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†Estelestes ensis (Mammalia, Metatheria) from the early Eocene of Baja California (Mexico) as a generalized polydolopimorphian †Estelestes ensis(哺乳纲,后生动物目)来自始新世早期的下加利福尼亚(墨西哥),是一种广义的多食动物
IF 1.4 4区 地球科学 Q3 PALEONTOLOGY Pub Date : 2023-01-16 DOI: 10.1017/jpa.2022.105
F. Goin, E. C. Vieytes, V. Crespo, É. Oliveira
Estelestes ensis Novaceck et al., 1991 is a curious Paleogene metatherian mammal recognized on the basis of a single specimen from Baja California (Mexico) in southern North America. It comes from early Eocene (Wasatchian age) levels of the Las Tetas de Cabra Formation at “Marsupial Hill” in the Lomas Las Tetas de Cabra site (also known as Punta Prieta; see Novaceck et al., 1991). The specimen consists of a fragmentary left mandible with the last premolar, the roots of the first two molars, and almost complete last two molars (Fig. 1). It was referred to the Didelphini (Marsupialia, Didelphimorphia, Didelphidae, Didelphinae) even though Novaceck et al. (1991) stated that the overall morphology of the type specimen poses intriguing problems regarding its relationships. For example, the very deep, robust jaw of Estelestes distinguishes it from any other Holarctic “didelphine” (at the time Novaceck et al., 1991 published their work, both the concept and extent of Didelphidae and Didelphinae were much broader than today). Interestingly, they concluded that Estelestes had close affinities with “Mirandotherium” (lapsus calami for Mirandatherium), from the early Eocene of Itaboraí, in southeastern Brazil. “Resemblance between the two taxa is nevertheless striking, once again raising the possibility of close relationships among certain early members of the Northern Hemisphere and South American Didelphinae” (Novaceck et al., 1991, p. 16). The affinities of Mirandatherium are contested, having been regarded as part of the Didelphimorphia (e.g., de Paula Couto, 1952a) or Microbiotheria (e.g., Marshall, 1987; McKenna and Bell, 1997; Oliveira and Goin, 2011), or even as an alphadontian (Carneiro, 2019).
Estelestes ensis Novaceck et al., 1991,是一种奇怪的古近纪后生哺乳动物,是在北美南部下加利福尼亚(墨西哥)的一个单一标本的基础上被识别出来的。它来自始新世早期(Wasatchian时代)的Las Tetas de Cabra组,位于Lomas Las Tetas de Cabra遗址(也称为Punta Prieta;参见Novaceck et al., 1991)。该标本由一个残缺的左下颌骨和最后的前磨牙、前两颗磨牙的根部以及几乎完整的后两颗磨牙组成(图1)。尽管Novaceck等人(1991)指出,该模式标本的整体形态对其关系提出了有趣的问题,但它被称为Didelphini(有袋目,didelphimorphiia, Didelphidae, Didelphinae)。例如,Estelestes非常深、强壮的颚使它与任何其他的全北极“didelphine”(在Novaceck等人1991年发表他们的工作时,Didelphidae和Didelphinae的概念和范围都比今天广泛得多)区分。有趣的是,他们得出结论,Estelestes与巴西东南部Itaboraí始新世早期的“Mirandatherium”(Mirandatherium的lapsus calami)有着密切的关系。“尽管如此,这两个分类群之间的相似之处还是惊人的,这再次提出了北半球和南美洲Didelphinae的某些早期成员之间存在密切关系的可能性”(Novaceck et al., 1991, p. 16)。Mirandatherium的亲缘关系是有争议的,被认为是didelphimorphiia(例如,de Paula Couto, 1952a)或Microbiotheria(例如,Marshall, 1987;麦肯纳和贝尔,1997;Oliveira and Goin, 2011),或者甚至是作为一个字母专家(Carneiro, 2019)。
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引用次数: 0
Miocene instead of Jurassic: the importance of sound fieldwork for paleontological data analysis 中新世而非侏罗纪:良好的实地调查对古生物数据分析的重要性
IF 1.4 4区 地球科学 Q3 PALEONTOLOGY Pub Date : 2023-01-16 DOI: 10.1017/jpa.2022.106
F. Fürsich, Suraj Bhosale, Matthias Alberti, D. Pandey
Abstract. A diverse molluscan assemblage dominated by turritellid gastropods found in Kachchh, western India, has been interpreted in the past as Late Jurassic (Oxfordian) in age, based on associated undoubtedly Oxfordian ammonites. Recently, several investigations focused on the assemblage dealing with taxonomic, paleoecological, and evolutionary aspects. An analysis of the associated bivalve fauna, foraminiferal assemblage, and the geological context strongly suggests a Miocene rather than a Jurassic age and invalidates several conclusions drawn from the alleged Jurassic age of the fossils.
