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Lagomorphs (Mammalia) from the early Pliocene of Dorkovo, Bulgaria 保加利亚多尔科沃上新世早期的Lagomorphs(哺乳类)
Q3 Earth and Planetary Sciences Pub Date : 2020-01-01 DOI: 10.37520/fi.2020.007
S. Sen
The vertebrate locality of Dorkovo yielded a rich mammalian assemblage of which some large mammals have been studied, but not smaller mammals. The previous investigations correlated this fauna to the early Pliocene (early Ruscinian, MN 14). The present paper describes lagomorph remains which consist of one fragment of mandible and 106 isolated teeth. Four species are identified: Ochotona cf. antiqua, Prolagus sorbinii, Alilepus spassovi n. sp. and Trischizolagus crusafonti. The new species is characterised by the derived pattern of its p3 which has trigonid more stretched, enamel infolding stronger than in any other species of Alilepus, and the p3 lacks anteroflexid. The presence of four species of lagomorphs at Dorkovo is in agreement with the diversity dynamics of lagomorphs toward the end of late Miocene and during the early Pliocene in Eurasia, and in particular in Eastern Europe. The time range of the Dorkovo lagomorphs encompasses the Miocene/Pliocene boundary, and this is in agreement with the previous age assignments, i.e., earliest Pliocene, or early part of MN 14 mammalian zone.
Dorkovo的脊椎动物种群产生了丰富的哺乳动物组合,其中一些大型哺乳动物被研究过,但没有研究过小型哺乳动物。先前的研究将该动物群与上新世早期(早俄罗斯,MN 14)联系起来。本文描述了由一块下颌骨碎片和106颗离体牙齿组成的lagomorph遗骸。已鉴定出4种:古斑蝽、sorbinii斑蝽、Alilepus spassovi n. sp.和Trischizolagus crusafonti。新物种的特征是其p3的衍生模式,其三角肌更拉伸,牙釉质内折叠比任何其他种类的阿利普斯都更强,p3缺乏前弯曲。在多尔科沃发现的4种lagomorphate与欧亚大陆,特别是东欧地区中新世晚期和上新世早期lagomorphate的多样性动态一致。Dorkovo lagomorphs的时间范围包括中新世/上新世的分界线,这与以往的年龄划分一致,即上新世早期或MN 14哺乳动物带早期。
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引用次数: 3
A new stereospondylomorph, Korkonterpeton kalnense gen. et sp. nov., from lower Permian of the Czech Krkonoše Piedmont Basin and a redescription of Intasuchus silvicola from the lower Permian of Russia (Temnospondyli, Amphibia) 捷克Krkonoše Piedmont盆地下二叠统Korkonterpeton kalnense gen. et sp. nov.新立体海绵形态及俄罗斯下二叠统Intasuchus silvicola的再描述(Temnospondyli,两栖目)
Q3 Earth and Planetary Sciences Pub Date : 2020-01-01 DOI: 10.37520/fi.2020.019
R. Werneburg, S. Štamberg, J. Steyer
A new temnospondyl, Korkonterpeton kalnense gen. et sp. nov., is described on the basis of an almost complete specimen from the lover Permian Prosečné Formation of the Czech Krkonoše Piedmont Basin. The exceptional preservation of the holotype allows a detailed description of the cranial and postcranial characters: Korkonterpeton kalnense gen. et sp. nov. is characterized by the presence of vomerine fangs located anteriorly to the posterior choana, much longer than wide tabulars – as long as the supratemporals, a deeply concave posterior margin of the skull table, extraordinarily elongated exoccipitals, anteriorly widened pterygoids and a parasphenoid cultriform process swollen at its mid-length. In order to clarify the relationship between this new taxon and Intasuchus silvicola Konzhukova, 1956 from the lower Permian (late Kungurian) of the Russian Inta Basin, the latter is also redescribed here: Intasuchus silvicola exhibits a lacrimal entering the septomaxilla, a widened jugal, a posterolaterally widened interpterygoid vacuity, a subtemporal window elongated anteriorly up to the anterior orbital region, an elongated basipterygoid ramus which is as long as the orbit width, a vomer with a parachoanal tooth row, and an ectopterygoid of about the same length as the palatine. The description of Korkonterpeton kalnense gen. et sp. nov. and the revision of Intasuchus silvicola complete our knowledge on the origin and evolution of early stereospondylomorph temnospondyls during the lower Permian.
