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Determining the gait of Miocene, Pliocene, and Pleistocene horses from fossilized trackways 根据足迹化石确定中新世、上新世和更新世马的步态
IF 1.4 4区 地球科学 Q3 Earth and Planetary Sciences Pub Date : 2021-06-10 DOI: 10.5194/FR-24-151-2021
A. Vincelette
Abstract. Much work has been done on the study of vertebrate gaits over the pastseveral decades and efforts undertaken to apply this to fossil tracks,especially dinosaurs and mammals such as cats, dogs, camels, and horses.This work seeks to expand upon such studies and in particular to studyfootprints laid down in sand by modern horses and apply such studies todetermine the gaits of fossil horse trackways. It thus builds upon the workof Renders (1984a, b) and Kienapfel et al. (2014) and suggests additionalmeasurements that can be taken on horse footprints. In this study thefootprints left in the sand by 15 horses of various breeds with variousgaits were videotaped, photographed, described, and measured in order todetermine characteristics useful in distinguishing gaits. These results werethen applied to two new sets of fossil footprints, those of the middleMiocene merychippine horse Scaphohippus intermontanus that I personally examined and measured andthose from the late Pleistocene horse Equus conversidens, previously illustrated and describedin the literature (McNeil et al., 2007). The latter horse exhibits a fastgallop of around 9.4 m/s, but it is the former whose footprints are quiteunique. The quantitative and visual features of these prints are suggestiveof a medium-fast gait involving apparent “understepping” of diagonalcouplets and hind feet that overlap the centerline. The gait that mostclosely matches the footprints of Scaphohippus is the “artificial” gait of a slowrack or tolt, or pace, around 1.9 m/s, though an atypical trot of a horsewith major conformation issues or which is weaving (swaying) from side toside is a less likely possibility. This intimates, along with the earlierstudy of Renders (1984a, b), who found the artificial gait of the running walkdisplayed by Pliocene hipparionine horses, that ancient horses possessed amuch greater variety of gaits than modern horses and that over time theylost these abilities with the exception of certain gaited breeds.
摘要在过去的几十年里,人们对脊椎动物的步态进行了大量的研究,并努力将其应用于化石足迹,特别是恐龙和哺乳动物,如猫、狗、骆驼和马。这项工作旨在扩展这些研究,特别是研究现代马在沙子上留下的脚印,并将这些研究应用于确定化石马足迹的步态。因此,它建立在渲染(1984a, b)和Kienapfel等人(2014)的工作基础上,并建议可以对马脚印进行额外的测量。在这项研究中,我们对15匹不同品种、不同步态的马在沙地上留下的脚印进行了录像、拍照、描述和测量,以确定区分步态的有用特征。然后将这些结果应用于两组新的化石足迹,一组是我亲自检查和测量的中新世中马Scaphohippus intermontanus,另一组是来自晚更新世马Equus conversidens,之前在文献中有说明和描述(McNeil et al., 2007)。后一匹马能以每秒9.4米的速度疾驰,但只有前一匹马的脚印才相当独特。这些脚印的数量和视觉特征表明,这是一种中速步态,包括对角对联的明显“下步”和后脚重叠在中心线上。与舟形马脚印最接近的步态是一种“人造”步态,大约1.9米/秒,尽管一匹有重大构象问题的非典型小跑或从一侧到另一侧编织(摇摆)的可能性较小。这表明,再加上早些时候的研究,发现上新世海马所展示的人造步态,古代马比现代马拥有更多种类的步态,随着时间的推移,除了某些有步态的品种外,它们失去了这些能力。
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引用次数: 3
A new enigmatic lineage of Dascillidae (Coleoptera: Elateriformia) from Eocene Baltic amber described using X-ray microtomography, with notes on Karumiinae morphology and classification 利用X射线显微成像技术描述了始新世波罗的海琥珀中一个新的神秘的Dascillidae谱系(鞘翅目:Elateriformia),并对Karumiinae的形态和分类进行了注释
IF 1.4 4区 地球科学 Q3 Earth and Planetary Sciences Pub Date : 2021-05-21 DOI: 10.5194/FR-24-141-2021
Robin Kundrata, M. Gimmel, Gabriela Packova, A. Bukejs, S. Blank
Abstract. Dascillidae are a species-poor beetle group with a scarcefossil record. Here, we describe Baltodascillus serraticornis gen. et sp. nov. based on a well-preservedspecimen from Eocene Baltic amber. It differs from all known Dascillidae byits reduced mandibles. After studying the specimen using light microscopyand X-ray microtomography, we tentatively place this genus in the poorlydefined subfamily Karumiinae based on the large eyes, serrate antennae, andlack of prosternal process. This is the first representative of theDascillidae formally described from Baltic amber and the first describedfossil member of the subfamily Karumiinae. We briefly discuss theproblematic higher classification of Dascillidae, along with the morphologyand biogeography of the group.