摘要在印度西部的Kachchh发现的一个以turritellid腹足类为主的多样化软体动物群落,在过去被解释为侏罗纪晚期(牛津阶),其年龄无疑是基于相关的牛津阶菊石。最近,一些研究集中在该组合的分类学、古生态学和进化方面。对相关双壳动物群、有孔虫组合和地质背景的分析有力地表明了中新世而非侏罗纪,并使从化石的侏罗纪时代得出的几个结论无效。
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引用次数: 2
Sympatric speciation driving evolution of Late Ordovician brachiopod Zygospira in eastern North America 同域物种形成驱动北美东部晚奥陶世腕足类合螺类进化
IF 1.4 4区 地球科学 Q3 PALEONTOLOGY Pub Date : 2023-01-16 DOI: 10.1017/jpa.2022.102
Colin D. Sproat, Jessica S.A. McLeod
Abstract. The morphology of Zygospira, an early atrypide brachiopod, was analyzed using a multivariate approach. Principal component analysis and discriminant analysis clearly differentiated species as they are currently defined primarily based on differences in shell size and ornamentation but not in terms of overall shell shape. The older Zygospira modesta was able to persist into the late Katian (Richmondian) while smaller early species in other brachiopod lineages mostly went extinct. This may have been possible through niche partitioning because the smaller shells have been found attached to other filter feeders and no larger species have been found in these associations so far. This could represent a rare example of sympatric speciation preserved in the fossil record. In the future, detailed study of the spiralia and their associated structures may provide clues as to the ultimate evolutionary affinities of this group in relation to the other atrypide brachiopods evolving at this time.
摘要采用多变量分析方法,对早期atrypide腕足动物Zygospira的形态进行了分析。主成分分析和判别分析明确区分了物种,因为它们目前主要根据壳的大小和纹饰的差异来定义,而不是根据整体壳的形状。较老的Zygospira modesa能够持续到Katian (Richmondian)晚期,而其他腕足动物谱系中较小的早期物种大多灭绝了。这可能是通过生态位划分实现的,因为较小的壳被发现附着在其他滤食性动物身上,到目前为止还没有在这些关联中发现较大的物种。这可能是化石记录中保存的同域物种形成的罕见例子。在未来,对螺旋体及其相关结构的详细研究可能会为这一群体与当时进化的其他非典型腕足动物的最终进化亲缘关系提供线索。
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引用次数: 1
JPA volume 97 issue 1 Cover and Front matter JPA第97卷第1期封面和封面事项
IF 1.4 4区 地球科学 Q3 PALEONTOLOGY Pub Date : 2023-01-01 DOI: 10.1017/jpa.2023.9
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引用次数: 0
The rangeomorph fossil Charnia from the Ediacaran Shibantan biota in the Yangtze Gorges area, South China 长江三峡地区埃迪卡拉纪石板滩生物群的轮虫化石
IF 1.4 4区 地球科学 Q3 PALEONTOLOGY Pub Date : 2022-12-06 DOI: 10.1017/jpa.2022.97
Chengxi Wu, K. Pang, Zhe Chen, Xiaopeng Wang, Chuanming Zhou, B. Wan, Xunlai Yuan, S. Xiao
The terminal Ediacaran Shibantan biota (~550–543 Ma) from the Dengying Formation in the Yangtze Gorges area of South China represents one of the rare examples of carbonate-hosted Ediacara-type macrofossil assemblages. In addition to the numerically dominant taxa—the non-biomineralizing tubular fossil Wutubus and discoidal fossils Aspidella and Hiemalora, the Shibantan biota also bears a moderate diversity of frondose fossils, including Pteridinium, Rangea, Arborea, and Charnia. In this paper, we report two species of the rangeomorph genus Charnia, including