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The ultrastructure of the tooth enamel of small Equus of the “tarpan” group and their possible phylogenetic connections “tarpan”类群小马牙釉质的超微结构及其可能的系统发育联系
Q3 Earth and Planetary Sciences Pub Date : 2021-01-01 DOI: 10.37520/fi.2021.007
Vitalii Demeshkant, L. Rekovets
The paper presents the results of a comparative analysis of the ultrastructure of tooth enamel (incisors, premolars and molars) in representatives of morphologically (and possibly phylogenetically) similar forms of small horses. These are both extinct forms of tarpan – Equus gmelini from various sites of the early Holocene of Ukraine (Myrne, Kamiana Mohyla and Hirzhevo) and modern forms – konik polski and E. przewalskii, and European wild ass – E. hydruntinus. Morphology of prisms in different layers of tooth enamel and its relative relationships in various taxa were thoroughly described. The data obtained in the form of tables and graphs, with the appropriate statistical processing thereof, show the morphological similarity of tarpan remains from different sites, and their similarity to the konik polski and Przewalski’s horse. However, the European wild ass differs significantly from the other groups studied. In equines, the radial enamel varieties (types І, ІІ and ІІІ) have evolutionary tendencies to strengthen the first and second types of hydroxyapatite prisms, while reducing the share of the IPM matrix in the enamel structure. In turn, the enamel of the incisors is characterized by lamellar enamel of the HSB uniserial type.
本文介绍了在形态上(和可能的系统发育上)相似的小型马的代表的牙釉质(门牙、前磨牙和臼齿)的超微结构的比较分析结果。这两种都是已经灭绝的tarpan——来自乌克兰全新世早期不同地点的Equus gmelini (Myrne, Kamiana Mohyla和Hirzhevo)和现代形式——konik polski和E. przewalskii,以及欧洲野驴E. hydruntinus。详细介绍了牙釉质不同层的棱柱形态及其在不同分类群中的关系。以表格和图表的形式获得的数据,经过适当的统计处理,显示了不同地点的焦油布遗迹的形态相似性,以及它们与波兰科尼人和普氏野马的相似性。然而,欧洲野驴与其他研究群体有很大不同。马的放射状牙釉质品种(І、ІІ和ІІІ)具有强化第一类和第二类羟基磷灰石棱柱的进化趋势,同时减少了IPM基质在牙釉质结构中的份额。反过来,门牙的牙釉质以HSB单列型的板层牙釉质为特征。
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引用次数: 1
Why Lycospora dominated many Pennsylvanian spore assemblages 为什么Lycospora在许多宾夕法尼亚孢子组合中占优势
Q3 Earth and Planetary Sciences Pub Date : 2021-01-01 DOI: 10.37520/fi.2021.002
B. Thomas
The microspore genus Lycospora in its wider sense is produced by both heterosporous cones, Flemingites, and homosporous cones Lepidostrobus. Calculations are made of the number of microspores that are produced by cones and then by individual Pennsylvannian arborescent lycophytes. The distribution of the rooting rhizohores, Stigmaria can be extrapolated to a square kilometre of forest. A calculation of the number of spores produced in such a forest is enormous which explains why some dispersed spore floras are dominated by these spores. There are dispersed spore assemblages prepared from coal with high percentages of Lycospora. However, the methods of preparation and counting of the spores do not give enough information to make a meaningful calculation of their total numbers in the samples.