摘要Dascillidae是一个物种贫乏的甲虫群,化石记录很少。在这里,我们描述了Baltodascillus serraticornis gen. et sp. 11 .基于一个保存完好的始新世波罗的海琥珀标本。它与所有已知的滑蝇科的不同之处在于它的下颌骨缩小了。在使用光学显微镜和x射线显微断层扫描对标本进行研究后,我们根据大眼睛,锯齿状触角和缺乏前胸骨突初步将该属归入定义不明确的Karumiinae亚科。这是在波罗的海琥珀中被正式描述的dascillidae的第一个代表,也是Karumiinae亚科的第一个被描述的化石成员。我们简要地讨论了有问题的Dascillidae高级分类,以及该群体的形态和生物地理。
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引用次数: 1
First record of the subfamily Sagrinae (Coleoptera: Chrysomelidae) from the Eocene of North America 北美始新世首次记录的金鱼亚科(鞘翅目:金鱼科)
IF 1.4 4区 地球科学 Q3 Earth and Planetary Sciences Pub Date : 2021-05-12 DOI: 10.5194/FR-24-135-2021
A. Legalov
Abstract. A new genus, Palaeatalasis gen. nov. (type species P. monrosi sp. nov.), from the tribe Megamerini(Chrysomelidae: Sagrinae) from the early-middle Eocene Green River Formationis described and illustrated. The new genus is similar to the RecentAtalasis Lacordaire, 1845 but differs from it in the subparallel sides of thepronotum, metafemora without teeth, and non-emarginate eyes. It differs fromthe Eocene Eosagra Haupt, 1950 in the large, convex, non-emarginate eyes, wideelytra, and transverse pronotum. The new genus is distinguished from the Paleocene Gallopsis Legalov, Kirejtshuk et Nel, 2019 in the wide forehead and convexeyes. It is the first record of the Sagrinae from North America and thefourth known species of the family Chrysomelidae from the Green River.
摘要一个新属,Palaetalasis gen.nov.(模式种P.monrosi sp.nov.),来自始新世中期早期绿河地层的Megamerini部落(金花虫科:Sagrinae)。这一新属与1845年的RecentAtalasis Lacordaire属相似,但与之不同的是,前肢的近平行侧、没有牙齿的后股骨和非微缺眼。它与1950年始新世的Eosagra Haupt的不同之处在于大而凸、非微缺的眼睛、宽的鞘翅和横向的前足。新属与古新世Gallopsis Legalov,Kirejtshuk et Nel,2019在宽额和凸起方面有所不同。这是北美洲首次记录到仙人掌科,也是绿河金花虫科已知的第四个物种。
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引用次数: 2
The polycentropodid genus Cernotina (Insecta, Trichoptera) in Miocene Dominican amber 中新世多米尼加琥珀中的多心足动物Cernotina属(昆虫亚目,毛翅目)
IF 1.4 4区 地球科学 Q3 Earth and Planetary Sciences Pub Date : 2021-05-10 DOI: 10.5194/FR-24-129-2021
W. Wichard, C. Neumann
Abstract. A new polycentropodid caddisfly species is described from MioceneDominican amber. The family Polycentropodidae is therefore represented inthe Dominican amber with two species belonging to the genus Cernotina: C. pulchra Wichard, 2007,and C. fossilinova sp. nov. The endemic C. danieli Flint & Sykora, 2004, is the onlyrepresentative of the genus occurring on Hispaniola today and is similar to thetwo fossil species. On the Caribbean islands altogether six extant speciesof the genus Cernotina are registered.
