In Bagaria (J Symb Log 81(2), 584–604, 2016), Bagaria and Väänänen developed a framework for studying the large cardinal strength of downwards Löwenheim-Skolem theorems and related set theoretic reflection properties. The main tool was the notion of symbiosis, originally introduced by the third author in Väänänen (Applications of set theory to generalized quantifiers. PhD thesis, University of Manchester, 1967); Väänänen (in Logic Colloquium ’78 (Mons, 1978), volume 97 of Stud. Logic Foundations Math., pages 391–421. North-Holland, Amsterdam 1979) Symbiosis provides a way of relating model theoretic properties of strong logics to definability in set theory. In this paper we continue the systematic investigation of symbiosis and apply it to upwards Löwenheim-Skolem theorems and reflection principles. To achieve this, we need to adapt the notion of symbiosis to a new form, called bounded symbiosis. As one easy application, we obtain upper and lower bounds for the large cardinal strength of upwards Löwenheim–Skolem-type principles for second order logic.