10.47894/mpal.67.1.03
F. Schlagintweit, M. Yazdi-Moghadam
Moncharmontia apenninca (De Castro 1966) is widely reported in the Upper Cretaceous (Turonian-Campanian) shallow-water successions of the Neotethyan realm (e.g., Italy, Croatia). With its "First Appearance Datum" (FAD) in the lowermost Turonian, it is considered a newcomer taxon in the aftermath of the Cenomanian/Turonian boundary (CTB) extinction event, and therefore a member of the Upper Cretaceous Global Community Maturation Cycle. Its common presence in the middle-pro parte upper Cenomanian Sarvak Formation and time-equivalent strata in Mexico (Valles-San Luis Potosi Platform) document that it represents a CTB extinction survivor taxon like Dicyclina, Nezzazata, Reticulinella and several others. The different local "FAD's" of Moncharmontia might possibly be linked to latitudinal differences, i.e., an earlier evolution of the species in the near-equatorial area (Mexico, Iran) compared to the low-middle latitudinal occurrences in the peri-Mediterranean realm (e.g., Italy, Turkey).
Moncharmontia apenninca (De Castro 1966)在新特提斯王国(如意大利、克罗地亚)的上白垩纪(Turonian-Campanian)浅水序列中被广泛报道。由于其“首次出现基准点”(First Appearance Datum, FAD)位于最底的Turonian,因此被认为是Cenomanian/Turonian边界(CTB)灭绝事件后的一个新分类单元,因此是上白垩纪全球群落成熟周期的成员。它在墨西哥(vales - san Luis Potosi台地)的上Cenomanian Sarvak组中后期和时间相当的地层中普遍存在,表明它代表了CTB灭绝幸存者的分类群,如Dicyclina, Nezzazata, Reticulinella和其他几个。Moncharmontia地区不同的“FAD”可能与纬度差异有关,即,与地中海周边地区(如意大利、土耳其)的低中纬度事件相比,近赤道地区(墨西哥、伊朗)的物种进化时间更早。
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引用次数: 13
Automating taxonomic and systematic search of benthic foraminifera in an online database 联机数据库中底栖有孔虫的自动分类和系统搜索
IF 1.5 4区 地球科学 Q3 PALEONTOLOGY Pub Date : 2021-01-01 DOI: 10.47894/mpal.67.6.06
A. Amao
Recent advances in the applications of deep neural networks in computer vision tasks such as image classification has seen a tremendous surge in interest. Several image classification algorithms can now be leveraged in automating some tedious tasks associated with benthic foraminifera research especially in sample picking, taxonomy and systematics. In this study, a small image identification model was built with 414 SEM micrographs representing twenty-one species of benthic foraminifera, using a convolutional neural network which achieved 84% model accuracy and 75% validation accuracy on previously unseen images. The model was also deployed through a web application to demonstrate how it may be useful in augmenting online databases such as the Ellis Messina catalogue and the World Register of Marine Species. These services although very valuable, can be modernized with image search functionalities to enhance their perpetual usefulness and continuity.
最近深度神经网络在计算机视觉任务(如图像分类)中的应用取得了巨大的进展,引起了人们的极大兴趣。现在,一些图像分类算法可以用来自动化一些与底栖有孔虫研究相关的繁琐任务,特别是在样本采集、分类和分类学方面。在这项研究中,使用卷积神经网络建立了一个小型图像识别模型,该模型具有84%的模型准确率和75%的先前未见过的图像验证准确率。该模型还通过一个网络应用程序进行了部署,以展示它如何在增加在线数据库方面发挥作用,例如埃利斯·梅西纳目录和世界海洋物种登记册。这些服务虽然非常有价值,但可以通过图像搜索功能进行现代化,以增强其永久的有用性和连续性。
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引用次数: 1
A Neotype for Neogloboquadrina atlantica (Berggren 1972) atlantica新球虫的一种新类型(Berggren 1972)
IF 1.5 4区 地球科学 Q3 PALEONTOLOGY Pub Date : 2021-01-01 DOI: 10.47894/mpal.67.1.08
M. Kaminski, W. Berggren
Neogloboquadrina atlantica (Berggren 1972) is one of the most common planktonic foraminiferal species in the Upper Miocene and Pliocene at DSDP/IODP sites in the northern Atlantic. It was originally described from the upper Pliocene at DSDP Site 116 in the Rockall Trough, northeastern Atlantic. In the original description of the species, the depository of the illustrated type specimens was not specified. One of us (WAB) was under the impression that the illustrated type specimens of Neogloboquadrina atlantica had been sent to the Cushman Collection at the Smithsonian Institution. Unfortunately there is no record of the specimen slides having ever arrived at the museum (B. Huber, personal communication, 2020). Fortunately, the second author is still in possession of the shipboard sample residues from DSDP Leg 12, and was able to select a neotype specimen to replace the missing illustrated holotype. The purpose of this note is to designate a neotype specimen for Neogloboquadrina atlantica and to specify a repository for the permanent archive of the neotype and paraneotypes.
Neogloboquadrina atlantica (Berggren, 1972)是北大西洋DSDP/IODP站点中新世和上新世最常见的浮游有孔虫物种之一。它最初是在大西洋东北部Rockall海槽的DSDP Site 116上新世被发现的。在该物种的原始描述中,所示模式标本的存放处没有指定。我们中的一个人(WAB)以为附有插图的大西洋新舌龙的模式标本已经送到史密森学会的库什曼收藏馆了。不幸的是,没有关于标本幻灯片到达博物馆的记录(B. Huber, personal communication, 2020)。幸运的是,第二作者仍然拥有DSDP第12腿的船上样本残留物,并能够选择一个新模式标本来取代缺失的图解全模。本说明的目的是为大西洋新舌龙指定一个新类型标本,并指定一个永久存档新类型和副类型的储存库。
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Micropaleontology
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