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Some rare helical Ammonoidea from the lower Aptian (Lower Cretaceous) of the Paris Basin (NE France) 法国东北部巴黎盆地(下白垩统)下阿普提亚(下白垩统)罕见的螺旋形菊石
IF 1 4区 地球科学 Q3 Earth and Planetary Sciences Pub Date : 2023-04-01 DOI: 10.1016/j.annpal.2023.102601
Stéphane Bersac , Didier Bert

In the present paper, we describe and figure some rare lower Aptian (Lower Cretaceous) helical ammonoids from the Deshayesites deshayesi Zone of the Argiles à Plicatules Formation (Paris Basin, NE France). Those are the only helical ammonoids known in this zone so far. They were only occasionally evoked in literature and this is probably on the basis of the specimens here described that, according to us, the genus Toxoceratoides (Helicancylidae) was wrongly regarded as possessing an early helical spiral part. We demonstrate here that these specimens are not belonging to Toxoceratoides nor Helicancylidae: they are interpreted as ‘abnormally’ helical representatives of the genus Ancyloceras (Ancyloceratidae) or a new taxon of Ancyloceratidae.

在本文中,我们描述并描绘了一些罕见的下阿普第阶(下白垩纪)螺旋菊石,它们来自ArgilesàPlicatules组(法国东北部巴黎盆地)的Deshayesites deshayesi带。到目前为止,这些是该地区已知的唯一的螺旋菊石。它们只是偶尔在文献中被提及,这可能是基于这里描述的标本,根据我们的说法,Toxoceratoides属(Heliancylide)被错误地认为拥有早期的螺旋状部分。我们在这里证明,这些标本既不属于弓形虫,也不属于螺旋虫科:它们被解释为钩角虫属(钩角虫科)的“异常”螺旋代表,或是钩角虫科新的分类单元。
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引用次数: 0
New data on the Devonian and Carboniferous Graptolithina (Dendroidea) from Belgium with notes on possible occurrences of Rhabdopleuridae in the Belgian Carboniferous 比利时泥盆纪和石炭纪Graptolithina(Dendroidea)的新资料以及比利时石炭纪Rhabdopleuridae可能出现的注释
IF 1 4区 地球科学 Q3 Earth and Planetary Sciences Pub Date : 2023-04-01 DOI: 10.1016/j.annpal.2023.102612
Bernard Mottequin , Jörg Maletz , Stijn Goolaerts

An old find of a graptolite from the Eifelian Jemelle Formation, combined with the recent discovery of an exceptionally well-preserved specimen from the Upper Frasnian (Matagne Formation), allow documentation of the geologically oldest and youngest dendroid fossils from the Devonian of Belgium. Both are ascribed to the long-ranging genus Callograptus (Acanthograptidae). These records enlarge the sparse knowledge of graptolite diversity and occurrences along the southern margin of Laurussia during the mid-Paleozoic. In Belgium, Devonian dendroids are exclusively known from the Dinant Synclinorium. The Devonian dendroid faunas previously reported from the Eifelian, Givetian and Frasnian of the Ardenne–Rhenish massifs (Belgium and Germany) are of low diversity (one to five species per locality). The specimens, generally inedaquatelly preserved, were ascribed to the genera Callograptus, Dictyonema, Palaeodictyota?, and Ruedemannograptus? In these massifs, Carboniferous dendroids are only known from the Viséan (Moliniacian) Denée Konservat-Lagerstätte. Other but undocumented reports of dendroids (and rhabdopleurids) are from the historical type area of the Tournaisian Stage (Tournai area, Brabant Parautochthon). Rhabdopleuridae from the Viséan–Serpukhovian of the Campine Basin (Turnhout borehole) remain incompletely known in the absence of the previously described material.