广义上的小孢子属Lycospora是由异孢子球果(Flemingites)和同孢子球果(Lepidostrobus)产生的。计算由球果产生的小孢子的数量,然后由单个宾夕法尼亚树状石松产生的小孢子的数量。根根根的分布,可以推断为一平方公里的森林。在这样的森林中产生的孢子数量的计算是巨大的,这就解释了为什么一些分散的孢子区系是由这些孢子主导的。煤中有分散的孢子组合,煤中含有高比例的石孢菌。然而,孢子的制备和计数方法并没有提供足够的信息来对样品中的孢子总数进行有意义的计算。
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引用次数: 0
Early complex tiering pattern: Upper Ordovician, Barrandian area, the Czech Republic 早期复杂层序模式:捷克共和国Barrandian地区上奥陶统
Q3 Earth and Planetary Sciences Pub Date : 2021-01-01 DOI: 10.37520/fi.2021.003
Pavel Bokr, R. Mikuláš, P. Budil, P. Kraft
Upper Ordovician shallow marine fine-grained sandstones and siltstones exposed in the Loděnice – vinice locality yielded a distinct and well-preserved tiering pattern of trace fossils. The two uppermost tiers are composed mainly of Bifungites and Nereites. Deeper in the sediment, tiers dominated by Thalassinoides, Zoophycos and Teichichnus occur. Most of the succession is completely bioturbated; however, several storm layers enabled study of a well-preserved frozen tiering pattern. Large portions of the bedding planes (ichnologic snapshots) showed a considerable patchiness of intensive surface bioturbation and a preferred orientation of Bifungites. The identified tiering pattern is one of the earliest examples of a well-documented complex tiering of burrows documented in detail.
在洛德涅斯-维尼斯地区暴露的上奥陶统浅海相细粒砂岩和粉砂岩中,发现了保存完好的明显的化石层序。最上面的两层主要由双辉石和内辉石组成。在沉积物的深处,以海藻纲、植藻纲和Teichichnus纲为主的分层出现。大部分演替完全受到生物扰动;然而,几个风暴层使研究保存完好的冰冻分层模式成为可能。大部分层理平面(技术快照)显示出明显的表面生物扰动斑块和双辉石的优先取向。所确定的分层模式是详细记录的有良好记录的复杂洞穴分层的最早例子之一。
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引用次数: 0
First record of intact equisetalean strobili from the Wealden (Lower Cretaceous) of the Isle of Wight, southern England 来自英格兰南部怀特岛威尔登(下白垩纪)的第一个完整的equisetalean strobili记录
Q3 Earth and Planetary Sciences Pub Date : 2021-01-01 DOI: 10.37520/fi.2021.005
C. Pott
Two excellently preserved small strobili were obtained from a Wealden plant debris bed in the Lower Cretaceous (Barremian) Wessex Formation, south-east of Chilton Chine, on the Isle of Wight, southern England. The strobili are preserved as compressions and show the characteristic morphology of sporangiophore heads of Equisetales. Based on the morphology of the strobili, attribution to a certain species is not warranted. Therefore, the strobili have been left unassigned in the fossil-genus Equisetostachys which is commonly used for isolated strobili of fossil sphenophytes. From their size, shape and constitution, the strobili are interpreted as immature; the absence of preserved sporangia and spores is consequently not unexpected. Affiliation with Equisetum burchardtii might be an option. The strobili represent the first record of any equisetalean or sphenophyte remains from the Wessex Sub-basin of the English Wealden and are thus of considerable importance. The find is especially significant because previously known specimens from the Weald Sub-basin and the German Wealden are confined to subterranean rhizomes, adventitious roots, tubers and bases of aerial shoots, commonly preserved in situ, together with only fragmentary remains of sporangiophore heads from disarticulated strobili. These strobili finds are thus the first intact equisetalean reproductive structures from the Wealden of either England or Germany.
