Pub Date : 2025-10-01DOI: 10.1016/j.annpal.2025.102889
Théo Delemar, Loïc Villier
A fossil goniasterid (Asteroidea, Echinodermata) from the lower Lutetian glauconitic sands of the Authevernes-Vesly quarry (Eure, France) defines a new species, Autavesniasterwozniaki gen. et sp. nov. This is the first discovery of a nearly complete starfish fossil from the Eocene of the Paris Basin, whereas isolated marginal plates are relatively common in these deposits. This fragmentary material is often identified as “Teichaster” poritoides (Des Moulins, 1832) in the literature and institutional paleontological collections. A review of these isolated marginals reveals that the name “Teichaster” poritoides also includes remains of Autavesniasterwozniaki gen. et sp. nov., Metopaster sp., “Metopaster” besanconi Valette, 1925, and “Pycinaster” grignonensis Valette, 1924. The available material for these species remains insufficient to revise the generic attributions. Although “Teichaster” poritoides is widely reported across Europe, the name should be restricted to a species that appears to be endemic to the Paris Basin in the Eocene.
在Authevernes-Vesly采石场（Eure, France）下Lutetian海绿石砂中发现的一种goniasterid化石（Asteroidea, Echinodermata）定义了一个新物种，Autavesniaster wozniaki gen. et sp. 11 .这是在巴黎盆地始新世首次发现的几乎完整的海星化石，而孤立的边缘板块在这些沉积物中相对常见。在文献和机构古生物收藏中，这种碎片材料通常被认定为“Teichaster”poritoides （Des Moulins, 1832）。对这些孤立的边缘的回顾表明，“Teichaster”poritoides的名称还包括Autavesniaster wozniaki gen. et sp., Metopaster sp.，“Metopaster”besanconi Valette， 1925年和“Pycinaster”grignonensis Valette， 1924年的遗骸。这些物种的现有资料仍不足以修正属属归属。尽管“Teichaster”poritoides在欧洲被广泛报道，但这个名字应该仅限于始新世巴黎盆地特有的一种物种。
{"title":"Autaveniaster wozniaki gen. et sp. nov. a new fossil goniasterid (Asteroidea, Echinodermata), and insight on the hidden sea star diversity in the Lutetian of the Paris Basin (France)","authors":"Théo Delemar, Loïc Villier","doi":"10.1016/j.annpal.2025.102889","DOIUrl":"10.1016/j.annpal.2025.102889","url":null,"abstract":"<div><div>A fossil goniasterid (Asteroidea, Echinodermata) from the lower Lutetian glauconitic sands of the Authevernes-Vesly quarry (Eure, France) defines a new species, <em>Autavesniaster</em> <em>wozniaki</em> gen. et sp. nov. This is the first discovery of a nearly complete starfish fossil from the Eocene of the Paris Basin, whereas isolated marginal plates are relatively common in these deposits. This fragmentary material is often identified as “<em>Teichaster</em>” <em>poritoides</em> (Des Moulins, 1832) in the literature and institutional paleontological collections. A review of these isolated marginals reveals that the name “<em>Teichaster</em>” <em>poritoides</em> also includes remains of <em>Autavesniaster</em> <em>wozniaki</em> gen. et sp. nov., <em>Metopaster</em> sp., “<em>Metopaster</em>” <em>besanconi</em> Valette, 1925, and “<em>Pycinaster</em>” <em>grignonensis</em> Valette, 1924. The available material for these species remains insufficient to revise the generic attributions. Although “<em>Teichaster</em>” <em>poritoides</em> is widely reported across Europe, the name should be restricted to a species that appears to be endemic to the Paris Basin in the Eocene.</div></div>","PeriodicalId":50780,"journal":{"name":"Annales de Paleontologie","volume":"111 4","pages":"Article 102889"},"PeriodicalIF":0.7,"publicationDate":"2025-10-01","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"145486406","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":4,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
Pub Date : 2025-10-01DOI: 10.1016/j.annpal.2025.102903
Olev Vinn , Oive Tinn , Liisa Lang , Mare Isakar , Magdy El Hedeny , Abdullah A. Alkahtane , Saleh Alfarraj
The Middle Ordovician cryptic encrusting community was numerically dominated by bryozoans (66.7%), followed by cornulitids (26.7%) and graptolites (6.6%). Bryozoans also occupied the largest encrustation area. Similarly, the Late Ordovician cryptic encrusting community was dominated by bryozoans (60%), while cornulitids (30%), brachiopods (8%), and graptolites (2%) formed a minor part of the community. In the studied samples, the disparity of Upper Ordovician cryptic encrusting fauna (including Bryozoa, Brachiopoda, Tentaculita, and Hemichordata) was greater than that of the Middle Ordovician, which consisted of Bryozoa, Hemichordata, and Tentaculita. There appears to be an increase in the maximum encrustation density within nautiloid shell interiors from the Darriwilian to the Katian. Additionally, a statistically significant increase in the percentage of encrusted nautiloid interiors from the Darriwilian to the Katian suggests a growing prevalence of cryptic encrusters over time. The increase was likely driven more by the evolutionary diversification of encrusting taxa than by the warming climate during the Katian in Baltica.