来自Eifelian Jemelle组的笔石的古老发现,加上最近发现的来自上弗拉斯纪(Matagne组)的保存异常完好的标本,使得比利时泥盆纪最古老和最年轻的树突化石的记录成为可能。两者都属于分布较广的棘蝗属(棘蝗科)。这些记录扩大了中古生代Laurussia南缘笔石多样性和产状的稀疏知识。在比利时,泥盆纪的树状体只在迪南向斜中被发现。先前在阿登-莱茵块体(比利时和德国)的埃弗勒、吉文和弗拉斯尼亚报道的泥盆纪树状动物群的多样性很低(每个地方1 - 5种)。这些标本一般保存不完整,归属于Callograptus属、Dictyonema属、Palaeodictyota?和Ruedemannograptus?在这些地块中,石炭纪的树状体只在vissaman (Moliniacian) densame Konservat-Lagerstätte中被发现。其他未记载的树突类(和横纹肌类)的报告来自Tournaisian阶段的历史类型区域(Tournai地区,Brabant Parautochthon)。由于缺乏先前描述的材料,Campine盆地vis - serpukhovian (Turnhout钻孔)的Rhabdopleuridae仍然不完全清楚。
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引用次数: 0
Records and palaeoenvironmental conditions of Early Miocene scleractinian corals from the Fırat formation (Diyarbakır, Turkey) Fırat组早中新世巩膜珊瑚的记录和古环境条件(土耳其迪亚巴克尔)
IF 1 4区 地球科学 Q3 Earth and Planetary Sciences Pub Date : 2023-04-01 DOI: 10.1016/j.annpal.2023.102613
İhsan Ekin , Nikolaos Tsaparas

Miocene scleractinian corals were documented for the first time in the Fırat Formation of Sarıca village, Eğil district of Diyarbakır, SE Turkey. The fauna consists of four hermatypic species belonging to three families: Diploastrea taurinensis (d’Achiardi, 1868), Tarbellastraea reussiana (Milne-Edwards and Haime, 1850), Echinopora gemmacea (Lamarck, 1816) and Montastraea pelouaensis (Chevalier, 1954). The morphospecies described here are typical of the early Miocene (Aquitanian-Burdigalian) and show that the Fırat Formation coral community was composed of shallow and subtropical reefs of Mediterranean Tethys palaeobiogeographical affinity. The examined species are reef components found in tropical and subtropical oceans, as are the majority of zooxanthellae. The species were upper photic zone related, had massive colonial growth morphology, and lived in a shallow tropical-subtropical marine palaeoenvironment at a depth of up to 50 m. They grew on hard ground, clean water, and normal salinity and required a minimum seawater temperature of 18 ̊C; however, they were most abundant at water temperatures ranging from 22 to 26 ̊C in high-moderate water energy. Consequently, this fossil assemblage will not only improve our knowledge of scleractinian coral diversity and morphological variation in SE Turkey but will also provide an important background for future studies of coral taxonomy, diversity and biogeography of the region.

