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Height increment patterns in Pinus pinaster seedlings emerging in naturally regenerated gaps 天然更新间隙中出现的松柏幼苗的高度递增模式
IF 2.8 2区 农林科学 Q1 FORESTRY Pub Date : 2024-01-31 DOI: 10.1093/forestry/cpae002
Sergio de Frutos, Rubén Manso, Sonia Roig-Gómez, Ricardo Ruiz-Peinado, Miren del Río, José Alfredo Bravo-Fernández
The group selection cutting system provides a promising technique for transforming monospecific even-aged stands into uneven-aged, mixed stands. However, this system has scarcely been used to regenerate Mediterranean shade-intolerant species. In this study, we analyze the height increment patterns of Pinus pinaster seedlings emerging in naturally regenerated gaps. P. pinaster is a shade-intolerant pine species distributed throughout the Mediterranean basin. An experimental trial was established in a 50-year-old P. pinaster (maritime pine) plantation located in the mountains of central Spain. Gaps of two different sizes (diameter equal to 1.5 and 2.5 times the dominant height of the stand, 26 and 44 m, respectively) were opened during winter 2017–18. A total of 531 subplots with a 1-m radius were installed in spring 2018, distributed within the gaps as well as in control plots adjacent to the gaps. Seedlings were tagged individually and variables including height, along with other ecological variables, were collected twice a year (in spring and autumn), providing six time intervals for the development of our models. We applied seemingly unrelated regression to analyze the height growth in this polycyclic species, allowing us to model spring and autumn height increments at the same time. Our results showed that seedling height growth depended on the silvicultural treatment, meteorological conditions during the spring growth season, inter-specific scrub competition, position within the gap, and seedling characteristics, such as the ontogenetic development or the size of the seedling itself. This work, along with germination and survival analysis, aims to provide a holistic perspective on the suitability of the group selection cutting system to regenerate maritime pine stands, taking into account current and future climate change scenarios.
群体选择砍伐系统为将单一的均匀树龄林分转变为不均匀的混合树龄林分提供了一种很有前景的技术。然而,这种系统很少用于地中海耐阴树种的再生。在这项研究中,我们分析了在自然再生隙地中出现的松柏幼苗的增高模式。松柏是一种不耐阴的松树,分布在整个地中海盆地。我们在西班牙中部山区的一个有 50 年树龄的松树(海洋松)种植园中进行了一项实验。在 2017-18 年冬季,开辟了两种不同大小的间隙(直径分别相当于林分主要高度的 1.5 倍和 2.5 倍,即 26 米和 44 米)。2018 年春季共设置了 531 个半径为 1 米的子地块,分布在间隙内以及间隙附近的对照地块中。幼苗被单独标记,包括高度在内的变量以及其他生态变量每年收集两次(春季和秋季),为我们建立模型提供了六个时间间隔。我们采用看似无关的回归方法来分析这种多环树种的高度增长,使我们能够同时建立春季和秋季的高度增长模型。我们的研究结果表明,幼苗高度的增长取决于造林处理、春季生长季节的气象条件、灌丛间的竞争、间隙内的位置以及幼苗的特征,如幼苗本身的生长发育或大小。这项工作以及发芽率和存活率分析旨在提供一个全面的视角,在考虑到当前和未来气候变化情景的情况下,研究组选砍伐系统是否适合于再生海洋松林。
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引用次数: 0
Comparing the effects of ground cultivation and protection against browsing upon the natural regeneration of Scots pine and birch in a Caledonian pinewood 比较加里东松林中苏格兰松树和桦树的地面栽培和防砍伐对其自然再生的影响
IF 2.8 2区 农林科学 Q1 FORESTRY Pub Date : 2024-01-29 DOI: 10.1093/forestry/cpad068
W L Mason, S Rao, J Agnew, V Stokes, A Painting, T-K Clarke, C Edwards
The Caledonian pinewoods of northern Scotland represent the remnants of once extensive forests found at the western edge of the natural range of Scots pine (Pinus sylvestris L.). The surviving pinewoods are of international significance and are of high conservation and cultural value. However, for many decades, there has been concern about their long-term future because of a lack of regenerating seedlings and young trees. This problem is thought to be driven by high deer populations resulting in intensive browsing and severe damage to young trees, as well as a lack of favourable micro-sites for seedling germination. An experiment was established on the Mar Lodge estate in 2007–08 adjacent to native pinewood stands to test the effects of protection and site cultivation upon seedling numbers and growth. The experiment was located within a regeneration zone where intensive deer control had reduced numbers to <5 animals km−2. Ten years after initiation, occurrence and numbers of both Scots pine and birch seedlings were substantially increased in cultivated treatments compared with uncultivated controls. There were no effects of protection upon seedling numbers. Protection improved height growth of Scots pine, but not of birch. Cultivation had no effect upon the growth of pine seedlings but did influence the height of birch seedlings due to the rapid growth of some pre-existing seedlings in the uncultivated control. Vegetation cover had increased from ~60%to at least 80% cover 6 years after cultivation. The results suggest that site cultivation may be desirable to boost seedling germination within a Caledonian pinewood, but sustained measures to control browsing pressure are a prerequisite for satisfactory establishment of these seedlings.