摘要一个新的多中心足类球童物种描述自中新世多明尼加琥珀。因此,多明尼加琥珀中有多足目动物科的代表,有两个物种属于Cernotina属:C.pulchra Wichard,2007年和C.石化新星属。当地的C.danieli Flint和Sykora,2004年,是该属目前在伊斯帕尼奥拉岛上的唯一代表,与这两个化石物种相似。在加勒比群岛上,共有六种现存的卷须属物种登记在册。
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引用次数: 3
Two new species of the family Rhynchitidae (Coleoptera: Curculionoidea) from Eocene Baltic amber, with key to species and assumed trophic relationships 始新世波罗的海琥珀纹蛛科二新种(鞘翅目:纹蛛总科)及其物种和营养关系
IF 1.4 4区 地球科学 Q3 Earth and Planetary Sciences Pub Date : 2021-05-04 DOI: 10.5194/FR-24-117-2021
A. Bukejs, A. Legalov
Abstract. Two new fossil species of Baltocar Kuschel, 1992 and Pseudomesauletes Legalov, 2001 (Curculionoidea:Rhynchitidae) are described from Eocene Baltic amber. Baltocar sontagae sp. nov. is similar toB. groehni Riedel, 2012 but differs in the shorter rostrum, tarsomere 1 shorterthan tarsomere 5 and 1.4× as long as tarsomere 2, longer elytra,and shorter body. Pseudomesauletes lobanovi sp. nov. is similar to P. culex (Scudder, 1893) and P. ibis(Wickham, 1912): the new species differs from P. culex in the larger eyes, longerpronotum, being 0.44× shorter than elytra, and slightly smaller bodysize; from P. ibis it differs in the smaller body size, rostrum shorter thanelytra, and weakly convex pronotum. This is the first record ofPseudomesauletes from Baltic amber and the sixth species of Baltocar. Keys to species of the genusBaltocar and to the Eocene species of the genus Pseudomesauletes are given. Assumed trophicrelationships of these fossil taxa are discussed (urn:lsid:zoobank.org:pub:260EBE6E-DA6C-4D6D-A1D2-2C258224622F).
摘要Baltocar Kuschel的两个新化石物种,1992年和Pseudomesauletes Legalov的两个化石物种(Curculinoidea:Rhynchitidae)描述自始新世波罗的海琥珀。Baltocar sontagae sp.nov.与B相似。groehni Riedel,2012,但不同之处在于较短的主席台,跗骨1比跗骨5短,是跗骨2的1.4倍长,鞘翅较长,身体较短。Pseudomesauletes lobanovi sp.nov.与P.culex(Scudder,1893)和P.ibis(Wickham,1912)相似:新物种与P.culexs的不同之处在于较大的眼睛,长额足,比鞘翅短0.44倍,体型略小;与朱鹭的不同之处在于体型较小,喙短于喙,前凸无力。这是波罗的海琥珀中首次记录到的假琥珀,也是波罗的海珊瑚的第六个物种。文中给出了巴托卡尔属物种和始新世假美苏莱属物种的分类索引。讨论了这些化石分类群的假定营养关系(urn:lsid:zoobank.org/pub:260EBE6E-DA6C-4D6D-A1D2-2C258224622F)。
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引用次数: 6
Extensive vertebrate tracksite from the Upper Red Formation (middle–late Miocene), west Zanjan, northwestern Iran 伊朗西北部赞詹西部上红组(中新世中晚期)广泛的脊椎动物足迹
IF 1.4 4区 地球科学 Q3 Earth and Planetary Sciences Pub Date : 2021-04-23 DOI: 10.5194/FR-24-101-2021
N. Abbassi, Masoumeh Khoshyar, S. Lucas, F. Esmaeili
Abstract. An extensive vertebrate tracksite from the middle–lateMiocene Upper Red Formation in western Zanjan Province, northwestern Iran,provides new records of paleobiogeographical significance. These are recordsof common footprints of felids referred to as Felipeda lynxi Panin and Avram, canidsreferred to as Canipeda longigriffa Panin and Avram and less common bird footprints referred to asIranipeda abeli Lambrecht. The Canipeda record establishes the late Miocene presence of canids onthe Iranian Plateau as part of the Eurasia-wide “Eucyon event”. TheFelipeda footprints are consistent with body fossil and footprint records elsewherein Eurasia that indicate a widespread distribution of felids by late Miocenetime. An unusual trace associated with the footprints is a large, shallowgrazing or locomotion trace similar to Megaplanolites in some features but distinctive invarious features and its occurrence in nonmarine facies, and it will be thesubject of further study.