the type species Charnia masoni Ford, 1958 emend. and Charnia gracilis new species, from the Shibantan biota. Most of the Shibantan Charnia specimens preserve only the petalodium, with a few bearing the holdfast and stem. Despite overall architectural similarities to other Charnia species, the Shibantan specimens of Charnia gracilis n. sp. are distinct in their relatively straight, slender, and more acutely angled first-order branches. They also show evidence that may support a two-stage growth model and a epibenthic sessile lifestyle. Charnia fossils described herein represent one of the youngest occurrences of this genus and extend its paleogeographic and stratigraphic distributions. Our discovery also highlights the notable diversity of the Shibantan biota, which contains examples of a wide range of Ediacaran morphogroups. UUID: http://zoobank.org/837216cd-4a4a-4e13-89e2-ee354ba48a4c
中国南部长江三峡地区灯影组的末期埃迪卡拉纪石板滩生物群(~550–543 Ma)是碳酸盐岩型埃迪卡拉纪大型化石组合的罕见实例之一。除了数量上占主导地位的分类群——非生物矿化的管状化石Wutubus和盘状化石Aspidella和Hiemalora之外,石板滩生物群还拥有适度多样性的叶化石,包括蕨类植物、兰芝属、乔木属和夏尼亚属。本文报道了野蛙属的两个种,包括模式种马索尼·福特蛙,1958年修订。和石板滩生物群中的细柄炭新种。石板滩茶的大部分标本只保存了花瓣状茎,少数标本带有茎和茎。尽管在整体结构上与其他Charnia物种相似,但细柄Charnia n.sp.的石板滩标本在相对笔直、细长和角度更尖锐的一级分支方面是不同的。它们还显示了可能支持两阶段生长模式和表海底固着生活方式的证据。本文描述的夏尼亚化石代表了该属最年轻的化石之一,并扩展了其古地理和地层分布。我们的发现还突出了石板滩生物群的显著多样性,其中包含了广泛的埃迪卡拉纪形态群的例子。UUID:http://zoobank.org/837216cd-4a4a-4e13-89e2-ee354ba48a4c
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引用次数: 0
Needmorella, a new trilobite genus of the Synphoriinae (Dalmanitidae) from the Lower–Middle Devonian of West Virginia 西维吉尼亚泥盆纪中下泥盆世三叶虫科三叶虫属Needmorella
IF 1.4 4区 地球科学 Q3 PALEONTOLOGY Pub Date : 2022-12-06 DOI: 10.1017/jpa.2022.96
D. Holloway, Brian M. Scott
Abstract. The trilobite Needmorella new genus, with type species N. simoni new genus new species from the late Emsian to mid-Eifelian Needmore Shale of West Virginia, is a distinctive member of the subfamily Synphoriinae. It also occurs in the same formation in Pennsylvania and Virginia. It is not very similar to other Devonian representatives of the subfamily and is considered to have its origins in a morphologically less-derived ancestor because it shares certain similarities with Silurian genera, including the very short anterior cephalic border unmodified by crenulations or spines, S2 that is not largely reduced to a deep pit adaxially, the relatively low inflation of L3, and the well-defined interpleural furrows on the pygidium. Other particularly distinctive characters of the genus include the very long genal spines and the abaxially inflated and expanded posterior pleural bands on the thorax and pygidium that project slightly distally. The conventional concept of the Devonian synphoriine Anchiopsis Delo, 1935 appears to be incompatible with the holotype of the type species, judging from the early illustrations of the specimen, and the genus could be a synonym of Synphoria Clarke, 1894.
