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Micro-CT analysis of Katian radiolarians from the Malongulli Formation, New South Wales, Australia, and implications for skeletogenesis 澳大利亚新南威尔士州Malongulli组Katian放射虫的显微ct分析及其对骨骼形成的意义
IF 1.4 4区 地球科学 Q3 PALEONTOLOGY Pub Date : 2023-04-05 DOI: 10.1017/jpa.2023.13
A diverse and well-preserved radiolarian assemblage from the Malongulli Formation, New South Wales, Australia, contains 13 species representing 10 genera and six families. One new genus, Wiradjuri, is introduced to accommodate pre-Devonian single-shelled entactiniid taxa, and one new species, Secuicollacta malongulliensis, is recorded together with some previously described forms. The microstructures of the “rotasphaerid structure/primary unit” and the “ectopic spicule” are investigated to validate their roles as fundamental units in the Secuicollactidae, together with comprehensive documentation of the previously enigmatic Pseudorotasphaera internal skeleton. The results of this investigation suggest that, among all radiolarian genera that survived the Late Ordovician Mass Extinction event (LOME) and transitioned into the Silurian, Secuicollacta, Haplotaeniatum, and Palaeoephippium maintained stable body plans during the transition and were more successfully established. The selective advantages these lineages had during the LOME were most likely spontaneous outcomes of the mode of structural development involving sequential skeletogenesis and a tendency to evolve toward simpler body plans. UUID: http://zoobank.org/264c026f-35af-4025-a4f7-de8f9a91f8b9
澳大利亚新南威尔士州Malogulli组的一个多样且保存完好的放射虫群落,包含13种,代表6科10属。引入了一个新属Wiradjuri,以适应前泥盆纪的单壳五足类分类群,并记录了一个新种Securicollacta malogulliensis,以及一些先前描述的形式。研究了“轮状体结构/主要单元”和“异位针状体”的微观结构,以验证它们作为Securicollactidae中基本单元的作用,并对以前神秘的假轮状体内部骨骼进行了全面的记录。研究结果表明,在晚奥陶世大灭绝事件(LOME)中幸存下来并过渡到志留纪的所有放射虫属中,Securicollacta属、Haplotaeniatum属和Palaeephippium属在过渡期间保持了稳定的身体计划,并更成功地建立起来。这些谱系在LOME期间具有的选择性优势很可能是结构发育模式的自发结果,包括顺序骨骼发育和向更简单的身体计划进化的趋势。UUID:http://zoobank.org/264c026f-35af-4025-a4f7-de8f9a91f8b9
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引用次数: 0
Crinoid calyx origin from stem radial echinoderms 来源于茎干放射状棘皮动物的蛛网膜肾盏
IF 1.4 4区 地球科学 Q3 PALEONTOLOGY Pub Date : 2023-03-29 DOI: 10.1017/jpa.2023.14
T. Guensburg, R. Mooi, N. Mongiardino Koch
Evidence from the earliest-known crinoids (Tremadocian, Early Ordovician), called protocrinoids, is used to hypothesize initial steps by which elements of the calyx evolved. Protocrinoid calyces are composed of extraxial primary and surrounding secondary plates (both of which have epispires along their sutures) that are unlike those of more crownward fossil and extant crinoids in which equivalent calycinal plating is strongly organized. These reductions inspired several schemes by which to name the plates in these calyces. However, the primary-secondary systems seen in protocrinoids first appeared among Cambrian stem radial echinoderms, with primaries representing centers around which secondaries were sequentially added during ontogeny. Therefore, the protocrinoid calyx represents an intermediate condition between earliest echinoderms and crownward crinoids. Position and ontogeny indicate certain primaries remained as loss of secondaries occurred, resulting in abutting of primaries into the conjoined alternating circlets characteristic of crinoids. This transformative event included suppression of secondary plating and modification or, more commonly, elimination of respiratory structures. These data indicate subradial calyx plate terminology does not correspond with most common usage, but rather, supports an alternative redefinition of these traditional expressions. Extension and adoral growth of fixed rays during calyx ontogeny preceded conjoined primaries in earliest crinoids. Restriction with modification or elimination of calyx respiratory structures also accompanied this modification. Phylogenetic analyses strongly support crinoid origination from early pentaradiate echinoderms, separate from blastozoans. Accordingly, all Tremadocian crinoids express a distinctive aggregate of plesiomorphic and apomorphic commonalities; all branch early within the crinoid clade, separate from traditional subclass-level clades. Nevertheless, each taxon within this assemblage expresses at least one diagnostic apomorphy of camerate, cladid, or disparid clades.