在英国南部怀特岛奇尔顿中国东南部的下白垩纪(巴雷米亚)韦塞克斯组的威尔登植物碎屑床上,发现了两个保存完好的小球。球果以压缩形式保存,表现出木贼属孢子囊头状花序的特征形态。根据球果的形态,不能将其归为某一物种。因此,在棘叶植物化石属(Equisetostachys)中,棘叶属(Equisetostachys)未被归属,而棘叶植物化石属通常用于分离棘叶。从它们的大小、形状和构造来看,它们被认为是不成熟的;因此,没有保存的孢子囊和孢子并不意外。与木贼(Equisetum burchardtii)的关系可能是一个选择。这些石柱代表了英国威尔登的威塞克斯次盆地中首次记录的马属植物或叶生植物遗骸,因此具有相当重要的意义。这一发现尤其重要,因为以前已知的来自威尔德亚盆地和德国威尔登的标本仅限于地下根茎、不定根、块茎和气生芽的基部,这些通常保存在原位,以及来自分离的球果的孢子囊头的碎片残骸。因此,这些strobili的发现是来自英格兰或德国威尔登的第一个完整的马类生殖结构。
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引用次数: 1
New leaf species from the upper Miocene flora of the leaf-bearing Wischgrund clay (Lower Lusatia, Brandenburg, Germany) 德国勃兰登堡下卢萨蒂亚地区Wischgrund粘土中新世上部植物区系的新叶种
Q3 Earth and Planetary Sciences Pub Date : 2021-01-01 DOI: 10.37520/fi.2021.009
U. Striegler
This is a brief outline of the fossil content of the Wischgrund leaf-bearing clay (Lower Lusatia, Brandenburg State, Germany), which belongs to the youngest Neogene layers of Lower Lusatia (upper Miocene, Tortonian). It is a deciduous temperate flora, with a small proportion of evergreen species. The reconstructed vegetation can be assigned to an azonal swamp forest with Taxodium and a riparian forest with Liquidambar and Acer as the most frequent species. Furthermore, a zonal Mixed Mesophytic Forest characterized by Quercus and Fagus is proposed in upland positions. The flora consists of 78 species of leaves, 51 carpological species and 82 palynological groups. The climate belongs to Cfa-type according to Köppen. Two new species of leaves are described from there, Pyracantha pseudococcinea sp. nov. and Ternstroemites klettwitzensis sp. nov.
这是Wischgrund含叶粘土(德国勃兰登堡州下卢萨蒂亚)化石含量的简要概述,它属于下卢萨蒂亚(上中新世,托尔顿期)最年轻的新近纪层。它是一个温带落叶植物区系,有一小部分常绿物种。重建的植被可划分为以紫杉属为主的地带性沼泽林和以枫木和槭木为主的河岸林。此外,在丘陵地带还可形成以栎、Fagus为主要树种的地带性混合中植物林。植物区系包括78种叶片、51种树状植物和82个孢粉类群。根据Köppen,气候属于cfa型。其中描述了两个新种:Pyracantha pseudococcinea sp. 11和Ternstroemites klettwitzensis sp. 11。
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引用次数: 0
New suoid fossils (Mammalia, Artiodactyla) from the Miocene of Moghara, Egypt, and Gebel Zelten, Libya: biochronological implications 埃及Moghara和利比亚Gebel Zelten中新世新类人猿化石(哺乳类,偶蹄类):生物年代意义
Q3 Earth and Planetary Sciences Pub Date : 2021-01-01 DOI: 10.37520/fi.2021.010
M. Pickford, M. A. Abdel Gawad, M. Hamdan, A. El-Barkooky, Mohammed H. Al Riaydh
Some undescribed suoid specimens from early and middle Miocene deposits at Moghara, Egypt, and Gebel Zelten, Libya, are of interest for biochronology. The fossils comprise maxillae and mandibles with incomplete dentitions, which are described and illustrated in detail. Three species of suids and one sanithere occur at Moghara. A huge edentulous suid mandible was collected at Gebel Zelten in 1997 during the Spanish-Libyan Palaeontology Expedition. In January, 2020, additional sanithere fossils were collected from Moghara by a team from Cairo University and the Muséum National d’Histoire Naturelle, Paris. A suid mandible with four teeth collected from Moghara in 1994, and now curated at the Cairo Geological Museum, confirms the presence of the species Libycochoerus massai at the site, previously represented by an isolated upper molar and two canines. A talus previously thought to be from Moghara is now known to have been collected at Wadi Natrun (late Miocene) and thus probably represents a tetraconodont rather than a kubanochoere. The age of the Moghara deposits is estimated to span the period ca. 19.5–16.5 Ma (late early Miocene, Faunal Sets PII–PIIIa) and the Zelten sequence is most likely to span the period ca. 17–14.5 Ma (late early Miocene to basal middle Miocene, Faunal Sets PIIIa–PIIIb).