摘要三叶虫Needmorella新属,模式种N.simoni新属新种,产于西弗吉尼亚州埃姆辛晚期至艾菲尔阶中期的Needmore页岩,是Synphoriinae亚科的一个独特成员。它也发生在宾夕法尼亚州和弗吉尼亚州的同一地层中。它与该亚科的其他泥盆纪代表不太相似,被认为起源于一个形态上不太同源的祖先,因为它与志留纪属有某些相似之处,包括非常短的未被细齿或棘修饰的头部前边界,S2在很大程度上没有正面缩小为深坑,L3的膨胀率相对较低,以及在pygidium上明确的胸膜间沟。该属的其他特别独特的特征包括非常长的genal棘和胸部和肿胀的后部胸膜带,这些胸膜带稍微向远侧突出。根据该标本的早期插图判断,1935年泥盆纪的同生珊瑚Anchiopsis Delo的传统概念似乎与模式物种的正模不兼容,该属可能是1894年的同生孢子虫Clarke的同义词。
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引用次数: 0
Cornulitid tubeworms and other calcareous tubicolous organisms from the Hirmuse Formation (Katian, Upper Ordovician) of northern Estonia 爱沙尼亚北部赫斯穆斯组(卡提亚,上奥陶统)的角质管虫和其他钙质管状生物
IF 1.4 4区 地球科学 Q3 PALEONTOLOGY Pub Date : 2022-12-05 DOI: 10.1017/jpa.2022.89
O. Vinn, A. Madison, Mark A. Wilson, U. Toom
Abstract. Seven species of cornulitids, one unidentified tubicolous shell, and the problematic bryozoan Lagenosypho Spandel, 1898 are here described from the Katian of Baltica. Three new species—Cornulites lindae new species, Cornulites meidlai new species, and Conchicolites kroegeri new species—are described. The unidentified tubicolous organism has punctate shell structure and setae-like structures that can best be affiliated with lophophorates. The Hirmuse fauna indicates that the diversity and number of cornulitids in the Ordovician of Baltica has been underestimated and it is likely that the Baltic cornulitid fauna was as diverse and abundant as the fauna of Laurentia. Clay mud-bottom environments supported the highest cornulitid diversity in the Late Ordovician of Baltica. The occurrence of intermediate forms indicates that some tentaculitid characters, e.g., regular annulation and a nearly straight shell, which were thought to be apomorphies of free-living tentaculitids, were actually inherited from ancestral cornulitids. The cornulitid fauna of the Late Ordovician of Laurentia somewhat resembles the cornulitid fauna of the Late Ordovician of Baltica, but there are fewer common faunal elements between Gondwana and Baltica.
摘要这里描述了七种玉米粒虫,一种未知的管状外壳,以及1898年有问题的苔藓虫Lagenosypho Spandel,来自Baltica的Katian。介绍了三个新种——林达麻麻新种、美德莱麻麻新种和克氏海螺新种。这种未经鉴定的管状生物具有点状外壳结构和刚毛状结构,这些结构最适合与扁孔虫属联系在一起。Hirmuse动物群表明,在波罗的海奥陶纪,人们低估了玉米粒的多样性和数量,波罗的海玉米粒动物群很可能与Laurentia动物群一样多样和丰富。粘土泥底环境支持了巴尔蒂卡晚奥陶世最高的麻粒岩多样性。中间形态的出现表明,一些被认为是自由生活的触手类的变形体的触手状特征,如规则环状和几乎直的外壳,实际上是从祖先的玉米粒继承而来的。Laurentia晚奥陶世的玉米粒动物群与Baltica晚奥陶纪的玉米粒植物群有些相似,但冈瓦纳大陆和Baltica大陆之间的常见动物群元素较少。
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引用次数: 1
A new Early Paleogene fossil mammal locality in the central-eastern Nemegt Basin, Gobi Desert, Mongolia, and notes on mammalian biostratigraphy 蒙古戈壁沙漠Nemegt盆地中东部早古近系哺乳动物化石新地点及哺乳动物生物地层学注释
IF 1.4 4区 地球科学 Q3 PALEONTOLOGY Pub Date : 2022-12-05 DOI: 10.1017/jpa.2022.85
K. Yo, Eva A. Hoffman, M. O'Leary, M. Novacek
Abstract. We report new, fossiliferous Paleogene Naran Bulak Formation localities from the central-eastern part of the Nemegt Basin of the Gobi Desert, Mongolia. Early Paleogene localities have been identified previously only in the western half of the Nemegt Basin. The new localities, near the town of Daus, are also noteworthy for their geographical proximity to Ukhaa Tolgod, a Late Cretaceous Djadokhta Formation locality known for its numerous dinosaur, mammal, and lizard fossils. The