{"title":"Cryptic fauna in nautiloid shells from the Ordovician of Estonia","authors":"Olev Vinn , Oive Tinn , Liisa Lang , Mare Isakar , Magdy El Hedeny , Abdullah A. Alkahtane , Saleh Alfarraj","doi":"10.1016/j.annpal.2025.102903","DOIUrl":"10.1016/j.annpal.2025.102903","url":null,"abstract":"<div><div>The Middle Ordovician cryptic encrusting community was numerically dominated by bryozoans (66.7%), followed by cornulitids (26.7%) and graptolites (6.6%). Bryozoans also occupied the largest encrustation area. Similarly, the Late Ordovician cryptic encrusting community was dominated by bryozoans (60%), while cornulitids (30%), brachiopods (8%), and graptolites (2%) formed a minor part of the community. In the studied samples, the disparity of Upper Ordovician cryptic encrusting fauna (including Bryozoa, Brachiopoda, Tentaculita, and Hemichordata) was greater than that of the Middle Ordovician, which consisted of Bryozoa, Hemichordata, and Tentaculita. There appears to be an increase in the maximum encrustation density within nautiloid shell interiors from the Darriwilian to the Katian. Additionally, a statistically significant increase in the percentage of encrusted nautiloid interiors from the Darriwilian to the Katian suggests a growing prevalence of cryptic encrusters over time. The increase was likely driven more by the evolutionary diversification of encrusting taxa than by the warming climate during the Katian in Baltica.</div></div>","PeriodicalId":50780,"journal":{"name":"Annales de Paleontologie","volume":"111 4","pages":"Article 102903"},"PeriodicalIF":0.7,"publicationDate":"2025-10-01","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"145486408","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":4,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
Pub Date : 2025-10-01DOI: 10.1016/j.annpal.2025.102901
France de Lapparent de Broin , Xabier Murelaga , Renaud Vacant
The turtle Marocokatognathus jimenezi n. g., n. sp., known by a dentary, unique by its long, narrow and upward curved beak, is defined as Chelonioidea, not Dermochelyoidea. The morphology of the dentary indicates its skull had a secondary palate. By comparison with Chelonioid turtles, the variations of that structure are classified here in morphotypes, of practical taxonomical use and partly of phyletic incidence, notably as a Chelonioidea member. The new fossil is attributed to Cheloniidae and related to extant forms by: - its general dentary morphology including a Carettine shape of the bony triturating surface and the same relative proportions of distances between structural elements (posterior border of the symphysis, coronoid process, foramen mentale, in relation to corresponding skull structures); - the assignation to a same morphotype 1 of secondary palate, by an elongated snout for an elongated lower jaw, the choanal border anterior to palatine fossae, palate not developed beyond ca. two thirds of the snout length, correlatively preserving long dentary branches anteriorly and posteriorly to the coronoid process; - and overall by the posterior implantation of the mandibular ramphotheca on the mandibular symphysis with a medial symphyseal facet, specific of extant Cheloniidae. The possible diet and prey capture processes are examined, proposing a seizure of various prey of shape adapted to the narrow and long beak, followed immediately by their aspiration and swallowing. By its novelty, the new form emphasizes the great diversity of fossil turtles from Phosphates of Morocco, which are inventoried, as of the whole world.