在土耳其东南部Diyarbakır Eğil地区Sarıca村Fırat组首次记录到中新世核状珊瑚。区系由3科4个雌雄同体种组成:taurinensis Diploastrea (d 'Achiardi, 1868)、Tarbellastraea reussiana (Milne-Edwards and Haime, 1850)、Echinopora gemmacea (Lamarck, 1816)和Montastraea pelouaensis (Chevalier, 1954)。本文所描述的形态种是典型的早中新世(阿基坦-布尔迪亚),表明Fırat组珊瑚群落由地中海特提斯古地理亲缘的浅海和亚热带珊瑚礁组成。研究的物种是在热带和亚热带海洋中发现的珊瑚礁成分,大多数虫黄藻也是如此。该物种与上层光带有亲缘关系,具有庞大的群体生长形态,生活在深度达50 m的热带-亚热带浅海古环境中。它们生长在坚硬的地面、干净的水和正常的盐度,需要最低18℃的海水温度;而在中高水能水温22 ~ 26℃时最为丰富。因此,这一化石组合不仅将提高我们对土耳其东南部石核期珊瑚多样性和形态变异的认识,而且将为该地区未来珊瑚分类、多样性和生物地理学的研究提供重要的背景。
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引用次数: 0
First record of hybodont egg capsules from the Jurassic of Thailand 泰国侏罗纪首次记录到双齿龙卵囊
IF 1 4区 地球科学 Q4 PALEONTOLOGY Pub Date : 2023-01-01 DOI: 10.1016/j.annpal.2023.102652
Gilles Cuny , Phornphen Chanthasit
Two egg capsules of Palaeoxyris sp. are reported from the Jurassic Phu Kradung Formation in the Khorat Plateau, northeastern Thailand. As the record of this genus remains rare in the Jurassic, it complements its stratigraphic distribution. Most importantly, it represents the first unambiguous evidence that some hybodont sharks from the Mesozoic of Thailand were able to reproduce in fresh waters, even if it is not yet possible to identify which species in particular.
在泰国东北部的呵叻高原发现了两枚卵形蒴果。这些标本来自侏罗纪的普卡东地层。该属在侏罗纪的化石记录仍然鲜为人知,此次发现完善了其地层分布。这些卵裂囊是中生代某些杂交鲨鱼在泰国淡水中繁殖的第一个不可否认的证据,尽管目前还无法确定具体是哪个物种。
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引用次数: 0
Insights into dwarf stegodon (Stegodon florensis florensis) palaeobiology based on rib histology 基于肋骨组织学对矮盗龙古生物学的启示
IF 1 4区 地球科学 Q4 PALEONTOLOGY Pub Date : 2023-01-01 DOI: 10.1016/j.annpal.2023.102654
Pauline Basilia , Justyna J. Miszkiewicz , Julien Louys , Unggul Prasetyo Wibowo , Gerrit D. van den Bergh
Stegodon florensis florensis is an example of a diminutive insular stegodon from the early Middle Pleistocene bonebeds of Mata Menge, So’a Basin (Flores, Indonesia). We report the first insights into this Stegodon's bone histology, which furthers our understanding of this species’ palaeobiology, and evaluate the micro-anatomy and metabolic microstructural products retained in well-preserved rib bone tissue. A proximal rib fragment selected from a larger assemblage was sampled at three anatomical regions from which histological sections were produced. The sampling regions differed in cortical width size ranging from relatively thick to thin (17.94 mm to 6.31 mm). Bone tissue arrangement and histomorphometric variables measuring histological products of adult bone remodelling processes (secondary osteon population density and area, bone area, and Haversian canal percentage) were examined. The preservation of bone histology was very good, showing widespread Haversian bone with indication of several generations of remodelling processes, multiple resorption cavities, and the presence of ‘super’ (unusually large) osteons. Secondary osteon density and geometry varied with cortical width through the serial sections whereby the section with the narrowest cortex produced the highest secondary osteon density but also the largest osteon area. The effects of medullary cavity infilling on the endocortical region in the Stegodon rib may also relate to the adjacent periocortical region. The rib histology of this intermediate-sized dwarfed species shows, for the first time, a degree of secondary bone active remodelling in a Stegodon that indicates this was possibly a long-lived individual. Microscopic bone characteristics shared with other extinct and extant proboscideans may be useful in identifying Stegodon from fragmented mixed fossil assemblages.
是印度尼西亚弗洛勒斯岛 So'a 盆地 Mata Menge 中更新世早期骨骼组合中的一个岛屿矮小盗齿兽的例子。我们报告了对这一物种骨骼组织学的初步认识,这加深了我们对这一物种古生物学的理解,并评估了保存完好的化石肋骨骨组织中保留的微解剖学和微结构代谢产物。从一大批化石中选取了一块近端肋骨片段,在三个解剖区域取样并进行组织学切片。取样区域的皮质宽度从相对较厚到较薄(17.94 毫米到 6.31 毫米)不等。对骨组织的布局和衡量成人骨重塑过程组织学特征的组织形态计量变量(次级骨细胞群的密度和表面积、骨表面积和哈弗氏管的百分比)进行了检查。骨组织学保存非常完好,显示出具有多代重塑过程的哈弗斯骨、多个吸收腔以及异常大的骨骨(超级骨骨)的存在。次生骨小梁的密度和几何形状随切片皮质宽度的变化而变化,因此皮质厚度越薄,次生骨小梁的密度越大,骨小梁覆盖的表面积也越大。髓腔充填对肋骨皮质内区域的影响可能会延伸到邻近的皮质周围区域。这种中型矮小动物的肋骨组织学首次表明,长寿动物的二次骨重塑具有一定程度的活跃性。与其他已灭绝和现存长鼻目动物共享的显微骨骼特征可能有助于从支离破碎的混合化石群中进行鉴定。
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引用次数: 0
Luzon predators: Clues from a fossil with bite marks 吕宋岛的掠食者带有咬痕的化石提供的线索
IF 1 4区 地球科学 Q4 PALEONTOLOGY Pub Date : 2023-01-01 DOI: 10.1016/j.annpal.2023.102649
Meyrick U. Tablizo , Rebekka Volmer , Allan Gil S. Fernando , Juan C. Rofes