一种新的temnospondyl, Korkonterpeton kalnense gen. et sp. nov.,是在捷克Krkonoše Piedmont盆地的lover Permian prose n组的一个几乎完整的标本的基础上描述的。完整型的特殊保存使得对颅骨和颅骨后特征的详细描述成为可能:Korkonterpeton kalnense gen. et p. 11 .的特征是位于后颈前部的毒牙,比宽的表骨长得多——与颞上一样长,颅骨表的后缘深凹,枕外骨异常延长,翼状骨向前加宽,副蝶状突在其中部肿胀。为了澄清该新分类群与1956年俄罗斯印塔盆地下二叠世(昆古里晚期)的Intasuchus silvicola Konzhukova的关系,对后者也进行了重新描述:silvicola的特征是泪道进入鼻中隔腋窝,颧部加宽,翼状间隙后外侧加宽,颞下窗向前延伸至眶前区,基底翼状分支延长至眶宽,翼状旁齿排,翼状外突与腭约等长。Korkonterpeton kalnense gen. et sp. 11 .的描述和Intasuchus silvicola的修正完善了我们对下二叠世早期立体脊柱体temnospondyls的起源和演化的认识。
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引用次数: 2
Independent water vole (Mimomys savini, Arvicola: Rodentia, Mammalia) lineages in Italy and Central Europe 独立的水田鼠(Mimomys savini, Arvicola:啮齿目,哺乳类)在意大利和中欧的血统
Q3 Earth and Planetary Sciences Pub Date : 2020-01-01 DOI: 10.37520/fi.2020.005
F. Masini, L. Maul, L. Abbazzi, D. Petruso, A. Savorelli
Water voles are important key fossils of the Quaternary. Given their wide distribution, regional differences were expected to exist in different areas. Early hints on possible independent evolutionary trends of water voles in Italy came from palaeontology and specifically from the comparison of enamel differentiation (SDQ value) of the first lower molars between specimens from Italy and Germany. The data available at that time indicated that in the early Middle Pleistocene there were only minor enamel differences between first lower molars of water voles from these two geographical regions, whereas from the late Middle Pleistocene onwards, two lineages were clearly distinguished. Examination of mitochondrial DNA of extant Arvicola populations from across Europe by Wust-Saucy led to a similar conclusion that Arvicola populations from the Italian Peninsula had undergone independent evolution during the last 0.2 million years. The inclusion of new fossil and extant Arvicola samples from Italy and Central Europe, together with the examination of additional morphological parameters of the occlusal surface (so-called van der Meulen indexes), have provided further support for the proposed evolutionary pattern. The combined analysis of length, SDQ and A/L index reveal a certain degree of intercorrelation and indicate an essentially continuous evolutionary trend. However, variations are discernible, related to the age and geographical origins of the samples, and become more clearly seen at least since the beginning of the Late Pleistocene. Italian samples have a characteristic tendency to grow larger, elongate the anteroconid, and have less derived SDQ. This corroborates the suggestion that Italian water voles underwent an evolutionary history distinct from that of their Central European counterparts. The differences in morphology may be related to a combination of etho-/ecological (aquatic or terrestrial habits) and palaeobiogeographical factors.
水鼠是第四纪重要的关键化石。由于它们分布广泛,预计在不同地区会存在区域差异。关于意大利水田鼠可能独立进化趋势的早期线索来自古生物学,特别是来自意大利和德国标本的第一下磨牙牙釉质分化(SDQ值)的比较。当时的资料表明,在中更新世早期,这两个地理区域的水田鼠第一下磨牙之间的牙釉质差异很小,而从中更新世晚期开始,两个谱系之间的牙釉质差异明显。Wust-Saucy对来自欧洲各地现存的Arvicola种群的线粒体DNA进行了检查,得出了类似的结论,即意大利半岛的Arvicola种群在过去的20万年中经历了独立的进化。来自意大利和中欧的新化石和现存的阿维科拉样本,以及对咬合表面的额外形态学参数(所谓的van der Meulen指数)的检查,为提出的进化模式提供了进一步的支持。对长度、SDQ和A/L指数的综合分析显示出一定程度的相关关系,并显示出基本连续的演化趋势。然而,变化是可辨的,与样品的年龄和地理来源有关,并且至少从晚更新世开始变得更加清晰。意大利的样本有一个特点,即长得更大,延长前结肠,并有较少的衍生SDQ。这证实了意大利水田鼠经历了与中欧同类不同的进化史。形态上的差异可能与动物/生态(水生或陆生习性)和古地理因素的结合有关。
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引用次数: 5
Allosorex stenodus Fejfar, 1966 (Eulipotyphla, Soricidae): re-description of type material and re-interpretation of its fossil record 异位巨齿兽,1966(巨齿兽总目,巨齿兽科):类型材料的重新描述及其化石记录的重新解释
Q3 Earth and Planetary Sciences Pub Date : 2020-01-01 DOI: 10.37520/fi.2020.006
O. Fejfar, W. Koenigswald, M. Sabol
The original fossil record of Allosorex stenodus Fejfar, 1966 from Ivanovce (late Ruscinian, MN 15b) is redescribed and supplemented by the description of so far unpublished fossil remains, including upper dentition (P4 and M3) and a humerus fragment as well as the enamel microstructure analysis of m2. Based on the critical evaluation of other fossil remains from sites in Romania, Hungary, and France, the Ivanovce fossils are so far the only unquestionable recorded finds of A. stenodus in Europe, which can be considered as a local early Pliocene (MN 15) endemic species. Its occurrence is connected with the forested karst area along a broad valley of “pre-Váh” River, ecologically and climatically resembling the environment of modern karst areas in south-eastern Asia. Based on enamel microstructure analysis and unique mandible and dentition morphological characters, a separate taxonomical status of the species is also discussed.