Marocokatognathus jimenezi n. g., n. sp.，以其长而窄且向上弯曲的喙而闻名，被定义为龟类，而不是皮龟类。牙齿的形态表明它的头骨有一个次级上颚。通过与类龟的比较，该结构的变化在形态类型、实际分类学用途和部分种系发生率方面进行了分类，特别是作为一个类龟纲成员。这一新化石被归为龟科，并与现存的形式相关：-其一般的牙齿形态包括骨磨砂表面的Carettine形状和结构元素之间距离的相对比例相同（联合后缘，冠状突，颏孔，相对于相应的头骨结构）；-下颌长鼻部与次上颚相同的形态型1，下颌前侧颊缘长鼻部，上颚发育不超过鼻部长度的约三分之二，相对保留冠突前后的长牙枝；-总的来说，通过在下颌联合和内侧联合关节突上的下颌下颌骨膜的后侧植入，这是现存的chelonidae特有的。研究了可能的饮食和猎物捕获过程，提出了各种形状适应窄而长的喙的猎物的捕获，紧接着是它们的吸气和吞咽。由于其新新性，新形式强调了摩洛哥磷酸盐化石龟的巨大多样性，这些化石龟在全世界都有记录。
{"title":"A chelonioid turtle with a long, upward curved, lower jaw beak, phosphates of Morocco, Paleocene-Eocene transition","authors":"France de Lapparent de Broin , Xabier Murelaga , Renaud Vacant","doi":"10.1016/j.annpal.2025.102901","DOIUrl":"10.1016/j.annpal.2025.102901","url":null,"abstract":"<div><div>The turtle <em>Marocokatognathus jimenezi</em> n. g., n. sp., known by a dentary, unique by its long, narrow and upward curved beak, is defined as Chelonioidea, not Dermochelyoidea. The morphology of the dentary indicates its skull had a secondary palate. By comparison with Chelonioid turtles, the variations of that structure are classified here in morphotypes, of practical taxonomical use and partly of phyletic incidence, notably as a Chelonioidea member. The new fossil is attributed to Cheloniidae and related to extant forms by: - its general dentary morphology including a Carettine shape of the bony triturating surface and the same relative proportions of distances between structural elements (posterior border of the symphysis, coronoid process, foramen mentale, in relation to corresponding skull structures); - the assignation to a same morphotype 1 of secondary palate, by an elongated snout for an elongated lower jaw, the choanal border anterior to palatine fossae, palate not developed beyond ca. two thirds of the snout length, correlatively preserving long dentary branches anteriorly and posteriorly to the coronoid process; - and overall by the posterior implantation of the mandibular ramphotheca on the mandibular symphysis with a medial symphyseal facet, specific of extant Cheloniidae. The possible diet and prey capture processes are examined, proposing a seizure of various prey of shape adapted to the narrow and long beak, followed immediately by their aspiration and swallowing. By its novelty, the new form emphasizes the great diversity of fossil turtles from Phosphates of Morocco, which are inventoried, as of the whole world.</div></div>","PeriodicalId":50780,"journal":{"name":"Annales de Paleontologie","volume":"111 4","pages":"Article 102901"},"PeriodicalIF":0.7,"publicationDate":"2025-10-01","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"145617999","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":4,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
Pub Date : 2025-10-01DOI: 10.1016/j.annpal.2025.102902
Zhi-Teng Chen , Xuhongyi Zheng
The mayfly family Isonychiidae is represented by the single extant genus Isonychia Eaton, 1871 and has long lacked a reliable fossil record, limiting insights into its evolutionary history. Here, we describe Fujiwaranychia chiyokoae gen. et sp. nov., a female imago preserved in mid-Cretaceous Kachin amber from northern Myanmar. This specimen represents the first reliable fossil evidence for Isonychiidae and confirms the family's presence in Laurasia during the Cretaceous. The new genus shares several morphological features with extant Isonychia, including forewing venation and a similar tarsal formula, but differs in the absence of cubital cross veins in forewings and the presence of an unforked media posterior vein in the hind wing. These differences justify its placement in a separate, extinct genus within Isonychiidae. The discovery extends the known history of the family by nearly 100 million years and reveals a previously unrecognized degree of morphological diversity. The fossil provides a critical calibration point for future phylogenetic studies and underscores the paleontological value of amber inclusions in tracing the evolutionary history of mayflies.