苏格兰北部的喀里多尼亚松林是苏格兰松树(Pinus sylvestris L.)自然分布区西部边缘曾经广阔森林的遗迹。现存的松林具有国际意义,具有很高的保护和文化价值。然而,几十年来,由于缺乏再生幼苗和幼树,人们一直担心它们的长远未来。据认为,造成这一问题的原因是鹿的大量繁殖,导致密集的啃食和对幼树的严重破坏,以及缺乏有利于幼苗发芽的微生境。2007-08 年,我们在玛尔洛奇庄园原生松林附近进行了一项实验,以测试保护和场地栽培对幼苗数量和生长的影响。该实验位于一个再生区内,在该区域内,密集的鹿控制已将鹿的数量减少到<5只/平方公里。试验开始 10 年后,与未开垦的对照组相比,开垦处理的苏格兰松树和桦树幼苗的出现率和数量都大幅增加。保护措施对幼苗数量没有影响。保护措施改善了苏格兰松树的高度生长,但没有改善桦树的高度生长。栽培对松树幼苗的生长没有影响,但对桦树幼苗的高度有影响,这是因为在未栽培对照组中,一些原有幼苗生长迅速。栽培 6 年后,植被覆盖率从 ~60% 提高到至少 80%。结果表明,在喀里多尼亚松林中进行现场栽培可能会促进幼苗发芽,但持续采取措施控制啃食压力是这些幼苗顺利成活的先决条件。
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引用次数: 0
Modeling basal area yield using simultaneous equation systems incorporating uncertainty estimators 利用包含不确定性估算器的同步方程系统建立基底面积产量模型
IF 2.8 2区 农林科学 Q1 FORESTRY Pub Date : 2024-01-25 DOI: 10.1093/forestry/cpae001
Simón Sandoval, Cristián R Montes, Bronson P Bullock
Over the last three decades, many growth and yield systems developed for the southeast USA have incorporated methods to create a compatible basal area (BA) prediction and projection equation. This technique allows practitioners to calibrate BA models using both measurements at a given arbitrary age, as well as the increment in BA when time series panel data are available. As a result, model parameters for either prediction or projection alternatives are compatible. One caveat of this methodology is that pairs of observations used to project forward have the same weight as observations from a single measurement age, regardless of the projection time interval. To address this problem, we introduce a variance–covariance structure giving different weights to predictions with variable intervals. To test this approach, prediction and projection equations were fitted simultaneously using an ad hoc matrix structure. We tested three different error structures in fitting models with (i) homoscedastic errors described by a single parameter (Method 1); (ii) heteroscedastic errors described with a weighting factor ${w}_t$ (Method 2); and (iii) errors including both prediction ($overset{smile }{varepsilon }$) and projection errors ($tilde{varepsilon}$) in the weighting factor ${w}_t$ (Method 3). A rotation-age dataset covering nine sites, each including four blocks with four silvicultural treatments per block, was used for model calibration and validation, including explicit terms for each treatment. Fitting using an error structure which incorporated the combined error term ($overset{smile }{varepsilon }$ and $tilde{varepsilon}$) into the weighting factor ${w}_t$ (Method 3), generated better results according to the root mean square error with respect to the other two methods evaluated. Also, the system of equations that incorporated silvicultural treatments as dummy variables generated lower root mean square error (RMSE) and Akaike’s index values (AIC) in all methods. Our results show a substantial improvement over the current prediction-projection approach, resulting in consistent estimators for BA.