来自已知最早的海百合(早奥陶世的Tremadocian)的证据,被称为原海百合,被用来假设花萼元素进化的最初步骤。原海百合花萼由外部初级板和周围的次级板组成(两者的缝合处都有针尖),这与更靠后的化石和现存海百合花萼不同,后者的等效花萼板组织强烈。这些减少启发了几种命名这些花萼中的板块的方案。然而,原海百合的初生-次生系统最早出现在寒武纪茎放射状棘皮动物中,初生系统代表在个体发育过程中依次添加次生系统的中心。因此,原海百合花萼代表了最早的棘皮动物和冠状海百合之间的中间状态。位置和个体发育表明,当次生损失发生时,某些原发植物仍然存在,导致原发植物接近成海百合特征的连体交替循环。这一变革性事件包括抑制二次电镀和修饰,或者更常见的是消除呼吸结构。这些数据表明,下径向萼板术语不符合最常见的用法,而是支持这些传统表达的另一种重新定义。在最早的海百合中,在花萼形成过程中,固定射线的延伸和顺口生长先于连体初生。改变或消除花萼呼吸结构的限制也伴随着这种改变。系统发育分析有力地支持海百合起源于早期的五源棘皮动物,与囊胚动物分离。因此,所有的Tremadocian海百合都表现出一种独特的半胚性和非胚性共性的集合;所有分支早期在海百合分支中,与传统的亚类水平分支分开。然而,这个组合中的每一个分类单元至少表现出一种照相机、有毛枝或分裂枝的诊断性形像。
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引用次数: 0
JPA volume 97 S92 Cover and Front matter JPA第97卷S92封面和封面
IF 1.4 4区 地球科学 Q3 PALEONTOLOGY Pub Date : 2023-03-23 DOI: 10.1017/jpa.2023.16
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引用次数: 0
Paleogeographical and paleoenvironmental significance of ostracodes from the Pennsylvanian Nagaiwa Formation, northeast Japan 日本东北部宾夕法尼亚-永洼组介形类的古地理和古环境意义
IF 1.4 4区 地球科学 Q3 PALEONTOLOGY Pub Date : 2023-03-23 DOI: 10.1017/jpa.2022.108
G. Tanaka
Abstract. The Early Pennsylvanian Nagaiwa Formation contains fossils such as corals, fusulinids, and ostracodes, and its age and depositional environments have been determined by fusulinids and sedimentology. In this study, I describe the ostracode assemblages from the Nagaiwa Formation. Moreover, I provide a reconstruction of the paleogeography of northeastern Japan during the Early Pennsylvanian by comparing this ostracode assemblage with assemblages from other regions during the same period. Thirty ostracode species, including 12 genera, have been identified, most of which are endemic species and 10 of which are new: Jordanites michinokuensis n. sp., Thuringobolbina ikeyai n. sp., Aechmina iwatensis n. sp., Pseudobythocypris asiatica n. sp., P. zipangu n. sp., P. siveteri n. sp., Platyrhomboides tohokuensis n. sp., P. japonica n. sp., Healdia ofunatensis n. sp., and H. rikutyuensis n. sp. Two of these species are also found in central Japan. The ostracodes from the Nagaiwa Formation are unique when compared with any other similarly aged assemblages.
