Intercity human mobility networks exhibit pronounced temporal heterogeneity, yet the structural and mechanistic reconfigurations that occur during high-demand episodes such as national holidays remain underexplored, particularly regarding their integrated dynamics. Addressing this gap, this study examines how China’s intercity mobility network reorganizes across regular and holiday periods through a multi-scale analytical framework that integrates network structural analysis with relational modeling. Drawing on nationwide location-based services data, the findings reveal a fundamental reconfiguration: the macro-scale network backbone shifts focus from established economic and administrative corridors toward alternative regional transport hubs, while meso-scale organization transitions from a dual-core hierarchy dominated by national centers to a more dispersed, tripolar pattern comprising supra-regional communities. This structural adaptation occurs alongside a significant recalibration of spatial interaction mechanisms: distance friction intensifies, economic complementarity such as income weakens, while non-economic factors like cultural proximity gain prominence, and notably, housing prices shift from deterrents to attractors. Synthesizing these results, the study demonstrates the coordinated evolution of network structure and relational mechanisms, revealing the temporally conditioned nature of spatial interaction principles. These insights challenge static frameworks, highlight the adaptive capacity of urban systems, and underscore the need for context-sensitive transport planning.
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