在过去的三十年里,为美国东南部开发的许多生长和产量系统都采用了创建兼容的基部面积(BA)预测和推算方程的方法。这项技术使实践者能够使用给定任意树龄的测量数据以及在有时间序列面板数据时的 BA 增量来校准 BA 模型。因此,无论是预测还是推算的模型参数都是兼容的。这种方法的一个注意事项是,用于向前预测的成对观测数据与单一测量年龄的观测数据具有相同的权重,而与预测时间间隔无关。为了解决这个问题,我们引入了一个方差-协方差结构,对不同时间间隔的预测给予不同的权重。为了测试这种方法,我们使用一个特别的矩阵结构同时拟合了预测方程和预测方程。我们在拟合模型时测试了三种不同的误差结构:(i) 用单一参数描述的同方差误差(方法 1);(ii) 用加权因子 ${w}_t$ 描述的异方差误差(方法 2);(iii) 在加权因子 ${w}_t$ 中包含预测误差($overset{smile }{varepsilon }$)和投影误差($tilde{varepsilon}$)的误差(方法 3)。用于模型校准和验证的轮伐期数据集涵盖九个地点,每个地点包括四个区块,每个区块有四种造林处理,包括每种处理的显式项。使用将组合误差项($overset{smile }{varepsilon}$和$tilde{varepsilon}$)纳入加权因子${w}_t$的误差结构(方法 3)进行拟合,根据均方根误差得出的结果优于其他两种评估方法。此外,在所有方法中,将造林处理作为虚拟变量的方程组产生的均方根误差(RMSE)和阿凯克指数值(AIC)都较低。我们的结果表明,与目前的预测-投影方法相比,我们的方法有了很大的改进,为 BA 提供了一致的估计值。
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引用次数: 0
More water, less light: how to improve silver fir seeding to convert Norway spruce monocultures into mixed stands in a drier region of Germany 更多的水,更少的光:如何改进银杉播种,将德国干旱地区的挪威云杉单一种植转变为混合种植
IF 2.8 2区 农林科学 Q1 FORESTRY Pub Date : 2024-01-20 DOI: 10.1093/forestry/cpad069
Christian Schattenberg, Ana Stritih, Dominik Thom
Adaptive forest management gains importance as climate and disturbance regimes continue to change. Norway spruce monocultures are particularly vulnerable to those changes. Thus, there is a strong demand to convert pure spruce stands toward better adapted forest ecosystems. Silver fir has similar wood properties as Norway spruce, but is less vulnerable to droughts. Yet, little is known how to efficiently admix silver fir into Norway spruce monocultures. We analyzed seedling abundance in mature Norway spruce monocultures 1 year after soil scarification and silver fir seeding in a relatively dry region of southern Germany. Our objectives were to study the effect of soil scarification and seeding on regeneration density and to identify drivers of seedling abundance. We collected data on regeneration, forest structure, as well as soil and light conditions at 103 scarified and 25 untreated control plots. We used non-parametric independence tests to compare scarified and untreated plots. Subsequently, we used boosted regression trees to investigate the drivers of seedling abundance. Norway spruce strongly benefited from soil scarification (P < .001), with seedling numbers even exceeding those of silver fir. Both species thrived in soils with high plant available water capacity. In contrast, seedling numbers were overall negatively affected by light. Moreover, we found a higher density of silver fir regeneration at greater distance from neighboring trees and in soils devoid of carbonate. Our study indicates that water, not light, is the limiting factor for both Norway spruce and silver fir seedlings in the study region. Seeding silver fir will likely be most effective underneath an intact canopy on deep, fine-textured soils without carbonate, while maximizing the distance to Norway spruce trees. In conclusion, silver fir seeding can be optimized to become an effective adaptive measure to diversify Norway spruce monocultures, and thus to create more resistant and resilient forest ecosystems.