Luzon Island remained isolated throughout its emergent history with its paleofauna often considered depauperate. However, the terrestrial vertebrate fossil record of the island is still poorly known. A new fossil-bearing paleochannel deposit was discovered in La Union Province, northwestern Luzon, Philippines. This paleochannel is inferred to be of Pleistocene age and has yielded fossils tentatively classified as a bovid (Bubalus sp.), a cervid (Rusa sp.), a suid (Sus sp.), and a large crocodile (Crocodylidae). Interestingly, a proximal fragment of the left tibia of a cervid shows multiple bite marks. This study analyzes the bite marks and modifications on the specimen to infer its possible trace maker and explore its implications in understanding the ancient predatory guild of Luzon. Three sets of modifications were recognized on the specimen, namely the bite marks, the broken proximal end and the dry bone fractures. A crocodilian agent for the bite marks is supported by the bisections, their occurrence in the paleofaunal assemblage and the fluvial setting of the deposit. However, an unknown carnivoran agent interpretation cannot be fully disregarded given the ambiguity of bisected marks and the possibly gnawed proximal end of the specimen. Unfortunately, given the lack of in situ context and apparent overprinting, the extent of the peri- and post-mortem modification remains uncertain which hinders the identification of the predatory agent. Verification of the observations reported in this study would require the systematic collection and detailed taphonomic analysis of additional fossil specimens from the deposit.
卢松岛在历史上一直与世隔绝,其古生物群通常被认为是贫乏的。然而,人们对该岛的陆生脊椎动物化石仍然知之甚少。在菲律宾吕宋岛西北部的拉乌尼翁省发现了一个新的古河道化石矿床。这条古河道被认为可以追溯到更新世,出土的化石被暂时归类为牛科(sp.)、鹿科(sp.)、麂科(sp.)和大型鳄鱼科(Crocodylidae)。值得注意的是,左侧颈鹿胫骨的近端碎片上有多处咬痕。我们对这些咬痕的分析表明了它们的来源。这对吕宋岛古代食肉动物群的组成具有重要意义。在描述的标本上发现了三个系列的变化:咬痕、近端断裂和干骨上的骨折。观察到的痕迹可能与动物群中鳄鱼的痕迹一致,并且与沉积物的水流环境相吻合。然而,由于咬痕和近端被啃咬的痕迹模糊不清,不能完全排除另一种未知食肉动物的解释。遗憾的是,由于缺乏背景资料和明显的叠印,这些痕迹的改变程度仍不确定,从而阻碍了对食肉动物的最终鉴定。需要对遗址中的其他化石标本进行进一步观察、系统收集和详细的岩石学分析。
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引用次数: 0
Morphology of teleosaurid osteoderms from the Phu Kradung Formation of Thailand 泰国普卡东地层远东龙类骨器的形态学
IF 1 4区 地球科学 Q4 PALEONTOLOGY Pub Date : 2023-01-01 DOI: 10.1016/j.annpal.2023.102653
Supanut Bhuttarach , Uthumporn Deesri , Prapasiri Warapeang , Nonthiwat Taesuk , Komsorn Lauprasert
This is a morphological analysis of the osteoderm patterns of Indosinosuchus from the Late Jurassic Phu Noi excavation site in Kalasin Province, Thailand. Data on 203 specimens indicated that 197 osteoderms belongs to the teleosaurid genus Indosinosuchus. Out of 203 total, 109 specimens are dorsal osteoderm morphotypes (M1 to M5) with sub-square, sub-rectangular, or rhomboidal in shape, including keel and peg. Another 88 are ventral osteoderms (M6 and M7) with rectangular or sub-square osteoderms, obscure keel and peg. Large subcircular ellipsoid pits with an anterior facet bar are discovered in both dorsal and ventral osteoderms. Data on morphological features enabled osteoderm positions of Indosinosuchus to be described. The anterior cervical osteoderms are characterized by an obscure keel and a small sub-square shape (M1). In the posterior trunk region, the osteoderms increase in width and adopt a sub-rectangular shape (M2). These osteoderms decrease in size at the sacral level (M3) with a more pronounced keel. The osteoderms gradually decrease in size as they adopt a rhombus shape towards the caudal region (M4) until the caudal end (M5). The ventral osteoderms, with a lack of keel and peg, exhibit a rectangular shape in the central region (M6), while a square or sub-square shape is observed along its lateral margin (M7). Six specimens are could not be assigned to Indosinosuchus and were described as morphotype 8 (M8). This morphotype has distinct characteristics resembling the ventral osteoderms of Mesoeucrocodylia that are yet to be described. The discovery of Indosinosuchus and Mesoeucrocodylia osteoderms suggests a diverse range of crocodyliformes in the Phu Noi fossil excavation site with potential for future studies.
本研究对泰国加拉信府 Phu Noi 上侏罗纪遗址出土的骨龙进行了形态分析。对 203 个标本收集的数据表明,197 个骨皮属于远程龙科。在这203件标本中,有109件是背侧的骨皮形态(M1至M5),形状为近方形、近矩形或斜方形,有一个突起和一个小齿。另外88个是腹部骨皮形态(M6和M7),呈长方形或近方形,没有明显的突缘或齿状突起。背侧和腹侧的骨膜具有大的近圆形椭圆杯和一个条形的前关节面。形态特征数据可用于描述骨膜的位置。颈椎前部骨膜的特征是有一个不明显的突起,形状近方形(M1)。在躯干的后部,骨膜的宽度增加,呈亚矩形(M2)。这些骨膜在骶骨处缩小(M3),其骨突更加明显。骨膜的尺寸逐渐减小,向尾部(M4)呈菱形,直至尾端(M5)。腹侧的骨膜没有突缘和齿状突起,在中央区域呈长方形(M6),而侧面的骨膜呈正方形或近正方形(M7)。有 6 个标本无法归入 M8 形态型,被描述为 M8 形态型。该形态型具有与中目栉水母腹侧骨膜相似的明显特征,但尚未被描述。Mesoeucrocodylia和M8骨膜的发现表明在富诺化石遗址中还存在几种鳄形目,这可能是未来研究的主题。
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引用次数: 0
A new genus of praeaulacid wasp (Hymenoptera: Evanioidea) from the mid-Cretaceous Kachin amber (Myanmar) 白垩纪中期克钦琥珀中的一个新属(膜翅目:Evanioidea)(缅甸)
IF 1 4区 地球科学 Q3 Earth and Planetary Sciences Pub Date : 2023-01-01 DOI: 10.1016/j.annpal.2023.102599
Corentin Jouault , Simon Rosse-Guillevic