摘要宾夕法尼亚早期的Nagaiwa组包含珊瑚、褐藻和介形虫等化石,其年龄和沉积环境已由褐藻和沉积学确定。在这项研究中,我描述了长岩组的介形虫组合。此外,我通过将这种介形虫组合与同一时期其他地区的组合进行比较,重建了宾夕法尼亚纪早期日本东北部的古地理。已鉴定出介形虫30种,包括12属,其中大多数是特有种,10种是新的:约旦虫(Jordanites michinokuensis n.sp.)。,和H.rikutyuensis n.sp.。其中两个物种也在日本中部发现。与任何其他年龄相似的组合相比,长岩组的介形虫是独特的。
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引用次数: 0
Nonmarine ostracod fauna from the Lower Cretaceous Shinekhudag Formation (southwest Mongolia): taxonomy, biostratigraphy, and paleoecology 蒙古西南部下白垩统Shinekhudag组非海相介形类动物群:分类、生物地层学和古生态学
IF 1.4 4区 地球科学 Q3 PALEONTOLOGY Pub Date : 2023-03-03 DOI: 10.1017/jpa.2023.5
Byung-Do Choi, Yaqiong Wang
Abstract. This work provides the detailed investigation (taxonomy, biostratigraphy, and paleoecology) of a nonmarine ostracod fauna from the Shinekhudag Formation in Gobi–Altai area, southwest Mongolia. The samples from two sections (Tsagaan Tsuvarga and Oshih Hollow East) yielded various ostracods assigned to nine species belonging to six genera: Cypridea verrucata Neustrueva, 1974, C. ihsienensis Hou, 1958, C. unicostata Galeeva, 1955, C. tumefacta Neustrueva, 1974, Yumenia cf. Y. oriformis Hou, 1958, Scabriculocypris subscalara Zhang and Chen in Ye et al., 2003, ?Trapezoidella sp., Candona sp., and Vlakomia ulanense Neustrueva, 1977. The biostratigraphic application of our taxonomic results suggests that the age of the Shinekhudag Formation is Barremian–Aptian. In addition, the ostracod fauna shows strong affinities to faunas from northeast China and potentially northwest China, providing evidence of faunal exchanges between these regions and southwest Mongolia. The new discovery of Vlakomia ulanense indicates that mid-Cretaceous (Albian) species in northeast China probably originated in Mongolia at the time of deposition of the Shinekhudag Formation. Two ostracod assemblages in the studied sections reflect that different paleoenvironment settings have existed during deposition; the Tsagaan Tsuvarga Section represents a shallow-littoral zone of athalassic (inland) saline lake under arid climate, and the dominance of cypridoideans of the Oshih Hollow East Section is indicative of ephemeral water bodies.
摘要本工作对蒙古西南戈壁-阿尔泰地区Shinekhudag组的一个非海洋介形虫动物群进行了详细的调查(分类学、生物地层学和古生态学)。来自两个剖面(Tsagaan Tsuvarga和Oshih Hollow East)的样本产生了各种介形虫,分为六属九种:Cypridea verrucata Neustrueva,1974,C.ihsienensis Hou,1958,C.unicostata Galeeva,1955,C.tumefacta Neustruva,1974,Yumenia cf.Y.oriformis Hou,1956,Scabriculocypis subscalara Zhang和Chen in Ye et al.,2003?Trapezoidella sp.、Candona sp.和Vlakomia ulanense Neustrueva,1977年。我们的分类学结果的生物地层学应用表明,Shinekhudag组的年龄为Barremian-Aptian。此外,介形虫动物群与中国东北和可能的中国西北的动物群表现出强烈的亲缘关系,为这些地区与蒙古西南部之间的动物群交流提供了证据。新发现的乌兰藻表明,我国东北地区白垩纪中期(阿尔比阶)物种可能起源于Shinekhudag组沉积时的蒙古。研究剖面中的两个介形虫组合反映了沉积过程中存在不同的古环境背景;Tsagaan-Tsuvarga剖面代表了干旱气候下古典(内陆)盐湖的浅水沿岸带,而Oshih Hollow东段的塞浦路斯类占主导地位,表明存在短暂水体。
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引用次数: 2
New insight into Cenozoic Orbitestellidae (Gastropoda: Heterobranchia) from the Magellanic Region based on lower Neogene and Recent species 麦哲伦地区新生代眶足目(腹足目:异鳃亚目)的新认识
IF 1.4 4区 地球科学 Q3 PALEONTOLOGY Pub Date : 2023-03-02 DOI: 10.1017/jpa.2023.8
Javier Di Luca, M. Griffin, G. Pastorino