摘要伊朗西北部赞詹省西部中新世中晚期上红组的一个广泛的脊椎动物足迹提供了具有古生物地理学意义的新记录。这些是被称为Felipeda lynxi Panin和Avram的猫科动物的常见足迹记录,被称为Canipeda longigriffa Panin和Avram的犬科动物的足迹记录,以及被称为Iranipeda abeli Lambrecht的不太常见的鸟类足迹记录。犬科动物记录表明,作为欧亚大陆范围内“Eucyon事件”的一部分,伊朗高原上存在中新世晚期的犬科动物。Felipeda足迹与欧亚大陆其他地区的身体化石和足迹记录一致,表明到中新世晚期,猫科动物分布广泛。与脚印相关的一个不寻常的痕迹是一个大型浅放牧或运动痕迹,在某些特征上类似于巨平橄榄岩,但其独特的不变特征及其在非海相中的出现,这将是进一步研究的主题。
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引用次数: 1
A new delphinid from the lower Pliocene of the North Sea and the early radiations of true dolphins 北海上新世晚期的一种新飞燕和真正海豚的早期辐射
IF 1.4 4区 地球科学 Q3 Earth and Planetary Sciences Pub Date : 2021-04-07 DOI: 10.5194/FR-24-77-2021
Alice Belluzzo, O. Lambert
Abstract. With a remarkable exception for the Mediterranean, the worldwidePliocene record of true dolphins (Cetacea, Odontoceti, Delphinidae) remainsscarce, in stark contrast with the large number of extant species testifyingto the evolutionary success of this family. Based on a fragmentary skulldiscovered in lower Pliocene deposits (Zanclean, 5 to 4.4 Ma) of theKattendijk Formation in the Antwerp harbour (Belgium, southern margin of theNorth Sea basin), we describe here a new delphinid species, Pliodelphis doelensis gen. et sp.nov. This small dolphin with cranial dimensions in the range of theshort-beaked common dolphin Delphinus delphis can be distinguished from other extinct andextant delphinids by a combination of cranial features including the maxillabeing significantly narrower than the premaxilla at a short distanceanterior to the antorbital notch in dorsal view, the maximum opening of themesorostral groove being located at the level of the antorbital notches, atransversely wide and anteroposteriorly long dorsal exposure of thepresphenoid anterior to the bony nares, and all dorsal infraorbital foraminabeing located posterior to the premaxillary foramina. P. doelensis constitutes the firstmember of the family described from the early Pliocene of the North Seabasin and, for the whole North Atlantic realm, only the third outside theMediterranean. This new record contributes thus to our understanding of thepoorly known Pliocene radiation(s) of true dolphins.