The diversity of praeaulacid wasps in Kachin amber is relatively poorly documented. Here, a new genus and species of praeaulacid wasp are described and illustrated from the mid-Cretaceous Kachin amber as Archeogastrinus kachinensis gen. et sp. nov., and placed in the subfamily Praeaulacinae. This new genus is distinguishable from the other Praeaulacidae because of its particular tibial spurs formula 1-2-2; its forewings with a long and thin pterostigma, a 1-M longer than 1-Rs, a 1-Rs longer than its distance to pterostigma, a crossvein 2r-rs aligned (or nearly) with 2rs-m and originating near the middle of pterostigma, a 2rs-m present, a cell 3rm long, a 1cu-a postfurcal to 1-M, a vein 2-M distinct; its hind wings cu-a reaching Cu distad M + Cu fork; and its metasoma with a first segment petiole-like (i.e., long and thin) and ventrally inflated in its distal half. The position of the genus Mesevania is briefly discussed, as well as the validity of the genus Paleosyncrasis, both described from the mid-Cretaceous Kachin amber.

克钦琥珀中前叶胡蜂的多样性文献记载较少。本文描述了中白垩世克钦琥珀中一个新属和新种,称为Archeogastrinus kachinensis gen. et sp. nov.,并将其归入预胡蜂亚科。这个新属由于其特殊的胫刺公式1-2-2而与其他前肢酸科区分开来;其前翅具一细长的翼柱头,长1 ~ 1米,长1 ~ 2米,距翼柱头1 ~ 2米,与翼柱头近正中平行(或近)的横脉2 ~ 2米,有1 ~ 2米长,室长3米,后径1 ~ 1米,脉2 ~ 2米明显;其后翅Cu -a伸至Cu - distad M + Cu叉;其下半部的瘤具有第一节叶柄状(即长而薄),腹侧膨大。本文简要讨论了Mesevania属的位置,以及古syncrasis属的有效性,这两个属都是在中白垩世克钦琥珀中描述的。
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引用次数: 0
Middle Eocene Ostracods from southeast Fayoum area, Egypt: Systematics, paleobathymetry and paleobiogeography 埃及法尤姆东南部始新世中期介形虫:系统学、古测深学和古生物地理学
IF 1 4区 地球科学 Q3 Earth and Planetary Sciences Pub Date : 2023-01-01 DOI: 10.1016/j.annpal.2023.102598
Sobhi A. Helal , Sherif M. El Baz