在埃及Moghara和利比亚Gebel Zelten的早中新世和中中新世沉积物中发现的一些未描述的水怪标本对生物年代学具有重要意义。这些化石包括带有不完整牙列的上颌骨和下颌骨,并对其进行了详细的描述和说明。在Moghara有三种suids和一种sani。1997年,西班牙-利比亚古生物学考察队在格贝尔泽尔滕发现了一块巨大的无牙鳄鱼下颌骨。2020年1月,开罗大学和巴黎国立自然历史博物馆的一个研究小组从Moghara收集了更多的蜥蜴化石。1994年在Moghara收集的一副带有四颗牙齿的suid下颌骨,现在收藏在开罗地质博物馆,证实了该遗址中存在利比亚人(Libycochoerus massai)这个物种,之前只有一颗孤立的上臼齿和两只犬齿。先前被认为是来自Moghara的一块距骨,现在已知是在Wadi Natrun(中新世晚期)收集的,因此可能代表四齿齿兽而不是kubanochoere。Moghara沉积的年龄估计为19.5-16.5 Ma(早中新世晚期,PII-PIIIa), Zelten层序的年龄估计为17-14.5 Ma(早中新世晚期至中新世中期基底,PIIIa-PIIIb)。
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引用次数: 0
The Amphicyoninae (Amphicyonidae, Carnivora, Mammalia) of the early Miocene from Tuchořice, the Czech Republic 捷克共和国Tuchořice中新世早期两栖动物科(两栖动物科,食肉目,哺乳纲)
Q3 Earth and Planetary Sciences Pub Date : 2021-01-01 DOI: 10.37520/fi.2021.011
J. Morales, O. Fejfar, E. Heizmann, J. Wagner, A. Valenciano, Juan Abella
The Amphicyoninae of the early Miocene from the locality of Tuchořice, the Czech Republic, are represented by three species. Two of them are classified within the tribe Amphicyonini: Paludocyon bohemicus (Schlosser, 1899) as the type species of Paludocyon n. gen., and a large-sized amphicyonid determined as Megamphicyon carnutense (Antunes et Ginsburg, 1977). Dehmicyon n. gen. aff. schlosseri is determined by two small teeth. This new genus has been proposed for the species Amphicyon schlosseri Dehm, 1950 from Wintershof-West and is tentatively included in the tribe Pseudarctini nov. together with the genera Ictiocyon and Pseudarctos. This association of Amphicyoninae provides valuable information on the taxonomy and systematics of this subfamily during the early Miocene, at which time important environmental changes were taking place in Europe, which undoubtedly affected the evolution of Amphicyonidae.