对1966年来自Ivanovce的Allosorex stenodus Fejfar的原始化石记录(晚俄罗斯人,MN 15b)进行了重新描述,并补充了迄今未发表的化石遗骸的描述,包括上牙列(P4和M3)和肱骨碎片以及m2的牙釉质微观结构分析。根据对罗马尼亚、匈牙利和法国遗址的其他化石遗骸的严格评估,Ivanovce化石是迄今为止在欧洲发现的唯一毫无疑问的a . stenodus化石,它可以被认为是当地上新世早期(MN 15)的特有物种。它的发生与“pre-Váh”河宽阔河谷沿线的森林喀斯特地区有关,生态和气候与东南亚现代喀斯特地区的环境相似。根据牙釉质微观结构分析和独特的下颌骨和牙列形态特征,讨论了该物种的单独分类地位。
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引用次数: 0
A new Oligocene flora from Ludvíkovice near Děčín (České středohoří Mts., the Czech Republic) 来自Děčín附近Ludvíkovice的渐新世新植物区系(České středohoří Mts.,捷克共和国)
Q3 Earth and Planetary Sciences Pub Date : 2020-01-01 DOI: 10.37520/fi.2020.024
V. Teodoridis, Z. Kvaček, Miroslav Radoň, Vladislav Raprich, A. Bruch
A recently recovered site of plant macrofossils, Ludvíkovice, in the České středohoří Mts. is situated on Sokolí vrch hill, belonging to the Děčín Formation (radiometrically dated to 30.8–24.7 Ma), according to regional stratigraphy. The flora has yielded a fern, Rumohra recentior, and several angiosperms, but no conifers. The prevailing foliage is preserved without cuticles. Noteworthy are records of Daphnogene cinnamomifolia, Laurophyllum cf. acutimontanum, Platanus neptuni, Sloanea artocarpites, Carya fragiliformis / C. quadrangula, Alnus rhenana, Trigonobalanopsis rhamnoides, Eotrigonobalanus furcinervis and cf. Quercus sp. Several foliage specimens of dicots could not be identified to species level, i.e., Leguminophyllum sp., Pungiphyllum cf. cruciatum and Dicotylophyllum sp. div. The fossil plant assemblage of Ludvíkovice corresponds to zonal mesophytic vegetation accompanied by riparian elements. This is corroborated by the Integrated Plant Record vegetation analysis, which reconstructs a zonal broad-leaved evergreen forest similar to the living broad-leaved evergreen sclerophyllous forest from Southern Hunan and Northern Guangxi in SE China. The vegetation thrived under a humid climate characterized by average values of MAT (14.6–24.1 °C), WMMT (24.7–28.3 °C), CMMT (2.2–18.8 °C) and MAP (979–1724 mm). The fossil flora of Ludvíkovice is similar in composition to the floras of Markvartice, Veselíčko, also from the Oligocene Děčín Formation of the České středohoří Mts. and the Hrazený hill.