随着气候和干扰机制的不断变化,适应性森林管理变得越来越重要。挪威云杉单一种植尤其容易受到这些变化的影响。因此,人们强烈要求将纯云杉林改造成适应性更强的森林生态系统。银杉的木材特性与挪威云杉相似,但不易受干旱影响。然而,人们对如何将银杉有效地混入挪威云杉单植林却知之甚少。我们在德国南部一个相对干旱的地区分析了土壤除痕和银杉播种1年后成熟的挪威云杉单一栽培中的幼苗丰度。我们的目标是研究土壤除痕和播种对再生密度的影响,并确定幼苗丰度的驱动因素。我们收集了 103 个已覆土地块和 25 个未覆土地块的再生、森林结构、土壤和光照条件数据。我们使用非参数独立性检验来比较疤痕化地块和未处理地块。随后,我们使用增强回归树来研究幼苗丰度的驱动因素。挪威云杉从土壤结疤中获益匪浅(P < .001),其幼苗数量甚至超过了银杉。这两个树种都在植物可用水量大的土壤中茁壮成长。相比之下,幼苗数量总体上受到光照的负面影响。此外,我们还发现,在与邻树距离较远的地方和不含碳酸盐的土壤中,银冷杉的再生密度更高。我们的研究表明,在研究地区,限制挪威云杉和银杉幼苗生长的因素是水,而不是光。在无碳酸盐的深层细粒土壤中,银杉的播种可能在完整树冠下最为有效,同时还能最大限度地拉近与挪威云杉的距离。总之,银杉播种可以优化成为一种有效的适应措施,使挪威云杉单一种植多样化,从而创造出更有抵抗力和复原力的森林生态系统。
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引用次数: 0
When economically optimal is ecologically complicated: modeling tree-by-tree cutting decisions to maximize financial returns from northern hardwood stands 当经济最优时生态却很复杂:建立逐树砍伐决策模型,使北方阔叶林的经济收益最大化
IF 2.8 2区 农林科学 Q1 FORESTRY Pub Date : 2024-01-17 DOI: 10.1093/forestry/cpad067
John D Foppert, Neal F Maker
This study challenges a long-standing and often uncontested assertion in the forestry discourse that maximizing financial returns always requires ecologically simplified stands. We developed a high-resolution simulation tool for northern hardwood stands in eastern North America and integrated advanced numerical optimization methods to model the tree-level harvest decisions that maximize financial returns. We modeled each individual tree’s growth and its probability of natural mortality, conditioned on the evolving neighborhood-scale competitive environment it resides in. We developed size-, species-, and grade-specific price functions to assign potential harvest revenue values to each discrete bole section of each standing tree, and we used an evolutionary search algorithm to specify the financially optimal timing of tree-by-tree removals. We modeled three different case studies, representing a broad range of northern hardwood stand conditions, including a hypothetical 50-year-old, even-aged stand and two inventoried stands in northern New York, USA, with contrasting management histories. We observed consistent results across all three cases: maximizing financial returns from northern hardwood forests requires silvicultural finesse and results in ecologically complicated stands.
这项研究对林业界长期以来的一种观点提出了质疑,这种观点认为,要实现经济收益最大化,就必须对林分进行生态简化。我们为北美东部的北部硬木林开发了一种高分辨率模拟工具,并整合了先进的数值优化方法,为实现经济收益最大化的树木级采伐决策建模。我们对每棵树的生长及其自然死亡概率进行了建模,并以其所处的不断变化的邻域竞争环境为条件。我们开发了特定大小、物种和等级的价格函数,为每棵活立木的每个离散树干部分分配潜在的采伐收益值,并使用进化搜索算法来指定逐棵树木移除的最佳经济时机。我们模拟了三个不同的案例研究,代表了广泛的北方硬木林分状况,包括一个假定树龄为 50 年、平均树龄的林分和两个位于美国纽约北部、管理历史截然不同的清查林分。我们在所有三个案例中都观察到了一致的结果:要从北部阔叶林中获得最大的经济回报,需要精细的造林技术,并导致生态复杂的林分。
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引用次数: 0
Modeling a new taper curve and form factor of tree branches using terrestrial laser scanning 利用陆地激光扫描建立树枝新锥度曲线和形状系数模型
IF 2.8 2区 农林科学 Q1 FORESTRY Pub Date : 2024-01-11 DOI: 10.1093/forestry/cpad064
Muluken N Bazezew, Lutz Fehrmann, Christoph Kleinn, Nils Nölke
Modeling branch taper curve and form factor contributes to increasing the efficiency of tree crown reconstructions: the branch taper, defined as the sequential measure of diameters along the course of the branch, is pivotal to accurately estimate key branch variables such as biomass and volume. Branch diameters or volumes have commonly been estimated from terrestrial laser scanning (TLS) based on automatized voxelization or cylinder-fitting approaches, given the whole branch length is sufficiently covered by laser reflections. The results are, however, often affected by ample variations in point cloud characteristics caused by varying point density, occlusions, and noise. As these characteristics of TLS can hardly be sufficiently controlled or eliminated in automatized techniques, we proposed a new branch taper curve model and form factor, which can be employed directly from the laser reflections and under variable point cloud characteristics. In this paper, the approach is demonstrated on primary branches using a set of TLS-derived datasets from a sample of 20 trees (six species). The results showed an R2 of 0.86 and a mean relative absolute error of 1.03 cm (29%) when validated with field-measured diameters. The approach improved the accuracy of diameter estimates for the fine branch scales (<10 cm) as compared to the quantitative structural model (QSM). Our approach also allowed branch diameter estimation for a relatively larger number of manually recognized primary branches (>85%) from point clouds when validated with panoramic images acquired simultaneously with laser scanning. Frequently used automatized crown reconstructions from QSM, on the other hand, were affected by gaps in the point clouds due to obstruction, with the crown-tops and finer branches being the most critical. Our approach reports mean form factors across the examined species of 0.35 and 0.49, with the diameters determined at 5% and 10% of the total branch length, respectively. Our approach may have the potential to produce branch volume information with reasonable accuracy from only knowing the length and respective diameter of each branch. Our model delivers a first approximation for the taper curve and form factor for primary branches but was developed on a relatively small set of samples. We believe that our approach holds the potential to improve the accuracy of the assessment of branch diameter and volume from TLS data. The approach may also be extended to other branch orders. This could expand the horizon for volumetric calculations and biomass estimates from non-destructive TLS proxies in tree crowns.