Daus section consists of the Zhigden, Naran, and Bumban members of the Naran Bulak Formation at three localities, and mammal and ostracode fossils were discovered in the Naran Member. Noteworthy discoveries are a dentary of the pantodont Archaeolambda cf. A. planicanina, postcrania of Pantolambdodon, a skull of the gliroid Gomphos, and a partial skull with a worn and damaged dentition provisionally identified as an arctostylopid. Biostratigraphy has been the primary means of dating Paleogene Asian faunas, however, the local fauna from the new localities does not fit easily with established patterns. The Naran Member and Archaeolambda planicanina and the arctostylopid Palaeostylops typically have been allied with the Gashatan Asian Land Mammal Age (ALMA) and attributed to the latest Paleocene. By contrast, Gomphos repeatedly has been found in the Bumban Member and assigned a Bumbanian ALMA, which has been considered as the earliest Eocene. Pantolambdodon has been reported from middle Eocene Arshatan and Irdinmanhan ALMA beds. The co-occurrence of these taxa in Naran Member beds complicates the temporal interpretation of the new localities and the reliability of broader biostratigraphic patterns.
摘要我们报道了来自蒙古戈壁沙漠涅美格特盆地中东部的新的、含化石的古近系纳兰布拉克组。古近纪早期的位置以前只在涅美格特盆地的西半部发现。Daus镇附近的新地区也因其地理位置接近Ukhaa Tolgod而引人注目,Ukhaa托尔god是晚白垩世Djadokhta组的一个地区,以其众多的恐龙、哺乳动物和蜥蜴化石而闻名。Daus段由三个位置的Naran Bulak组的Zhigden、Naran和Bumban成员组成,在Naran成员中发现了哺乳动物和介形虫化石。值得注意的发现是Pantodon Archeolambda的一个牙齿。a.planicanina,Pantolambdodon的后颅,一个齿列体Gomphos的头骨,以及一个部分头骨,其齿列磨损和受损,暂时被确定为弓形齿。生物地层学一直是确定古近系亚洲动物群年代的主要手段,然而,新地区的当地动物群并不容易符合既定的模式。纳兰成员、平原古龙目和弓形古柱龙目通常与加沙坦亚洲陆地哺乳动物时代(ALMA)有关,并被认为是最新的古新世。相比之下,Gomphos在Bumban段多次被发现,并被指定为Bumbanian ALMA,这被认为是最早的始新世。在始新世中期的阿尔沙坦和Irdinmanhan ALMA地层中,已经报道了Pantlambdodon。这些分类群在纳兰成员层中的共存使新地点的时间解释和更广泛的生物地层学模式的可靠性变得复杂。
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引用次数: 0
Revision of Eothinoceras and the status of the Eothinoceratidae (Cyrtocerinida, Multiceratoidea, Cephalopoda) 青角鲸的修订和青角鲸科的地位(圆角鲸目,多角总目,头足目)
IF 1.4 4区 地球科学 Q3 PALEONTOLOGY Pub Date : 2022-12-05 DOI: 10.1017/jpa.2022.99
D. H. Evans, Marcela Cichowolski
Abstract. The type specimens of Eothinoceras americanum Ulrich et al., 1944, type species of the genus Eothinoceras Ulrich et al., 1944, are revised based on new photographic material. The resulting interpretation of the conch shape of Eothinoceras shows that the type species is an endogastric cyrtocone, necessitating the restriction of Eothinoceras to the type species and requiring the revision of the family Eothinoceratidae and the order Cyrtocerinida. A new scheme for classifying the genera of the order Cyrtocerinida into its three families is proposed: (1) Eothinoceratidae Ulrich et al., 1944, containing the genera Eothinoceras; Protothinoceras Chen and Teichert, 1987; Mesothinoceras Chen and Teichert, 1987; and Conothinoceras Chen and Teichert, 1987; (2) Cyrtocerinidae Flower, 1946, including Cyrtocerina Billings, 1865; Tangshanoceras Chen, 1976; and Centrocyrtocerina Stait, 1983; (3) Bathmoceratidae Gill, 1871, containing Bathmoceras Barrande, 1865; Saloceras Evans, 2005; Sacerdosoceras Evans, 2005; Margaritoceras Cecioni and Flower, 1985; and Mutveiceras Cichowolski et al., 2014.
