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Latest Oligocene to earliest Pliocene deep-sea benthic foraminifera from ODP Sites 752, 1168 and 1139, southern Indian Ocean 南印度洋ODP遗址752、1168和1139的渐新世至上新世早期深海底栖有孔虫
IF 2 3区 地球科学 Q1 PALEONTOLOGY Pub Date : 2019-12-10 DOI: 10.25500/EDATA.BHAM.00000411
Dana Ridha, I. Boomer, K. Edgar
Count data for most abundant benthic foraminifera in the late Oligocene-Neogene in ODP Holes 752A, 1139A and 1168A
ODP 752A、1139A和1168A孔晚渐新世—新近纪底栖有孔虫最丰富的计数资料
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引用次数: 2
Benthic foraminifera indicate Glacial North Pacific Intermediate Water and reduced primary productivity over Bowers Ridge, Bering Sea, since the Mid-Brunhes Transition Benthic有孔虫表明,自Mid-Brunhes过渡以来,白令海Bowers山脊上的北太平洋中层水和初级生产力降低
IF 2 3区 地球科学 Q1 PALEONTOLOGY Pub Date : 2019-12-03 DOI: 10.5194/jm-38-177-2019
S. Kender, A. Aturamu, J. Zalasiewicz, M. Kaminski, M. Williams
Abstract. The Mid-Brunhes Transition (MBT) saw an increase in the amplitude of glacial cycles expressed in ice core and deep ocean records from about 400 ka, but its influence on high-latitude climates is not fully understood. The Arctic Ocean is thought to have warmed and exhibited reduced sea ice, but little is known of sea ice marginal locations such as the Bering Sea. The Bering Sea is the link between the Arctic and Pacific Ocean and is an area of high productivity and CO2 ventilation; it hosts a pronounced oxygen minimum zone (OMZ) and is thought to be the location of Glacial North Pacific Intermediate Water (GNPIW) formation in the Pleistocene. To understand palaeoceanographic change in the region, we analysed benthic foraminiferal faunas from Bowers Ridge (Site U1342, 800 m of water depth) over the past 600 kyr, as they are uniquely well preserved and sensitive to changes in deep and surface ocean conditions. We identified and imaged 71 taxa and provide a full taxonomy. Foraminiferal preservation is markedly higher during glacials, indicating the presence of less corrosive GNPIW. The most abundant species are Bulimina exilis, Takayanagia delicata, Alabaminella weddellensis, Gyroidina sp. 2, Cassidulina laevigata, Islandiella norcrossi, and Uvigerina bifurcata, consistent with broadly high net primary production throughout the last 600 kyr. Correspondence analysis shows that the most significant Assemblage 1 comprises B. exilis, T. delicata, Bolivina spissa, and Brizalina, which occur sporadically within intervals of laminated, biogenic-rich sediment, mostly during glacials and also some deglacials, and are interpreted as indicating very high productivity. Other assemblages contain the phytodetritivore species A. weddellensis, I. norcrossi, and C. laevigata, indicative of seasonal phytoplankton blooms. Before the MBT, more numerous intervals of the very high-productivity Assemblage 1 and A. weddellensis occur, which we suggest reflect a time of more sea-ice-related seasonal stratification and ice edge blooms. Our inference of a decrease in sea ice meltwater stratification influence in the central Bering Sea after the MBT is consistent with records showing that the Arctic and Pacific Ocean warmed during glacials and suggests that high-latitude productivity and sea ice changes were an important feature of this climate event.