树枝锥度曲线和形状因子建模有助于提高树冠重建的效率:树枝锥度是指沿着树枝走向对直径进行的连续测量,对于准确估算生物量和体积等关键树枝变量至关重要。树枝直径或体积通常是通过陆地激光扫描(TLS)估算出来的,这种估算基于自动体素化或圆柱拟合方法,前提是整个树枝长度被激光反射充分覆盖。然而,由于点密度、遮挡物和噪声的不同而导致点云特征变化很大,往往会影响结果。由于 TLS 的这些特性很难在自动化技术中得到充分控制或消除,我们提出了一种新的树枝锥度曲线模型和形式因子,它可以在点云特性可变的情况下直接从激光反射中使用。本文使用一组从 20 棵树(6 个树种)样本中提取的 TLS 数据集,在主枝上演示了该方法。结果表明,在与实地测量直径进行验证时,R2 为 0.86,平均相对绝对误差为 1.03 厘米(29%)。与定量结构模型(QSM)相比,该方法提高了细枝尺度(<10 厘米)直径估算的准确性。在与激光扫描同时获取的全景图像进行验证时,我们的方法还可以从点云中估算出相对较多的人工识别主枝直径(>85%)。而常用的 QSM 自动树冠重建则会受到点云中因障碍物造成的间隙的影响,其中树冠顶端和较细的枝条最为关键。我们的方法报告了受检物种的平均形式因子,分别为 0.35 和 0.49,直径分别按树枝总长度的 5%和 10%确定。我们的方法有可能在只知道每个枝条的长度和各自直径的情况下,以合理的准确度得出枝条的体积信息。我们的模型提供了主枝锥度曲线和形状因子的第一近似值,但它是在一组相对较小的样本上开发出来的。我们相信,我们的方法有可能提高通过 TLS 数据评估树枝直径和体积的准确性。这种方法还可以扩展到其他枝序。这将扩大树冠非破坏性 TLS 代用数据的体积计算和生物量估算的范围。
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引用次数: 0
A method for identifying and segmenting branches of Scots pine (Pinus sylvestris L.) trees using terrestrial laser scanning 利用地面激光扫描技术对苏格兰松(Pinus sylvestris L.)树枝进行识别和分割
IF 2.8 2区 农林科学 Q1 FORESTRY Pub Date : 2023-12-05 DOI: 10.1093/forestry/cpad062
Tuomas Yrttimaa, Ville Kankare, Ville Luoma, Samuli Junttila, Ninni Saarinen, Kim Calders, Markus Holopainen, Juha Hyyppä, Mikko Vastaranta
Terrestrial laser scanning (TLS) has been adopted as a feasible technique to characterize tree stems while the characterization of trees’ branching architecture has remained less explored. In general, branching architecture refers to the spatial arrangement of branches and their characteristics that are important when exploring the eco-physiological functioning of trees or assessing tree biomass and wood quality. Our aim was to develop a point cloud processing method for identifying and segmenting individual branches from TLS point clouds. We applied a Cartesian-to-cylinder coordinate transformation and a simple morphological filtering for stem surface reconstruction and stem-branch separation. Then branch origins were identified as their intersections with the stem surface, and individual branches were segmented based on their connectivity with the branch origins. The method, implemented in MATLAB and openly available, was validated on a 0.4-ha mature and managed southern boreal forest stand. The branch identification performance was assessed based on visual interpretation of 364 randomly sampled stem sections from 100 Scots pine (Pinus sylvestris (L.)) trees that were inspected for branch identification accuracy. The results showed that the branches could only be identified up to the height where the stem could be reconstructed. For 90% of the trees, this threshold ranged between 59.3% and 81.2% relative tree heights. Branches located below this threshold were identified with a recall of 75%, a precision of 92%, and an F1-score of 0.82. Based on our study, it appears that in a managed Scots pine stand, most of the branches can be identified with the developed method for the most valuable stem part eligible for logwood. The findings obtained in this study promote the feasibility of using TLS in applications requiring detailed characterization of trees. The developed method can be further used in quantifying the characteristics of individual branches, which could be useful for biomass and wood quality assessment, for example.