蜉蝣科异爪蝇科的代表是1871年仅存的异爪蝇属伊顿，长期以来缺乏可靠的化石记录，限制了对其进化史的了解。在这里，我们描述了Fujiwaranychia chiyokoae gen. et sp. nov.，一种保存在缅甸北部克钦琥珀中白垩纪中期的女性形象。这个标本代表了异爪龙科的第一个可靠的化石证据，并证实了该家族在白垩纪时期在劳亚岛的存在。新属与现存的异爪目有几个相同的形态特征，包括前翼脉和类似的跗骨公式，但在前翼没有肘交叉脉和后翼有未分叉的中位后脉方面有所不同。这些差异证明了它属于异爪目中一个单独的、已灭绝的属。这一发现将该家族的已知历史延长了近1亿年，并揭示了此前未被认识到的形态多样性程度。该化石为未来的系统发育研究提供了一个关键的定标点，并强调了琥珀包裹体在追踪蜉蝣进化史中的古生物学价值。
{"title":"First reliable fossil record of the family Isonychiidae (Insecta, Ephemeroptera) from Cretaceous Kachin amber","authors":"Zhi-Teng Chen , Xuhongyi Zheng","doi":"10.1016/j.annpal.2025.102902","DOIUrl":"10.1016/j.annpal.2025.102902","url":null,"abstract":"<div><div>The mayfly family Isonychiidae is represented by the single extant genus <em>Isonychia</em> Eaton, 1871 and has long lacked a reliable fossil record, limiting insights into its evolutionary history. Here, we <em>describe Fujiwaranychia chiyokoae</em> gen. et sp. nov., a female imago preserved in mid-Cretaceous Kachin amber from northern Myanmar. This specimen represents the first reliable fossil evidence for Isonychiidae and confirms the family's presence in Laurasia during the Cretaceous. The new genus shares several morphological features with extant <em>Isonychia</em>, including forewing venation and a similar tarsal formula, but differs in the absence of cubital cross veins in forewings and the presence of an unforked media posterior vein in the hind wing. These differences justify its placement in a separate, extinct genus within Isonychiidae. The discovery extends the known history of the family by nearly 100 million years and reveals a previously unrecognized degree of morphological diversity. The fossil provides a critical calibration point for future phylogenetic studies and underscores the paleontological value of amber inclusions in tracing the evolutionary history of mayflies.</div></div>","PeriodicalId":50780,"journal":{"name":"Annales de Paleontologie","volume":"111 4","pages":"Article 102902"},"PeriodicalIF":0.7,"publicationDate":"2025-10-01","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"145486407","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":4,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
Pub Date : 2025-07-01DOI: 10.1016/j.annpal.2025.102888
Urlain Gaël Yakouya-Moubamba , Thomas Saucède , Wilma Linda-Belle Mougola , Benjamin Musavu Moussavou
This study presents a detailed systematic analysis of the irregular echinoid Mecaster fourneli (Deshayes, 1847) from Turonian (Late Cretaceous) deposits of the Gabonese coastal basin. The morphological and biometric study of 26 well-preserved specimens has allowed a precise characterisation of this species, previously mentioned but never described or illustrated in this region. The specimens studied come from three different localities (Gabon Store, Pétro Gabon and Maison Georges Rawiri). Comparative morpho-biometric analysis with populations from Algeria and Peru reveals a remarkable morphological homogeneity despite the geographical distance. The presence of M. fourneli in the Gabonese coastal basin confirms the geographical expansion of this species during the Upper Cretaceous and strengthens the hypothesis of a palaeobiogeographical link between the Mediterranean and West African provinces during the Turonian-Coniacian, probably via a trans-Saharan passage during the major transgressive event of this period.
本文对加蓬海岸盆地晚白垩世Turonian（晚白垩世）沉积中的不规则针孔类Mecaster fourneli （Deshayes, 1847）进行了详细的系统分析。对26个保存完好的标本进行形态学和生物计量学研究，可以对该物种进行精确的表征，该物种以前曾提到过，但从未在该地区描述或说明过。研究的标本来自三个不同的地方（加蓬商店、p加蓬和Maison Georges Rawiri）。阿尔及利亚和秘鲁种群的形态-生物统计学比较分析显示，尽管地理距离较远，但形态同质性显著。M. fourneli在加蓬海岸盆地的存在证实了该物种在上白垩纪的地理扩张，并加强了在Turonian-Coniacian期间地中海和西非省之间存在古生物地理联系的假设，可能是通过该时期主要海侵事件中的跨撒哈拉通道。