Abstract. Patagorbitestella new genus is here proposed to include two extant and one fossil species: P. ponderi (Linse, 2002) new combination, P. patagonica (Simone and Zelaya, 2004) new combination, and P. leonensis new species, the last described from the Punta Entrada Member of Monte León Formation (50°21′25.4”S, 68°53′05.9″W, Aquitanian to lower Burdigalian, lower Miocene). A protoconch sculptured with distinctive microscopic spiral threads serves as a per se diagnostic shell character for the new genus. Patagorbitestella n. gen. constitutes a distinctive lineage of orbitestellid gastropods inhabiting exclusively the Magellanic Region at least since the early Miocene. This is the first fossil record of Orbitestellidae in South America.
摘要Patagorbitestella新属在这里被提议包括两个现存的和一个化石物种:黄杨(Linse,2002)新组合、巴塔哥尼亚杨(Simone和Zelaya,2004)新组合和里昂杨新物种,最后一个描述于蒙特莱昂组的Punta Entrada成员(50°21′25.4“S,68°53′05.9”W,阿基坦阶至下布尔迪加阶,下中新世)。一个由独特的微观螺旋线雕刻而成的原球茎,本身就是这个新属的诊断外壳特征。Patagorbitestella n.gen.构成了一个独特的轨道星腹足类谱系,至少自中新世早期以来,它只生活在麦哲伦地区。这是南美洲第一个轨道虫科的化石记录。
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引用次数: 0
JPA volume 97 issue 2 Cover and Front matter JPA第97卷第2期封面和封面事项
IF 1.4 4区 地球科学 Q3 PALEONTOLOGY Pub Date : 2023-03-01 DOI: 10.1017/jpa.2023.18
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引用次数: 0
Systematic paleontology of macroalgal fossils from the Tonian Mackenzie Mountains Supergroup 托尼安-麦肯齐山脉超群大型藻类化石的系统古生物学
IF 1.4 4区 地球科学 Q3 PALEONTOLOGY Pub Date : 2023-03-01 DOI: 10.1017/jpa.2023.4
Katie M. Maloney, Dakota P. Maverick, J. Schiffbauer, G. Halverson, S. Xiao, M. Laflamme
Abstract. Proterozoic eukaryotic macroalgae are difficult to interpret because morphological details required for proper phylogenetic studies are rarely preserved. This is especially true of morphologically simple organisms consisting of tubes, ribbons, or spheres that are commonly found in a wide array of bacteria, plants, and even animals. Previous reports of exceptionally preserved Tonian (ca. 950–900 Ma) fossils from the Dolores Creek Formation of Northwestern Canada feature enough morphological evidence to support a green macroalgal affinity. However, the affinities of two additional forms identified on the basis of the size distribution of available specimens remain undetermined, while the presence of three unique algal forms supports other reports of increasing algal diversity in the early Neoproterozoic. Archaeochaeta guncho new genus new species is described as a green macroalga on the basis of its well-preserved morphology consisting of an unbranching, uniseriate thallus with uniform width throughout and possessing an elliptical to globose anchoring holdfast. A larger size class of ribbon-like forms is interpreted as Vendotaenia sp. A third size class is significantly smaller than Archaeochaeta n. gen. and Vendotaenia, but in the absence of clear morphological characters, it remains difficult to assign. As Archaeochaeta n. gen. and Vendotaenia represent photoautotrophic taxa, these findings support the hypothesis of increasing morphological complexity and phyletic diversification of macroalgae during the Tonian, leading to dramatic changes within benthic marine ecosystems before the evolution of animals.