根据区域地层学,最近发现的一个大型植物化石遗址Ludvíkovice位于České středohoří mt .的Sokolí vrch山上,属于Děčín组(辐射测定年代为30.8-24.7 Ma)。这里的植物群已经生长出了一种蕨类植物和几种被子植物,但没有针叶树。盛行的叶子被保留下来,没有角质层。值得注意的是,有记录的植物有:cinnamomifolia、Laurophyllum cf. acutimontanum、Platanus neptuni、Sloanea artocarpites、Carya fragiliformis / C. quadrangula、Alnus rhenana、Trigonobalanopsis rhamnoides、Eotrigonobalanus furcinervis和cf. Quercus sp.。Ludvíkovice的化石植物组合对应于带状的叶生植被,并伴有河岸元素。综合植物记录(Integrated Plant Record)植被分析证实了这一点,重建了一个类似于湘南和桂北常绿硬叶阔叶林的地带性常绿阔叶林。植被生长在湿润气候下,平均MAT(14.6 ~ 24.1°C)、WMMT(24.7 ~ 28.3°C)、CMMT(2.2 ~ 18.8°C)和MAP (979 ~ 1724 mm)。Ludvíkovice化石区系的组成与同样来自České středohoří mt .和Hrazený hill渐新世Děčín组的Markvartice, Veselíčko的区系相似。
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引用次数: 1
New genus of amphicyonid carnivoran (Mammalia, Carnivora, Amphicyonidae) from the phosphorites of Quercy (France) 法国Quercy磷化岩中两栖类食肉动物新属(哺乳纲,食肉目,两栖类科)
Q3 Earth and Planetary Sciences Pub Date : 2020-01-01 DOI: 10.37520/fi.2020.013
L. Bonis
An isolated mandible of Carnivora (Mammalia) from the phosphorites of Quercy (France) is described as a new genus. It is compared with the amphicyonid genus Cynodictis, some primitive North American amphicyonids, and with European and North American Eocene carnivoraforms. I conclude that it is a primitive amphicyonid which may be dated to the middle or late Eocene.
从法国Quercy的磷化石中分离出的一种食肉目(哺乳纲)的下颌骨被描述为一个新属。将其与两栖动物属Cynodictis、北美一些原始两栖动物以及欧洲和北美始新世食肉动物进行了比较。我认为这是一种原始的两栖类动物,可以追溯到始新世中晚期。
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引用次数: 0
Unexpected inhibitory cascade in the molariforms of sloths (Folivora, Xenarthra): a case study in xenarthrans honouring Gerhard Storch’s open-mindedness 树懒molariforms (Folivora, Xenarthra)中意想不到的抑制级联:xenarthrans的案例研究,以表彰Gerhard Storch的开放思想
Q3 Earth and Planetary Sciences Pub Date : 2020-01-01 DOI: 10.37520/fi.2020.002
Luciano Varela, P. Tambusso, R. Fariña
The inhibitory cascade (IC) represents a developmental model that explains the evolution of molar relative sizes, originally described in rodents but later validated in several mammalian groups. The IC comprises signalling molecules produced by the first molar buds that inhibit the development of subsequent molars and molecules from surrounding tissues that have opposite effects. Sloths, as xenarthrans, present many peculiarities in their dentition, like tooth and enamel loss, homodonty, and changes in the typically mammalian dental formula. Here, we test the existence of an IC and explore the evolution of the lower dentition in sloths. We studied the variability of molariform proportions in 20 specimens of the Late Pleistocene ground sloth Lestodon armatus. We also analysed molariforms proportions in 53 sloth genera to explore evolutionary trends. Our results show that the lower dentition of most sloths complies with the IC model, despite the difficulties of assessing dental homologies with other mammals. Furthermore, we tested the existence of different patterns among families, obtaining support for models taking mylodontids and orophodontids separately from the rest of sloths. Also, members of Mylodontidae show a unique IC pattern, with a slope considerably higher than 2 and an mf1 ≤ mf2 << mf3 configuration. This pattern could be related to the morphological adaptations to grazing showed by mylodontids during most of their evolutionary history.