{"title":"Mecaster fourneli (Deshayes, 1847) (Echinodermata: Echinoidea) from the Turonian (Late Cretaceous) of the Gabonese Coastal Basin: systematics and palaeobiogeographical implications","authors":"Urlain Gaël Yakouya-Moubamba , Thomas Saucède , Wilma Linda-Belle Mougola , Benjamin Musavu Moussavou","doi":"10.1016/j.annpal.2025.102888","DOIUrl":"10.1016/j.annpal.2025.102888","url":null,"abstract":"<div><div>This study presents a detailed systematic analysis of the irregular echinoid <em>Mecaster fourneli</em> (Deshayes, 1847) from Turonian (Late Cretaceous) deposits of the Gabonese coastal basin. The morphological and biometric study of 26 well-preserved specimens has allowed a precise characterisation of this species, previously mentioned but never described or illustrated in this region. The specimens studied come from three different localities (Gabon Store, Pétro Gabon and Maison Georges Rawiri). Comparative morpho-biometric analysis with populations from Algeria and Peru reveals a remarkable morphological homogeneity despite the geographical distance. The presence of <em>M</em>. <em>fourneli</em> in the Gabonese coastal basin confirms the geographical expansion of this species during the Upper Cretaceous and strengthens the hypothesis of a palaeobiogeographical link between the Mediterranean and West African provinces during the Turonian-Coniacian, probably via a trans-Saharan passage during the major transgressive event of this period.</div></div>","PeriodicalId":50780,"journal":{"name":"Annales de Paleontologie","volume":"111 3","pages":"Article 102888"},"PeriodicalIF":0.7,"publicationDate":"2025-07-01","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"145020963","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":4,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
Pub Date : 2025-07-01DOI: 10.1016/j.annpal.2025.102867
Sylvain Charbonnier , Alessandro Garassino , Thomas Laville , Damien Gendry , Philippe Courville , Jean-Philippe Pezy , Julien Devillez
Decapod crustaceans (Crustacea, Decapoda) are particularly abundant and diverse in the Jurassic of France. Based on historical collections from French museums and universities, about 25 species belonging to six infraorders (Axiidea, Anomura, Brachyura, Glypheidea, Erymida, Achelata) are reported from the Sinemurian to the Kimmeridgian, among which three are new: Palaeastacus ruteni sp. nov. (Erymidae, Bathonian), Stenodactylina souconnae sp. nov. (Erymidae, Bajocian), and Linuparus petrocorii sp. nov. (Palinuridae, Kimmeridgian). These new occurrences enlarge the stratigraphic and palaeogeographic distributions of most of the studied species in all the main French sedimentary platforms and basins.
{"title":"New records of decapod crustaceans in the Jurassic of France","authors":"Sylvain Charbonnier , Alessandro Garassino , Thomas Laville , Damien Gendry , Philippe Courville , Jean-Philippe Pezy , Julien Devillez","doi":"10.1016/j.annpal.2025.102867","DOIUrl":"10.1016/j.annpal.2025.102867","url":null,"abstract":"<div><div>Decapod crustaceans (Crustacea, Decapoda) are particularly abundant and diverse in the Jurassic of France. Based on historical collections from French museums and universities, about 25 species belonging to six infraorders (Axiidea, Anomura, Brachyura, Glypheidea, Erymida, Achelata) are reported from the Sinemurian to the Kimmeridgian, among which three are new: <em>Palaeastacus ruteni</em> sp. nov. (Erymidae, Bathonian), <em>Stenodactylina souconnae</em> sp. nov. (Erymidae, Bajocian), and <em>Linuparus petrocorii</em> sp. nov. (Palinuridae, Kimmeridgian). These new occurrences enlarge the stratigraphic and palaeogeographic distributions of most of the studied species in all the main French sedimentary platforms and basins.</div></div>","PeriodicalId":50780,"journal":{"name":"Annales de Paleontologie","volume":"111 3","pages":"Article 102867"},"PeriodicalIF":0.7,"publicationDate":"2025-07-01","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"144997694","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":4,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
One hermit crab and two brachyuran crabs from the Eocene of Pakistan are described. The hermit crab, recovered from the middle Eocene of the Drazinda Formation in the Sulaiman Range (northern Pakistan), represents a new species of diogenid (Diogenidae): ?Diogenes jafrii. The two brachyuran crabs, recovered from two different localities of the Kirthar Range (southern Pakistan), have been identified as a portunid (Portunidae) and a possibly dromioid (Dromioidea), respectively, since the fragmentary condition does not allow a more specific systematic assignment. In addition, a specimen from the late Paleocene of the southern Kirthar Range, figured by Charbonnier et al. (2013), is re-examined herein.