摘要除了地中海的一个显著例外,上新世世界范围内真正的海豚(鲸目、Odontoceti、Delphinidae)的记录仍然很少,这与证明该科进化成功的大量现存物种形成了鲜明对比。基于在上新世晚期矿床中发现的碎屑岩(Zanclean,5至4.4 Ma),我们在这里描述了一种新的飞燕属物种,Pliodelphis doelensis gen.et sp.nov。这种头骨尺寸在短喙普通海豚Delphinus delphis范围内的小海豚可以通过多种头骨特征的组合与其他已灭绝和近端的delphinids区分开来,包括上颌骨明显比前颌骨窄,鼻内侧沟的最大开口位于眶内切口的水平面上,在骨鼻孔前方有一个横向宽且前后长的乳突背侧暴露,所有眶下背孔位于上颌前孔的后方。P.doelensis是北海盆地上新世早期描述的家族中的第一个成员,在整个北大西洋地区,它只是地中海以外的第三个成员。因此,这一新记录有助于我们理解真正海豚已知的上新世辐射。
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引用次数: 0
A new Eocene Bacanius species (Histeridae: Dendrophilinae) from Baltic amber 波罗的海琥珀始新世一新种
IF 1.4 4区 地球科学 Q3 Earth and Planetary Sciences Pub Date : 2021-04-07 DOI: 10.5194/FR-24-93-2021
V. Alekseev, A. Bukejs
Abstract. A study of Baltic amber revealed a new fossil species of the genusBacanius LeConte, 1853. Bacanius gorskii sp. nov. differs from the congener described previously fromEocene Rovno amber, B. kirejtshuki Sokolov and Perkovsky, in the shape of meso-metaventralsuture, incomplete subhumeral stria, and the impunctate prosternal lobe (urn:lsid:zoobank.org:pub:EE9C6859-607A-4134-9037-8385601CF42F).
摘要对波罗的海琥珀的研究发现了一种新的化石物种bacanius LeConte, 1853年。Bacanius gorskii sp. 11 .与之前在meocene Rovno琥珀,B. kirejtshuki Sokolov和Perkovsky中描述的同类不同,在中-中心缝,不完整的肱骨下纹和不完整的前胸骨叶的形状(urn:lsid:zoobank.org:pub: ee9c6859 - 607a: 4134-9037- 8385601cf42f)。
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引用次数: 4
Selenogonus narinoensis Stirton, 1947 (Tayassuidae, Cetartiodactyla, Mammalia): taxonomic status and paleobiogeographic implications 纳氏Selenogonus narinoensis Stirton,1947(Tayasuidae,Cetartiodactyla,Mammalia):分类地位和古生物地理学意义
IF 1.4 4区 地球科学 Q3 Earth and Planetary Sciences Pub Date : 2021-03-29 DOI: 10.5194/FR-24-65-2021
G. Gasparini, Oscar F. Moreno-Mancilla, J. L. Cómbita
The species Selenogonus narinoensis was described by Stirton (1947) based on a single specimen which comes from sediments cropping out in the Cocha Verde locality, Nariño Department (Colombia), tentatively referred to the late Pliocene–Pleistocene (MGN 931; IGM p002118, Museo Geológico Nacional, Servicio Geológico Colombiano, Bogotá). However, morphological studies and comparative morphometric observations of the specimen suggest that (1) no diagnostic character supports the validity of the species Selenogonus narinoensis (here considered species inquirenda); (2) a combination of features (e.g., the mandibular condyle located behind the posterior edge of the vertical mandibular ramus, the angular process which projects laterally outwards, a bunolophodont crown morphology, a mesodont crown height, and a simple crown morphology of the third lobe of m3) indicates it belongs to the genus Platygonus; (3) this specimen corresponds to one of the largest South American peccaries; (4) taking into account certain anatomical characters as well as its morphometric range, this specimen is assigned to Platygonus cf. marplatensis. Even though the stratigraphic provenance of the specimen is still doubtful, it can be proposed that (1) it could be one of the most ancient records of tayassuids in South America, as would be expected given its geographical position, and (2) considering the new taxonomic proposal, this specimen represents the first record of Platygonus cf. marplatensis in Colombia and represents one of the northernmost South American records of the genus. This new interpretation would be of great relevance in the Great American Biotic Interchange due to its strategic geographical proximity to the Isthmus of Panama.