地面激光扫描(TLS)是一种可行的树干表征技术,但对树木分枝结构的表征研究较少。通常,分枝建筑学是指树枝的空间排列及其特征,在探索树木的生态生理功能或评估树木生物量和木材质量时具有重要意义。我们的目标是开发一种点云处理方法,用于从TLS点云中识别和分割单个分支。我们应用了笛卡尔到圆柱的坐标变换和简单的形态学滤波来进行茎表面重建和茎枝分离。然后根据分支原点与茎表面的交点来识别分支原点,并根据分支原点与分支原点的连通性对单个分支进行分割。该方法在MATLAB中实现,并在一个0.4 ha的成熟管理的南方北方森林林分上进行了验证。通过对100棵苏格兰松(Pinus sylvestris (L.))的364个随机取样的茎段进行视觉判读,评估了树枝识别的准确性。结果表明,树枝只能被识别到可以重建茎的高度。对于90%的树木,该阈值范围在相对树高59.3%至81.2%之间。低于这个阈值的分支被识别出来,召回率为75%,准确率为92%,f1得分为0.82。根据我们的研究,在管理的苏格兰松林中,大多数分支可以用所开发的方法识别出最具价值的符合原木条件的茎部分。本研究的结果促进了在需要详细表征树木的应用中使用TLS的可行性。所开发的方法可进一步用于量化单个树枝的特征,例如,这可能对生物量和木材质量评价有用。
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引用次数: 0
Assessing the potential of synthetic and ex situ airborne laser scanning and ground plot data to train forest biomass models 评估综合和非原位机载激光扫描和地面数据训练森林生物量模型的潜力
IF 2.8 2区 农林科学 Q1 FORESTRY Pub Date : 2023-12-05 DOI: 10.1093/forestry/cpad061
Jannika Schäfer, Lukas Winiwarter, Hannah Weiser, Jan Novotný, Bernhard Höfle, Sebastian Schmidtlein, Hans Henniger, Grzegorz Krok, Krzysztof Stereńczak, Fabian Ewald Fassnacht
Airborne laser scanning data are increasingly used to predict forest biomass over large areas. Biomass information cannot be derived directly from airborne laser scanning data; therefore, field measurements of forest plots are required to build regression models. We tested whether simulated laser scanning data of virtual forest plots could be used to train biomass models and thereby reduce the amount of field measurements required. We compared the performance of models that were trained with (i) simulated data only, (ii) a combination of simulated and real data, (iii) real data collected from different study sites, and (iv) real data collected from the same study site the model was applied to. We additionally investigated whether using a subset of the simulated data instead of using all simulated data improved model performance. The best matching subset of the simulated data was sampled by selecting the simulated forest plot with the highest correlation of the return height distribution profile for each real forest plot. For comparison, a randomly selected subset was evaluated. Models were tested on four forest sites located in Poland, the Czech Republic, and Canada. Model performance was assessed by root mean squared error (RMSE), squared Pearson correlation coefficient (r$^{2}$), and mean error (ME) of observed and predicted biomass. We found that models trained solely with simulated data did not achieve the accuracy of models trained with real data (RMSE increase of 52–122 %, r$^{2}$ decrease of 4–18 %). However, model performance improved when only a subset of the simulated data was used (RMSE increase of 21–118 %, r$^{2}$ decrease of 5–14 % compared to the real data model), albeit differences in model performance when using the best matching subset compared to using a randomly selected subset were small. Using simulated data for model training always resulted in a strong underprediction of biomass. Extending sparse real training datasets with simulated data decreased RMSE and increased r$^{2}$, as long as no more than 12–346 real training samples were available, depending on the study site. For three of the four study sites, models trained with real data collected from other sites outperformed models trained with simulated data and RMSE and r$^{2}$ were similar to models trained with data from the respective sites. Our results indicate that simulated data cannot yet replace real data but they can be helpful in some sites to extend training datasets when only a limited amount of real data is available.