描述了巴基斯坦始新世的一种寄居蟹和两种短爪蟹。寄居蟹，发现于巴基斯坦北部苏莱曼山脉德拉津达组始新世中期，是菊科寄居蟹新种：？提奥奇尼斯jafrii。这两种短爪蟹是在巴基斯坦南部基尔塔山脉的两个不同地点发现的，由于碎片状况不允许更具体的系统分配，它们被分别鉴定为一种梭子蟹科（梭子蟹科）和一种可能的梭子蟹科（梭子蟹总科）。此外，本文还对Charbonnier et al.（2013）绘制的南Kirthar Range晚古新世标本进行了重新检验。
{"title":"New data on Paleogene decapod crustaceans from Pakistan","authors":"Àlex Ossó , Sylvain Charbonnier , Alessandro Garassino , Didier Merle , Rafique Ahmed Lashari , Annachiara Bartolini , Grégoire Métais","doi":"10.1016/j.annpal.2025.102887","DOIUrl":"10.1016/j.annpal.2025.102887","url":null,"abstract":"<div><div>One hermit crab and two brachyuran crabs from the Eocene of Pakistan are described. The hermit crab, recovered from the middle Eocene of the Drazinda Formation in the Sulaiman Range (northern Pakistan), represents a new species of diogenid (Diogenidae): ?<em>Diogenes jafrii</em>. The two brachyuran crabs, recovered from two different localities of the Kirthar Range (southern Pakistan), have been identified as a portunid (Portunidae) and a possibly dromioid (Dromioidea), respectively, since the fragmentary condition does not allow a more specific systematic assignment. In addition, a specimen from the late Paleocene of the southern Kirthar Range, figured by Charbonnier et al. (2013), is re-examined herein.</div></div>","PeriodicalId":50780,"journal":{"name":"Annales de Paleontologie","volume":"111 3","pages":"Article 102887"},"PeriodicalIF":0.7,"publicationDate":"2025-07-01","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"144997695","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":4,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
Pub Date : 2025-07-01DOI: 10.1016/j.annpal.2025.102869
Olev Vinn , Andrej Ernst , Oive Tinn , Liisa Lang , Mare Isakar , Magdy El Hedeny , Mansour I. Almansour , Saleh Alfarraj
Six new symbiotic associations between erect branching bryozoan colonies and tentaculitoid tubeworm-like organisms have been described from argillaceous carbonate rocks of the Ludfordian of Saaremaa Island, Estonia. Several symbiotic associations occur in the Kuressaare Formation: Conchicolites sp.– Fistulipora sp. A and Fistulipora sp. B, Palaeoconchus sp. – Fistulipora sp. A, Conchicolites sp. – ?Eridotrypella sp., Conchicolites sp.– ?Anisotrypa proavus, Conchicolites sp. –?Leptotrypella versimilis and Monotrypa sp. – unknown endozoobiont association. The colonization of bryozoans by cornulitids and microconchids likely occurred because the former provided a suitable hard substrate on an otherwise soft clay sea floor. Most cornulitids had endobiotic life modes and were completely intergrown with their host bryozoans. There is no evidence of how cornulitids might have provided some advantage to the bryozoan host. However, given the likelihood of feeding competition, these associations are more appropriately qualified as mildly parasitic.
{"title":"Symbiosis in late Silurian (Ludfordian) bryozoans from Saaremaa Island, Estonia","authors":"Olev Vinn , Andrej Ernst , Oive Tinn , Liisa Lang , Mare Isakar , Magdy El Hedeny , Mansour I. Almansour , Saleh Alfarraj","doi":"10.1016/j.annpal.2025.102869","DOIUrl":"10.1016/j.annpal.2025.102869","url":null,"abstract":"<div><div>Six new symbiotic associations between erect branching bryozoan colonies and tentaculitoid tubeworm-like organisms have been described from argillaceous carbonate rocks of the Ludfordian of Saaremaa Island, Estonia. Several symbiotic associations occur in the Kuressaare Formation: <em>Conchicolites</em> sp.– <em>Fistuli</em><em>pora</em> sp. A and <em>Fistulipora</em> sp. B, <em>Palaeoconchus</em> sp. – <em>Fistulipora</em> sp. A, <em>Conchicolites</em> sp. – ?<em>Eridotrypella</em> sp., <em>Conchicolites</em> sp.– ?<em>Anisotrypa proavus</em>, <em>Conchicolites</em> sp. –?<em>Leptotrypella versimilis</em> and <em>Monotrypa</em> sp. – unknown endozoobiont association. The colonization of bryozoans by cornulitids and microconchids likely occurred because the former provided a suitable hard substrate on an otherwise soft clay sea floor. Most cornulitids had endobiotic life modes and were completely intergrown with their host bryozoans. There is no evidence of how cornulitids might have provided some advantage to the bryozoan host. However, given the likelihood of feeding competition, these associations are more appropriately qualified as mildly parasitic.</div></div>","PeriodicalId":50780,"journal":{"name":"Annales de Paleontologie","volume":"111 3","pages":"Article 102869"},"PeriodicalIF":0.7,"publicationDate":"2025-07-01","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"144997627","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":4,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