Selenogonus narinoensis是由Stirton(1947)根据在哥伦比亚Nariño省Cocha Verde地区的沉积物中发现的单一标本描述的,暂定为晚上新世-更新世(MGN 931;IGM p002118, Geológico国家博物馆,Geológico哥伦比亚博物馆,波哥大)。然而,对标本的形态学研究和比较形态学观察表明:(1)没有诊断特征支持种Selenogonus narinoensis(这里被认为是种inquirenda)的有效性;(2)下颌髁突位于下颌垂支后缘后方,有向外侧突出的角状突起,齿冠呈束状,齿冠高度为中齿状,齿冠形态为m3第三叶的简单齿冠;(3)该标本属于南美洲最大的一种鱼类;(4)考虑到某些解剖特征及其形态测量范围,该标本归属于Platygonus cf. marplatensis。尽管该标本的地层来源仍有疑问,但可以提出:(1)它可能是南美洲最古老的tayassuids记录之一,正如其地理位置所预期的那样;(2)考虑到新的分类建议,该标本代表了哥伦比亚Platygonus cf. marplatensis的第一个记录,并且代表了南美洲最北端的Platygonus属记录之一。这种新的解释将是非常相关的大美国生物交换由于其战略地理位置接近巴拿马地峡。
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引用次数: 1
Osteology of the Permian temnospondyl amphibian Glanochthon lellbachae and its relationships 二叠纪temnospondyl两栖动物Glanochthon lellbachae的骨学及其关系
IF 1.4 4区 地球科学 Q3 Earth and Planetary Sciences Pub Date : 2021-03-23 DOI: 10.5194/FR-24-49-2021
R. Schoch
Abstract. The early Permian Meisenheim Formation of the Saar–Nahe Basin(Germany) is famous for its richness in vertebrate fossils, among which thetemnospondyls were present with microvores and fish-eating apex predators.The latter trophic guild was occupied exclusively by the genusSclerocephalus in that basin within a long time interval up to M8, whereas in M9, a newtaxon, Glanochthon lellbachae, appeared. This taxon is defined by (1) a preorbital region 1.8–2.0 times as long as the postorbital skull table, (2) dermal ornament with tallradial ridges, (3) a prefrontal anteriorly wider with straight lateral margin,(4) a squamosal posteriorly only half as wide as the quadratojugal, (5) phalangesof manus and pes long and gracile, (6) carpals unossified in adults, and (7) tail substantially longer than skull and trunk combined. Phylogeneticanalysis finds that G. lellbachae forms the basal sister taxon of the stratigraphicallyyounger G. angusta and G. latirostre and that this clade nests within the paraphyletic taxonSclerocephalus, with S. nobilis forming the sister taxon of the genus Glanochthon (urn:lsid:zoobank.org:act:3038F794-17B9-4FCA-B241-CCC3F4423651; registration date: 15 March 2021).
摘要德国萨尔-纳河盆地早二叠世Meisenheim组以丰富的脊椎动物化石而闻名,其中temnospondyls存在微食动物和食鱼的顶端捕食者。在M8之前的很长一段时间间隔内,该盆地的后一个营养类群完全被硬头类占据,而在M9中出现了一个新的分类群Glanochthon lellbachae。这个分类群的定义是:(1)眶前区域是眶后颅骨表的1.8-2.0倍长,(2)真皮装饰有高的径向脊,(3)前额前部宽,有直的外侧边缘,(4)鳞片后部仅为方形颚的一半宽,(5)手和足的指骨长而柔软,(6)成年未分化的腕骨,(7)尾巴比颅骨和躯干加起来长得多。系统发育分析发现,G. lellbachae是地层较年轻的G. angusta和G. latirostre的基础姐妹分类群,该分支群在副进化分类群sclerocephalus中巢,S. nobilis形成Glanochthon属的姐妹分类群(urn:lsid:zoobank.org:act: 3038f794 - 17b9 - 4fca - b244 - ccc3f4423651;注册日期:2021年3月15日)。
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引用次数: 3
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Fossil Record
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