捷克共和国Tuchořice地区中新世早期的两栖动物科有3个种。其中两种属于两栖类:Paludocyon bohemicus (Schlosser, 1899)作为Paludocyon n. gen.的模式种,另一种大型两栖类被确定为Megamphicyon carnutense (Antunes et Ginsburg, 1977)。Dehmicyon n. gen. affer . schlosseri是由两颗小牙齿决定的。这个新属被认为是来自Wintershof-West的Amphicyon schlosseri Dehm, 1950,并暂定与Ictiocyon属和pseudoarctos属一起归入pseudoarctini 11 .族。这一组合为该亚科在中新世早期的分类学和系统学研究提供了有价值的信息,这一时期欧洲发生了重要的环境变化,这无疑影响了两栖科的进化。
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引用次数: 3
New actinopterygians from the Permian of the Brive Basin, and the ichthyofaunas of the French Massif Central 布里夫盆地二叠纪的新放光翼目动物和法国中部地块的鱼系动物
Q3 Earth and Planetary Sciences Pub Date : 2021-01-01 DOI: 10.37520/fi.2021.012
S. Štamberg, J. Steyer
A new actinopterygian fauna from the Permian of the Brive Basin (Corrèze, Massif Central, France) is described in detail. It is represented by a new pygopterid taxon, Briveichthys chantepieorum gen. et sp. nov., erected and described here on the basis of several well-preserved specimens showing peculiar characters in the bones of the skull roof, parasphenoid, bones of the cheek and opercular apparatus. Other specimens from this new fauna are also described; they are referred to Progyrolepis heyleri Poplin, 1999, also known in the Bourbon-l’Archambault Basin, and Aeduella cf. blainvillei, which is relatively common in the Permian of France. These discoveries show the importance of the Brive Basin which was previously considered as a rather barren basin in term of palaeontology. This new fauna from Brive is compared with the other Permian ichthyofaunas from France: it is less rich in terms of specimen numbers than the faunas of Bourbon-l’Archambault and Autun, but it is more diversified in terms of number of taxa than the faunas of Lodève and L’Argentière (Ardêche). These actinopterygians, together with other aquatic vertebrates (e.g., acanthodians, sharks etc.), were widespread and diversified in the Hercynian Mountain Chain during the Permian. They indicate that the French basins were connected in time and space. The wide distribution of these aquatic taxa may have been favoured by numerous hydric systems (lakes, rivers etc.) which were well developed under the tropical climate.
详细描述了法国中部地块布里夫盆地(corr)二叠系一种新的放光翼类动物群。它的代表是一个新的拟翅目分类群,Briveichthys chantepieorum gen. et sp. nov.,它是在几个保存完好的标本的基础上建立和描述的,这些标本在头盖骨、副棘骨、颊骨和眼器官上表现出独特的特征。本文还描述了这一新动物群的其他标本;它们被称为Progyrolepis heyleri Poplin, 1999年,在Bourbon-l 'Archambault盆地也被称为,以及Aeduella cf. blainvillei,这在法国的二叠纪盆地相对常见。这些发现显示了布里夫盆地的重要性,该盆地以前在古生物学方面被认为是一个相当贫瘠的盆地。这个来自Brive的新动物群与来自法国的其他二叠纪鱼类动物群进行了比较:就标本数量而言,它不如Bourbon-l 'Archambault和autumn的动物群丰富,但就分类群数量而言,它比lod和L ' argenti的动物群更多样化(Ardêche)。在二叠纪期间,这些放光翼动物与其他水生脊椎动物(如棘足动物、鲨鱼等)一起在海西山脉广泛分布和多样化。它们表明法国盆地在时间和空间上是相通的。这些水生分类群的广泛分布可能得益于热带气候下发达的众多水系(湖泊、河流等)。
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引用次数: 2
The occurrence of pollen of Sciadopityaceae Luerss. through time 山菖蒲科植物花粉的发生。经过漫长的时间
Q3 Earth and Planetary Sciences Pub Date : 2021-01-01 DOI: 10.37520/fi.2021.019
C. Hofmann, N. Odgerel, L. Seyfullah
Here we present LM and SEM data of pollen of extant Sciadopitys verticillata, fossil Cerebropollenites from Aptian/Albian strata (Austria, Mongolia) and Sciadopityspollenites from Campanian/Maastrichtian and Oligocene/Miocene strata (Siberia, Germany). Measurements and image comparisons show that the investigated fossil pollen taxa range from somewhat comparable to very similar to extant Sciadopitys verticillata, and that a previous affiliation of Cerebropollenites taxa with Tsuga cannot be corroborated. Additionally, it can be speculated that either the Rhaetian to Lower Cretaceous Cerebropollenites taxa are the pollen equivalent of the Eurasian Miroviaceae macrofossils that have unresolved relationships with Sciadopityaceae, or that they might belong to a basal group in the Sciadopityaceae, which is quite recognizable due to the similarity of the pollen morphology of Cerebropollenites thiergartii and Sciadopitys verticillata. However, until in situ pollen within cones attributed to the Miroviaceae are found, we will not know for sure.