Pub Date : 2025-07-01DOI: 10.1016/j.annpal.2025.102872
Kalangi Rodrigo , Wijerathne Bohingamuwa
The evolutionary history of lions (Panthera leo) reveals a dynamic trajectory shaped by environmental shifts, geographic dispersal, and adaptations to prey availability. Fossil evidence, spanning over a million years, underscores their origins, range expansions, and eventual divergence into various subspecies, including modern lions. This study provides a comparative analysis of lion species, with a focus on the extinct Panthera leo sinhaleyus, through an examination of dental morphological traits — specifically crown height and crown breadth. Panthera leo sinhaleyus, discovered in 1938 at Kuruvita, in the Rathnapura district, Sri Lanka, is known from the holotype specimen (i.e., a left lower carnassial). This specimen was found in association with extinct fauna assigned to the Late Pleistocene. Since its discovery, there has been limited research addressing its chronological and taxonomic range in relation to contemporary lion species, from around Asia, Africa and Europe. Our statistical analyses reveal that Panthera leo sinhaleyus exhibits distinct intermediate dental characteristics. Its crown height (29 mm) surpasses that of the modern Panthera leo (26.9 mm on average) but remains smaller than that of the Middle Pleistocene lion Panthera spelaea (32.36 mm). Similarly, its crown breadth (14.5 mm) is broader than many subspecies of Panthera leo yet narrower than Panthera spelaea (18.19 mm). These findings suggest that Panthera leo sinhaleyus exhibits adaptations potentially reflective of its environment in Pleistocene Sri Lanka, indicating a distinct ecological niche and evolutionary trajectory. This study underscores the need for further investigation into the chronological and taxonomic placement of Panthera leo sinhaleyus, contributing to a broader understanding of morphological diversity and evolutionary pathways among prehistoric lion species.
狮子（Panthera leo）的进化史揭示了一个由环境变化、地理分散和对猎物的适应所塑造的动态轨迹。跨越一百万年的化石证据强调了它们的起源、活动范围的扩展，以及最终分化成各种亚种，包括现代狮子。本研究提供了狮子物种的比较分析，重点是已灭绝的Panthera leo sinhaleyus，通过检查牙齿形态特征-特别是冠高和冠宽。Panthera leo sinhaleyus于1938年在斯里兰卡Rathnapura地区的Kuruvita被发现，它是通过完整模式标本（即左下卡尼塞尔）而闻名的。这个标本是与属于晚更新世的灭绝动物群一起发现的。自从它被发现以来，关于它与亚洲、非洲和欧洲当代狮子物种的时间和分类范围的研究一直很有限。我们的统计分析显示，黑豹具有明显的中间牙齿特征。其冠高（29 mm）超过了现代美洲豹（Panthera leo）的平均冠高（26.9 mm），但仍低于中更新世美洲豹（Panthera spelaea）的冠高（32.36 mm）。同样，它的冠宽（14.5 mm）比黑豹（Panthera leo）的许多亚种宽，但比黑豹（Panthera spelaea）的冠宽（18.19 mm）窄。这些发现表明，斯里兰卡更新世Panthera leo sinhaleyus表现出可能反映其环境的适应性，表明其具有独特的生态位和进化轨迹。本研究强调了进一步研究史前狮子的年代和分类位置的必要性，有助于更广泛地了解史前狮子物种的形态多样性和进化途径。
{"title":"Revisiting Panthera leo sinhaleyus: Morphological insights and evolutionary implications from the holotype specimen from Sri Lanka","authors":"Kalangi Rodrigo , Wijerathne Bohingamuwa","doi":"10.1016/j.annpal.2025.102872","DOIUrl":"10.1016/j.annpal.2025.102872","url":null,"abstract":"<div><div>The evolutionary history of lions (<em>Panthera leo</em>) reveals a dynamic trajectory shaped by environmental shifts, geographic dispersal, and adaptations to prey availability. Fossil evidence, spanning over a million years, underscores their origins, range expansions, and eventual divergence into various subspecies, including modern lions. This study provides a comparative analysis of lion species, with a focus on the extinct <em>Panthera leo sinhaleyus</em>, through an examination of dental morphological traits — specifically crown height and crown breadth. <em>Panthera leo sinhaleyus</em>, discovered in 1938 at Kuruvita, in the Rathnapura district, Sri Lanka, is known from the holotype specimen (i.e., a left lower carnassial). This specimen was found in association with extinct fauna assigned to the Late Pleistocene. Since its discovery, there has been limited research addressing its chronological and taxonomic range in relation to contemporary lion species, from around Asia, Africa and Europe. Our statistical analyses reveal that <em>Panthera leo sinhaleyus</em> exhibits distinct intermediate dental characteristics. Its crown height (29<!