抑制级联(IC)代表了一种解释臼齿相对大小进化的发育模型,最初在啮齿动物中描述,但后来在几种哺乳动物群体中得到验证。IC包括由第一磨牙芽产生的抑制后续磨牙发育的信号分子和来自周围组织的具有相反作用的分子。树懒,作为异种动物,在它们的牙齿上表现出许多特点,比如牙齿和牙釉质的脱落,同质性,以及典型的哺乳动物牙齿配方的变化。在这里,我们测试了树懒中IC的存在,并探索了树懒下齿的进化。本文对20个晚更新世地懒(Lestodon armatus)标本的臼齿比例进行了研究。我们还分析了53个树懒属的齿形比例,以探索进化趋势。我们的研究结果表明,大多数树懒的下齿符合IC模型,尽管很难评估与其他哺乳动物的牙齿同源性。此外,我们还测试了不同科间存在的不同模式,得到了将齿状突和齿状突从其他树懒中分离出来的模型的支持。此外,Mylodontidae成员具有明显高于2且mf1≤mf2 << mf3的独特IC模式。这种模式可能与齿状纲在其大部分进化史中表现出的对放牧的形态适应有关。
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引用次数: 4
First record of the stem amniote Discosauriscus (Seymouriamorpha, Discosauriscidae) from the Krkonoše Piedmont Basin (the Czech Republic) Krkonoše皮埃蒙特盆地(捷克共和国)干羊膜动物dissauriscus (Seymouriamorpha, dissauriscdae)的首次记录
Q3 Earth and Planetary Sciences Pub Date : 2020-01-01 DOI: 10.37520/fi.2020.020
Jozef Klembara, Marika Mikudíková, S. Štamberg, M. Hain
The first record of the seymouriamorph stem amniote Discosauriscus from the Krkonoše Piedmont Basin (the Czech Republic) is described. The specimen is identified as D. pulcherrimus on the basis of the following features which are absent in D. austriacus: 1) the pointed tip of the ventrolateral process of the postorbital lies anteriorly to the tip of the wedge-shaped dorsomedial process of the jugal; and 2) the rows of small denticles diverge anteromedially and anterolaterally from the midwidth of the ventral surface of the palatal ramus of the pterygoid. This new record increases our knowledge of the occurrence of this seymouriamorph in the Permo-Carboniferous basins of Europe.
本文描述了在捷克共和国Krkonoše皮埃蒙特盆地发现的首个seymriamorph干羊膜动物dissauriscus。根据以下特征,该标本被确定为D. pulcherrimus,而这些特征在D. austracus中是不存在的:1)后颊腹外侧突的尖尖位于下颌楔形后内侧突尖的前面;(2)小齿行从翼状腭支腹面中宽处向前内侧和前外侧发散。这一新记录增加了我们对这种西摩变形在欧洲二叠-石炭系盆地中发生的认识。
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引用次数: 2
Bibliography of Prof. RNDr. Zlatko Kvaček, DrSc. (2017 – 2020) RNDr教授参考书目。Sweetie Kvaček,博士。(2017-2020)
Q3 Earth and Planetary Sciences Pub Date : 2020-01-01 DOI: 10.37520/fi.2020.017
J. Sakala
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引用次数: 0
Small early Miocene listriodont suid (Artiodactyla: Mammalia) from Sabuncubeli (Manisa, SW Anatolia), Turkey 早中新世小齿兽(偶蹄目:哺乳纲),产自土耳其Sabuncubeli (Manisa, SW Anatolia)
Q3 Earth and Planetary Sciences Pub Date : 2020-01-01 DOI: 10.37520/fi.2020.026
M. Pickford, T. Kaya, Erhan Tarhan, Derya Erylmaz, Serdar Mayda
Turkey is known for the wealth of fossil suids found in deposits of middle Miocene, late Miocene and Plio-Pleistocene levels but material of this family from early Miocene and Palaeogene deposits is rare in the country, one of the few published occurrences being from Şemsettin (Kumartaş Formation, MN 4, Çankiri-Çorum Basin). For this reason, it is interesting to record the presence of small suid remains in the Soma Formation at Sabuncubeli (Manisa, SW Anatolia) in deposits correlated to MN 3 (early Miocene) and thus the earliest known Turkish members of the family. The upper and lower teeth are herein attributed to a new genus and species (Prolistriodon smyrnensis) of Listriodontinae because, in a nascent way, they show a suite of derived morphological features such as upper central incisors with apical sulci, and upper molars with lingual precrista, found in listriodonts but not in Kubanochoerinae, Palaeochoerinae, Tetracondontinae, Hyotheriinae, Namachoerinae, Cainochoerinae or Suinae.
土耳其以在中中新世、晚中新世和上新世-更新世沉积层中发现丰富的化石流体而闻名,但在早中新世和古近纪沉积层中发现的这一科的材料在该国是罕见的,为数不多的已发表的发现之一是在Şemsettin (Çankiri-Çorum盆地MN 4 kumartau组)。因此,在Sabuncubeli (Manisa, SW Anatolia)的Soma组(与mn3(早中新世)相关的沉积物中,记录小型suid遗迹的存在是很有趣的,因此是已知最早的土耳其家族成员。本文将上、下齿归为Listriodontinae的一个新属和新种(Prolistriodon smyrnensis),因为它们在新生时表现出一系列衍生形态特征,如上颌中央门齿有尖沟,上磨牙有舌前嵴,这些特征在listriodondonts中发现,而在Kubanochoerinae、Palaeochoerinae、Tetracondontinae、Hyotheriinae、Namachoerinae、Cainochoerinae或Suinae中没有。
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引用次数: 0
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