机载激光扫描数据越来越多地用于预测大面积的森林生物量。生物质信息不能直接从机载激光扫描数据中获得;因此,建立回归模型需要对森林样地进行实地测量。我们测试了虚拟森林样地的模拟激光扫描数据是否可以用于训练生物量模型,从而减少所需的实地测量量。我们比较了使用(i)模拟数据训练的模型的性能,(ii)模拟数据和真实数据的组合,(iii)从不同研究地点收集的真实数据,以及(iv)从模型应用的同一研究地点收集的真实数据。我们还研究了使用模拟数据的子集而不是使用所有模拟数据是否可以提高模型性能。选取与每个真实森林样地的回归高度分布曲线相关性最高的模拟森林样地作为模拟数据的最佳匹配子集。为了比较,随机选择一个子集进行评估。这些模型在波兰、捷克共和国和加拿大的四个森林地点进行了测试。模型性能通过观测和预测生物量的均方根误差(RMSE)、平方Pearson相关系数(r$^{2}$)和平均误差(ME)来评估。我们发现,仅用模拟数据训练的模型并没有达到用真实数据训练的模型的精度(RMSE增加52 - 122%,r$^{2}$减少4 - 18%)。然而,当只使用模拟数据的一个子集时,模型性能得到了改善(与真实数据模型相比,RMSE增加了21 - 118%,r$^{2}$减少了5 - 14%),尽管使用最佳匹配子集与使用随机选择的子集相比,模型性能的差异很小。使用模拟数据进行模型训练总是导致对生物量的严重低估。使用模拟数据扩展稀疏真实训练数据集降低了RMSE,增加了r$^{2}$,只要真实训练样本不超过12-346个,具体取决于研究地点。对于四个研究站点中的三个站点,使用从其他站点收集的真实数据训练的模型优于使用模拟数据训练的模型,RMSE和r$^{2}$与使用各自站点的数据训练的模型相似。我们的研究结果表明,模拟数据还不能取代真实数据,但在某些站点,当只有有限数量的真实数据可用时,模拟数据可以帮助扩展训练数据集。
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引用次数: 0
Growth and quality of 16-year-old sessile oak (Quercus petraea (Matt.) Liebl.) planted in traditional and alternative row planting patterns 16年无柄栎(Quercus petaea)的生长和品质)种植在传统和替代行种植模式
IF 2.8 2区 农林科学 Q1 FORESTRY Pub Date : 2023-11-27 DOI: 10.1093/forestry/cpad059
Tadeusz Andrzejczyk, Mateusz Liziniewicz, Leszek Bolibok
Traditional oak silviculture is costly due to high initial planting density required to obtain management goals of producing high quality timber. New methods therefore reduce the initial planting density and use a planting pattern that allows the inclusion of naturally regenerated trees of other species. The study presents the results of a 13-year experiment on the growth and quality of sessile oak planted in traditional and three alternative planting patterns. We hypothesized that increasing of initial spacing and use of natural regeneration to compensate for lower intra-specific competition does not deteriorate growth and quality parameters of oak. Breast height diameter, height, slenderness, height of the first live branch, diameter of the thickest branch, stem shape and potential future crop oak trees (PFCT) were measured and analysed. The local competition of admixture species was estimated on the basis of the relationship between the height of the oak and the height of admixture trees in its close proximity. The type of planting pattern had no significant effect on oak diameter, but oak height was greater in the traditional pattern than in alternative patterns. Height of the first live branch was the only qualitative trait that differed significantly between the compared planting patterns. The probability of an oak being selected as a PFCT was similar in traditional and alternative patterns, but PFCT absolute number was increasing with an increase of oak planting density. Analysis of competition of admixture species in alternative methods shows that it can reduce height, diameter and stability of oaks. On the other hand, the probability of trees being selected as a PFCT increases significantly with increasing interspecific competition. We conclude that, with controlled competition and at least moderate natural regeneration, the alternative planting patterns can produce oaks with similar growth and quality to those in the traditional pattern.