摘要元古代真核巨藻很难解释,因为系统发育研究所需的形态学细节很少被保存下来。对于由管状、带状或球形组成的形态简单的生物体来说尤其如此,这些生物体通常存在于各种各样的细菌、植物甚至动物中。先前关于加拿大西北部Dolores Creek组保存完好的Tonian(约950-900 Ma)化石的报道,有足够的形态学证据支持绿色大藻的亲缘关系。然而,根据现有标本的大小分布确定的另外两种形式的亲和力仍未确定,而三种独特的藻类形式的存在支持了新元古代早期藻类多样性增加的其他报告。古毛藻(Archaeochaeta guncho)新属新种被描述为一种保存完好的绿色大藻,其形态由无分支、单列的菌体组成,整个菌体具有均匀的宽度,并具有椭圆形到球形的锚定支架。带状形态的一个较大的尺寸类被解释为Vendotaenia sp.。第三个尺寸类明显小于Archaeochaeta n. gen.和Vendotaenia,但由于缺乏明确的形态特征,仍然难以确定。由于Archaeochaeta n. gen和Vendotaenia是光自养分类群,这些发现支持了大型藻类在Tonian时期形态复杂性和种类多样化的增加,导致底栖海洋生态系统在动物进化之前发生了巨大变化的假设。
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引用次数: 1
Typhlocybinae leafhoppers (Hemiptera, Cicadellidae) from Eocene Rovno amber reveal a transition in wing venation and a defensive adaptation 始新世Rovno琥珀中斑叶蝉科叶蝉(半翅目,斑叶蝉科)揭示了翅脉的转变和防御性适应
IF 1.4 4区 地球科学 Q3 PALEONTOLOGY Pub Date : 2023-03-01 DOI: 10.1017/jpa.2023.3
C. Dietrich, S. A. Simutnik, E. Perkovsky
Abstract. The first fossil Typhlocybinae inclusions from Eocene Rovno amber are described and illustrated. They include two new monotypic genera of the extinct tribe Protodikraneurini, Retrorsotettix n. gen. with type species R. vlaskini n. sp. and Protoparallaxis n. gen. with type species P. clavatus n. sp. Also described and illustrated are two new monotypic genera of the extant tribe Dikraneurini, Eodikraneura n. gen. with type species E. obscura n. sp. and Rovnodikra n. gen. with type species Rovnodikra longipes n. sp. Retrorsotettix is the oldest leafhopper known to exhibit a false eyespot and false leg markings on the forewing, representing an early acquisition of a defensive strategy against visual predators. Appearance of a small insect with false eyespots in the Eocene fossil record may reflect increased pressure by visual predators, especially crown ornithuromorph insectivorous birds. Such birds and small insect prey with false eyespots remain unknown in the Cretaceous. Eodikraneura exhibits a unique condition of the hind wing venation in which radius posterior and media anterior veins are confluent for a short distance but then diverge before separately reaching the submarginal vein. This presumably represents a morphological transition between the Protodikraneurini, in which hind-wing radius posterior and media anterior veins are completely separate and connected by a radial-medial crossvein, and Dikraneurini, in which radius posterior and media anterior veins are completely confluent distally. A key to genera of Protodikraneurini is provided.