本文介绍了奥地利、蒙古Aptian/Albian地层(奥地利、蒙古)和德国西伯利亚Campanian/Maastrichtian和渐新世/中新世地层(Campanian/Maastrichtian和渐新世/中新世)现存sciadopityverticillata的花粉、化石脑花粉的LM和SEM数据。测量和图像比较表明,所调查的化石花粉分类群与现存的花粉类群有些相似或非常相似,并且先前的脑花粉分类群与Tsuga的隶属关系无法得到证实。此外,可以推测,Rhaetian至下白垩世的脑花粉分类群可能是与Sciadopityaceae关系尚未确定的欧亚Miroviaceae大化石的花粉等量,或者它们可能属于Sciadopityaceae的一个基群,由于thiergartii和Sciadopitys verticillata的花粉形态相似,这一点很容易识别。然而,直到在球果内的原花粉被发现,我们将无法确定。
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引用次数: 1
Stratigraphy, chronology and palaeontology of the Tertiary rocks of the Cheringoma Plateau, Mozambique 莫桑比克切林戈马高原第三纪岩石的地层学、年代学和古生物学
Q3 Earth and Planetary Sciences Pub Date : 2021-01-01 DOI: 10.37520/fi.2021.014
M. Bamford, M. Pickford
The discovery of fossil plants, marine molluscs and mammals in the Mazamba Formation, Cheringoma Plateau, Mozambique, opens a new chapter in the study of this part of the African Rift System. The evidence suggests that the Mazamba Formation is older than previously reported, probably late Eocene rather than Miocene. The fossil wood and stems indicate a frost-free tropical humid environment and a high water table soon after deposition, and the marine molluscs and mammals indicate proximity to the sea. There is also evidence for the occurrence of pans in the area during the late Eocene which also suggest a near-surface water table. This paper discusses the history of interpretation of the geology of the Cheringoma Plateau and describes and interprets the fossil plants, molluscs and mammals collected in 2012 and 2013. It is concluded that the Mazamba Formation, which overlies the fully marine Lutetian-Bartonian Cheringoma Limestone, is a coastal facies (fluvio-deltaic, lagoonal and onshore deposits) that accumulated on top of the marine limestones as sea level dropped late in the Bartonian. Mammalian bones from the White Patch sites represent a heavily built species about the dimensions of a pygmy hippopotamus, probably belonging to the order Embrithopoda. If so, then the Mazamba Formation is likely to correlate to the latest Bartonian or early Priabonian rather than to the Miocene as previously assumed.
在莫桑比克切林戈马高原的马赞巴组发现了植物、海洋软体动物和哺乳动物化石,为研究非洲裂谷系的这一部分打开了新的篇章。证据表明,马赞巴组比先前报道的更古老,可能是始新世晚期,而不是中新世。化石木材和茎表明一个无霜的热带潮湿环境和沉积后不久的高地下水位,海洋软体动物和哺乳动物表明靠近大海。还有证据表明,该地区在始新世晚期出现了平底锅,这也表明地下水位接近地表。本文讨论了切林戈马高原地质解释的历史,并对2012年和2013年收集的植物、软体动物和哺乳动物化石进行了描述和解释。结果表明,马赞巴组位于完全海相的卢腾—巴顿期切林戈马灰岩之上,是在巴顿期晚期海平面下降时沉积在海相灰岩之上的海岸相(河流三角洲、泻湖和陆上沉积)。从白色斑块遗址发现的哺乳动物骨骼表明,这是一种体型庞大的物种,大约有侏儒河马那么大,可能属于肢足目。如果是这样的话,那么马赞巴组很可能与最新的巴顿世或早期的普里亚伯世有关,而不是像之前假设的那样与中新世有关。
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引用次数: 0
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