为了达到生产高质量木材的管理目标,需要很高的初始种植密度,传统的橡树造林成本很高。因此,新的方法减少了最初的种植密度,并使用了一种允许包括其他物种的自然再生树木的种植模式。本研究介绍了一项为期13年的无柄橡树生长和质量试验的结果,该试验以传统和三种替代种植模式种植。我们假设,增加初始间距和利用自然再生来弥补较低的种内竞争不会损害橡木的生长和质量参数。测定并分析了栎树胸高直径、栎树高、栎树长细、第一活枝高、最粗枝直径、栎树茎形和潜在未来作物(PFCT)。根据栎树高度与其附近混合树种高度的关系,估算了混合树种的局部竞争。不同种植模式对栎树直径的影响不显著,但传统模式对栎树高度的影响大于交替模式。第一活枝高度是两种种植方式间唯一存在显著差异的质量性状。在传统模式和替代模式下,栎树被选为PFCT的概率相似,但PFCT的绝对数量随着栎树种植密度的增加而增加。对不同方法下混合树种的竞争分析表明,混合树种会降低栎树的株高、株径和稳定性。另一方面,随着种间竞争的增加,树被选择为PFCT的概率显著增加。研究结果表明,在竞争可控和适度自然更新的条件下,不同种植模式下的栎树生长速度和质量与传统模式相近。
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引用次数: 0
The effects of arginine phosphate (ArGrow® Granulat) on growth of Scots pine and Norway spruce seedlings planted in varying soil layer structures simulating site preparation 磷酸精氨酸(ArGrow®Granulat)对不同土层结构模拟立地条件下种植的苏格兰松和挪威云杉幼苗生长的影响
IF 2.8 2区 农林科学 Q1 FORESTRY Pub Date : 2023-11-24 DOI: 10.1093/forestry/cpad060
Jaana Luoranen, Timo Saksa
The pressure to plant seedlings in unprepared instead of prepared soil in forest regeneration is increasing, although seedlings’ growth in unprepared soil is poorer. One way to improve seedlings’ growth could be to add arginine phosphate to the planting hole at the time of planting. In field studies, abiotic and biotic damage normally disturbs studies which seek to determine growth effects, especially in unprepared soil. The study’s aims were (i) to clarify whether it was possible to investigate the differences between varying soil layer structures mimicking site preparation methods and seedling treatments in simulated planting places in partly controlled conditions; and (ii) to investigate the effects of arginine phosphate (arGrow® Granulat) on the growth, especially root growth, of Norway spruce (Picea abies (L.) H. Karst.) and Scots pine (Pinus sylvestris L.) container seedlings in those simulated planting places. Seedlings of both tree species were planted in 40 l pots sunk partly into the soil in a sandy field. Before planting, pots were filled with soil layers mimicking layers in soil without any site preparation treatment and in spot mounds. After one growing season, the differences in growth parameters between treatments were small in Norway spruce seedlings, and no significant benefits of arginine phosphate were observed. The soil in the spruce pots was fine-textured and contained a thick organic layer. More studies in more common forest soil types suitable for Norway spruce seedlings are needed to be sure about the growth responses. In Scots pine seedlings, adding arginine phosphate granules to the planting holes improved the growth of seedlings at least in the first summer after planting in soil layers mimicking no mechanical site preparation situation, and the growth improvement may even have compensated the mounding effects. In conclusion, mimicking site preparation methods in large pots was a promising method by which the differences between site preparations methods as well as effects of seedling treatments, in our case arginine phosphate, on the growth of conifer seedlings, can be tested.
在森林更新中,幼苗在预备土壤中生长较差,但在预备土壤中种植幼苗的压力越来越大。一种改善幼苗生长的方法是在种植时向种植孔中添加精氨酸磷酸。在实地研究中,非生物和生物损害通常会干扰那些试图确定生长效应的研究,特别是在未准备好的土壤中。该研究的目的是:(i)阐明是否有可能在部分受控条件下,在模拟的种植地点调查不同土层结构模拟场地准备方法和幼苗处理之间的差异;(ii)研究磷酸精氨酸(arGrow®Granulat)对挪威云杉(Picea abies (L.))生长,特别是根系生长的影响。(H. Karst.)和苏格兰松(Pinus sylvestris L.)的容器苗。两种树种的幼苗种植在40 l盆中,部分埋在沙土中。在种植前,在花盆里填满土层,模仿土壤中的土层,没有任何现场准备处理和现场土堆。经过一个生长季节后,挪威云杉幼苗的生长参数在处理之间的差异很小,精氨酸磷酸没有观察到显著的效益。云杉花盆里的土壤质地细腻,含有一层厚厚的有机土层。需要对适合挪威云杉幼苗的更常见的森林土壤类型进行更多的研究,以确定其生长反应。在苏格兰松幼苗中,在种植孔中添加磷酸精氨酸颗粒,至少在种植后的第一个夏天,在模拟无机械立地情况的土层中,幼苗的生长得到改善,甚至可能补偿了堆积效应。综上所述,在大型盆栽中模拟立地制备方法是一种很有前途的方法,通过这种方法可以测试不同立地制备方法的差异以及幼苗处理(以精氨酸磷酸酯为例)对针叶树幼苗生长的影响。
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