摘要对始新世罗夫诺琥珀中的第一个Typhlocybinae包裹体化石进行了描述和说明。它们包括已灭绝部落Protodikraniurini的两个新的单型属,Retrorsotetix n.gen.与模式种R.vlaskini n.sp.和Protoparalaxis n.gen.和模式种P.clavatus n.sp.。还描述和说明了现存部落Dikraniurii的两个新单型属,Eodikraeneura n.gen.与模式种E.obscura n.sp.和Rovnodikra n.gen.,与模式种Rovnodikara longipes n.sp.Retrorsotetix是已知最古老的叶蝉,前翅上有假眼点和假腿纹,代表着早期获得了对抗视觉捕食者的防御策略。始新世化石记录中出现的一种带有假眼点的小型昆虫可能反映了视觉捕食者的压力增加,尤其是冠鸟形食虫鸟。在白垩纪,这种带有假眼点的鸟类和小型昆虫猎物仍然不为人知。Eodikraeneura表现出一种独特的后翅静脉状况,其中桡骨后静脉和中前静脉在短距离内汇合,但在分别到达缘下静脉之前分叉。这可能代表了原始脊神经和脊神经之间的形态转变,前者的后翼桡骨后静脉和中前静脉完全分离,并通过径向-内侧交叉静脉连接,后者的桡骨后静脉与中前静脉在远端完全汇合。本文提供了一个原双脊藻属的检索表。
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引用次数: 2
First Report of Sphenothallus Hall (Cnidaria, Medusozoa) from the Mesozoic Erathem (Upper Triassic, Slovenia) 中生代Erathem(上三叠纪,斯洛文尼亚)鞘翅目Hall(Cnidaria,Medusozoa)的首次报告
IF 1.4 4区 地球科学 Q3 PALEONTOLOGY Pub Date : 2023-02-16 DOI: 10.1017/jpa.2023.1
Heyo Van Iten, R. Gašparič, T. Hitij, T. Kolar-Jurkovšek, B. Jurkovšek
Sphenothallus Hall, 1847, one of the most widely distributed and longest ranging genera in the fossil record, has been documented from all systems of the Paleozoic Erathem except the Permian (Table 1), although it has been stated (e.g., Choi, 1990; Bolton, 1994; Fatka et al., 2012) that the genus also occurs in that system. At present the first appearance of this epibenthic, polypoid medusozoan cnidarian lies in Cambrian Stage 3, while the previously known youngest occurrences are in the Pennsylvanian System. Sphenothallus has been found in numerous formations on all continents except Australia and Antarctica. It occurs in a variety of marine facies ranging from shallow nearshore to deep offshore and has even been found in strata of coastal lacustrine origin, probably as an allochthonous element (Lerner and Lucas, 2011). Many of the rock units known to contain Sphenothallus also contain conulariids (Table 1), an extinct group of marine scyphozoans that may have been closely related to Sphenothallus (Van Iten et al., 1992, 1996). Van Iten et al. (1992) interpreted Sphenothallus as a medusozoan cnidarian of uncertain class-level affinities, but later Dzik et al. (2017) documented internal peridermal structures that may be homologous to similar features in the periderm of coronate scyphozoans (see for example illustrations in Van Iten, 1992, and Van Iten et al., 1996).
Sphenothallus Hall,1847,是化石记录中分布最广、范围最长的属之一,除二叠纪外,古生代的所有系统都有记录(表1),尽管有人指出(例如,Choi,1990;Bolton,1994;Fatka等人,2012)该属也出现在该系统中。目前,这种表海底、息肉状水母类的首次出现在寒武纪第3阶段,而之前已知的最年轻的出现在宾夕法尼亚系。除澳大利亚和南极洲外,在所有大陆的许多地层中都发现了鞘翅目化石。它存在于从浅近海到深近海的各种海洋相中,甚至在沿海湖泊成因的地层中发现,可能是一种外来元素(Lerner和Lucas,2011)。许多已知含有鞘翅目的岩石单元也含有圆锥虫(表1),圆锥虫是一种已灭绝的海洋镰刀形目动物,可能与鞘翅目有密切关系(Van Iten等人,19921996)。Van Iten等人(1992)将Sphenothallus解释为具有不确定类水平亲缘关系的水母类,但后来Dzik等人(2017)记录了内部皮周结构,这些结构可能与冠状病毒scyphozoans的皮周中的类似特征同源(例如,见Van Iten,1992和Van Iten et al.,1996中的插图)。
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引用次数: 1
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Journal of Paleontology
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