This study deals with the Middle Eocene ostracods from southeast Fayoum area, Egypt. To carry out this investigation, three outcrops are examined, Gebel Elwe El Breig, Gebel Sath El Hadid and Gebel Munqar El-Shinnara. This study is based on the Midawara Formation that consists mainly of fossiliferous limestone, marly limestone and clay. The studied ostracod assemblage includes thirty-one species and subspecies, belonging to twenty-three genera and seventeen families. The ostracod group belonging to Cytheroidea Baird, 1850 represents the most abundant group that reaches 81.99% in Munqar El-Shinnara section, 81.88% in Sath El Hadid section, and 61.12% in Elwe El Breig section. The structure of the investigated assemblages (abundance of ostracods, number of taxa, Fisher's alpha, Shannon index, and equitability) indicates deposition in an environment belonging to the neritic zone. Moreover, this study tries to use the multivariate analyses to distinguish the paleobiogeographic provinces in Tethyan region during the Middle Eocene. These analyses are applied on a matrix consists of some nominated Middle Eocene species from 13 countries. The results point to two distinctive provinces, Southern Tethys Province (Algeria, Tunisia, Libya, Egypt and Jordan) and the Northern Tethys Province (England, Germany, Spain, France, Belgium, Ukraine, Hungary and Turkey). The reasonable similarities between these provinces may reflect a possible connection between both sides of the Tethys during the Middle Eocene age.

本文研究了埃及法尤姆东南部地区中始新世介形类。为了进行这项调查,考察了三个露头:Gebel Elwe El Breig、Gebel Sath El Hadid和Gebel Munqar El- shinnara。本研究以Midawara组为研究对象,该组主要由化石灰岩、泥灰岩和粘土组成。所研究的介形类包括17科23属31种和亚种。属Cytheroidea Baird, 1850的介形虫群数量最多,Munqar El- shinnara剖面为81.99%,Sath El Hadid剖面为81.88%,Elwe El Breig剖面为61.12%。所调查组合的结构(介形虫丰度、分类群数量、Fisher’s alpha、Shannon指数和公平性)表明沉积环境属于浅海带。此外,本研究尝试运用多元分析方法来区分特提斯地区中始新世的古生物地理省。这些分析应用于一个由来自13个国家的一些指定的中始新世物种组成的矩阵。结果指向两个不同的省份,南特提斯省(阿尔及利亚、突尼斯、利比亚、埃及和约旦)和北特提斯省(英国、德国、西班牙、法国、比利时、乌克兰、匈牙利和土耳其)。这些省份之间合理的相似性可能反映了始新世中期特提斯河两岸之间可能存在的联系。
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引用次数: 0
A new species of glypheid lobster, Glyphea pisuergae (Crustacea, Glypheoidea), from the Early Jurassic of Palencia, Basque-Cantabrian Basin, Spain 西班牙巴斯克-坎塔布里亚盆地帕伦西亚早侏罗世的一种新的雕纹龙虾,皮纹龙虾(甲壳纲,雕纹总科)
IF 1 4区 地球科学 Q3 Earth and Planetary Sciences Pub Date : 2023-01-01 DOI: 10.1016/j.annpal.2023.102596
Sylvain Charbonnier , Alessandro Garassino , Mikel A. López-Horgue

A new glypheid lobster (Crustacea, Glypheidae), Glyphea pisuergae sp. nov. is described from the Early Jurassic (late Pliensbachian–early Toarcian) of Salinas de Pisuerga, Palencia, Spain. This species represents the second record for the genus in the Early Jurassic.

西班牙Palencia的Salinas de Pisuerga早侏罗世(pliensbachia - Toarcian早期)描述了一种新的形虾(甲壳纲,形虾科),Glyphea pisuergae sp. nov.。该物种代表了早侏罗世该属的第二个记录。
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引用次数: 0
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Annales de Paleontologie
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