摘要根据新的摄影材料,对美洲Eothinoceras Ulrich等人,1944年的模式标本和Eothinocaras Ulriche等人,1944的模式种进行了修订。由此对Eothinoceras海螺形状的解释表明,该模式物种是胃内cyrtocone,因此必须将Eothinocaras限制在该模式物种范围内,并需要对Eothanoceridae科和Cyrtocerinida目进行修订。提出了一种将Cyrtocerinida目属分为三科的新方案:(1)Eothinoceridae Ulrich et al.,1944,包含Eothinocaras属;原细角类Chen和Teichert,1987;Mesothinoceras Chen和Teichert,1987;以及Conothinoceras Chen和Teichert,1987年;(2) Cyrtocerinidae Flower,1946年,包括Cyrtocrina Billings,1865年;Tangshanoceras Chen,1976;和Centrocyroccerina Stait,1983年;(3) Bathmoceratidae Gill,1871年,包含Bathmoceras Barrande,1865年;Saloceras Evans,2005年;Sacerdoseceras Evans,2005年;Margaritoceras Cecioni和Flower,1985年;以及Mutweiceras Cichowolski等人,2014年。
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引用次数: 0
Evidence of large sturgeons in the Paleocene of North America 北美古新世大型鲟鱼的证据
IF 1.4 4区 地球科学 Q3 PALEONTOLOGY Pub Date : 2022-11-17 DOI: 10.1017/jpa.2022.87
C. Brownstein
Abstract. Ray-finned fishes comprise nearly half of extant vertebrate species and include several ancient lineages with fossil records that stretch over 200 Myr in time. One of these old clades, the sturgeons and paddlefishes, is distributed across the Northern Hemisphere and includes some of the largest known freshwater fishes. Yet, the fossil record of this lineage (Acipenseriformes) is poor compared to similarly ancient ray-finned fish clades. Here, I describe sturgeon fossils from two geological units in North America < 10 Myr younger than the Cretaceous-Paleogene boundary. Both come from individuals of ∼1.5 m in length. These Paleogene forms establish the long history of large body size in Acipenseriforms and reveal sturgeons were some of the largest inhabitants of freshwater ecosystems that were still recovering from the Cretaceous-Paleogene extinction.
摘要射线鳍鱼类占现存脊椎动物的近一半,包括几个古老的谱系,化石记录的时间跨度超过200 Myr。其中一个古老的分支,鲟鱼和白鲟,分布在北半球,包括一些已知最大的淡水鱼。然而,与同样古老的射线鳍鱼类分支相比,这一谱系(鲟形目)的化石记录很差。在这里,我描述了北美洲两个地质单元的鲟鱼化石,它们比白垩纪-古近纪边界年轻<10 Myr。两者都来自身长约1.5米的个体。这些古近纪形态奠定了鲟鱼体型庞大的悠久历史,并揭示了鲟鱼是淡水生态系统中最大的居民之一,这些生态系统仍在从白垩纪-古近纪灭绝中恢复。
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引用次数: 1
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Journal of Paleontology
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