--> <!-->mm) surpasses that of the modern <em>Panthera leo</em> (26.9<!--> <!-->mm on average) but remains smaller than that of the Middle Pleistocene lion <em>Panthera spelaea</em> (32.36<!--> <!-->mm). Similarly, its crown breadth (14.5<!--> <!-->mm) is broader than many subspecies of <em>Panthera leo</em> yet narrower than <em>Panthera spelaea</em> (18.19<!--> <!-->mm). These findings suggest that <em>Panthera leo sinhaleyus</em> exhibits adaptations potentially reflective of its environment in Pleistocene Sri Lanka, indicating a distinct ecological niche and evolutionary trajectory. This study underscores the need for further investigation into the chronological and taxonomic placement of <em>Panthera leo sinhaleyus</em>, contributing to a broader understanding of morphological diversity and evolutionary pathways among prehistoric lion species.</div></div>","PeriodicalId":50780,"journal":{"name":"Annales de Paleontologie","volume":"111 3","pages":"Article 102872"},"PeriodicalIF":0.7,"publicationDate":"2025-07-01","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"144997696","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":4,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
Pub Date : 2025-04-01DOI: 10.1016/j.annpal.2025.102757
Mohamed Benzaggagh
In the External Rif, northwest Morocco, the Barremian Stage is represented by a thick (200 m) marl and marly limestone series. This series, particularly in the western part of the External Rif, is locally rich in pyritic ammonites. Four Barremian outcrops (Malha, Jebel Meguedrouz, Jebel Bourhzine, and Mrejet), located between Mjara and Mrejet have been studied. Collected ammonites (931 specimens) from the four outcrops allow the identification of 38 species, characterizing the Hauterivian and the lowermost Barremian, at the Mrejet section; the uppermost lower Barremian, at Mrejet and Bourhzine sections, and the upper Barremian, at Mrejet and Malha sections. These species are described and illustrated for the first time from the External Rif. Most of them are reported for the first time from the studied area. They are common to several Lower Cretaceous basins of the southern and northern margins of the Tethys, mainly NE Algeria, SE Spain, SE France, Italy, eastern Serbia, Hungary, Slovakia, and Romania. Stratigraphic and paleogeographic distributions of each species are discussed.
{"title":"Barremian pyritic ammonites from the western External Rif (northwest Morocco)","authors":"Mohamed Benzaggagh","doi":"10.1016/j.annpal.2025.102757","DOIUrl":"10.1016/j.annpal.2025.102757","url":null,"abstract":"<div><div>In the External Rif, northwest Morocco, the Barremian Stage is represented by a thick (200<!--> <!-->m) marl and marly limestone series. This series, particularly in the western part of the External Rif, is locally rich in pyritic ammonites. Four Barremian outcrops (Malha, Jebel Meguedrouz, Jebel Bourhzine, and Mrejet), located between Mjara and Mrejet have been studied. Collected ammonites (931 specimens) from the four outcrops allow the identification of 38 species, characterizing the Hauterivian and the lowermost Barremian, at the Mrejet section; the uppermost lower Barremian, at Mrejet and Bourhzine sections, and the upper Barremian, at Mrejet and Malha sections. These species are described and illustrated for the first time from the External Rif. Most of them are reported for the first time from the studied area. They are common to several Lower Cretaceous basins of the southern and northern margins of the Tethys, mainly NE Algeria, SE Spain, SE France, Italy, eastern Serbia, Hungary, Slovakia, and Romania. Stratigraphic and paleogeographic distributions of each species are discussed.</div></div>","PeriodicalId":50780,"journal":{"name":"Annales de Paleontologie","volume":"111 2","pages":"Article 102757"},"PeriodicalIF":1.0,"publicationDate":"2025-04-01","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"143759974","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":4,"RegionCategory":"地球科学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
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