摘要在中布鲁内斯过渡期(MBT),冰芯和深海记录中表达的冰川周期的振幅从大约400增加 ka,但其对高纬度气候的影响并不完全。北冰洋被认为已经变暖,海冰减少,但对白令海等海冰边缘地区知之甚少。白令海是连接北冰洋和太平洋的纽带,是一个生产力高、二氧化碳排放量大的地区;没有明显的氧最低带(OMZ),被认为是更新世北太平洋冰川中间水(GNPIW)形成的位置。为了了解该地区的古海洋地理变化,分析了鲍尔斯山脊(U1342800)的底栖有孔虫动物群 mof水深) kyr,因为它们保存完好,对深海和表层海洋条件的变化非常敏感。我们鉴定和分析了71个分类群,并提供了一个完整的分类学。在冰川期,有孔虫的保存率明显较高,表明存在腐蚀性较小的GNPIW。最丰富的物种是出口Bulimina exilis、美味Takayanagia delicata、韦德尔Alabaminella weddellensis、Gyroidina sp.2、金钱子Cassidulina laevigata、北十字Islandiella和分叉Uvigerina divoca,在过去600年中与大致较高的净初级产量一致 kyr。对应分析表明,最显著的组合1包括B。exilis、T.delicata、Bolivina spissa和Brizalina,它们偶尔出现在富含生物成因的层状沉积物的间隔内,主要发生在冰川期和一些退冰川期,被解释为表明生产力非常高。其他组合包括植物碎屑动物物种A.weddellensis、I.norcrossi和C.laevigata,这表明季节性的hytopankton开花。在MBT之前,出现了更多的超高产组合1和A.weddellensis的间隔,我们认为这反映了与海冰相关的季节性分层和冰缘水华增多的时期。我们对MBT后白令海中部海冰融水分层影响减少的推断与北极和太平洋在冰川作用期间变暖的记录一致,并表明高纬度生产力和海冰变化是这一气候事件的一个重要特征。
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引用次数: 7
Early Oligocene dinocysts as a tool for palaeoenvironment reconstruction and stratigraphical framework – a case study from a North Sea well 早渐新世恐龙囊肿作为古环境重建和地层格架的工具——以北海油井为例
IF 2 3区 地球科学 Q1 PALEONTOLOGY Pub Date : 2019-09-02 DOI: 10.5194/jm-38-143-2019
K. Śliwińska
Abstract. The lower Oligocene (Rupelian) successions are climate record archives of the early icehouse world in the Cenozoic. Even though the number of studies focussing on the generally cold Oligocene is increasing, little is known about climatic variations in the mid-latitudes to high latitudes of the Northern Hemisphere. One of the major obstacles is the lack of stratigraphically complete uppermost Eocene to Oligocene successions in these regions. This study focusses on dinoflagellate cysts (dinocysts) from a thick nearly complete Rupelian succession in the Syracuse Oils Norge A/S well 11/10-1 drilled in 1969 in the Norwegian part of the North Sea basin. The well provides a record of mid-latitude dinocyst assemblages, which yield key biostratigraphical and palaeoenvironmental information. All the analyses were undertaken on ditch cutting samples. The dinocyst assemblages confirm that the well penetrates about 600 m of Rupelian sediments and (as supported by correlation with the Nini-1 well) that the lowermost Rupelian (below the top or the last occurrence of Areosphaeridium diktyoplokum) is expanded. These assemblages also indicate the presence of two hiatuses: the first extends from the Lutetian to the Priabonian (equivalent to the D9nb–D12nb zones), and the second spans the Rupelian–Chattian boundary (equivalent to the D14nb subzone or the NSO-5 zone). Despite the risk of caving, the dinocyst assemblages support the existing sequence stratigraphic framework. The assemblages reflect a clear transition from distal to proximal deposition in the vicinity of the site (across the regional seismic sequences OSS-1 – OSS meaning Oligocene seismic sequence – to OSS-2). The proximal deltaic deposits of the OSS-2 regressive system tract (RST) are characterised by pulses of high sea-surface productivity and pronounced shifts in the dinocyst assemblages, reflecting a highly dynamic environment in a restricted marine to marginal marine setting. The Rupelian succession penetrated by well 11/10-1 yields one new species, Areoligera? barskii sp. nov., which is described here in detail. The cold-water-tolerant dinocyst Svalbardella cooksoniae is present in two intervals in the studied succession. These intervals are related to the early Oligocene cooling maxima (the Oi-1a and the Oi-2 events). Furthermore, these two intervals correlate with two local sequence boundaries, suggesting that they are most probably of glacioeustatic origin. From these observations, I postulate that the early icehouse climate played an important role in the depositional development of the Oligocene succession in the North Sea basin. Even though the Eocene–Oligocene transition interval is not complete (i.e. Lutetian to Priabonian is either missing or condensed), well 11/10-1 merits high-resolution studies of the early icehouse climate for the North Sea region. Although any detailed studies should ideally be undertaken on conventional cores instead of ditch cuttings, no such samples spa
摘要下渐新世(卢比安)序列是新生代早期冰库世界的气候记录档案。尽管关注普遍寒冷的渐新世的研究数量正在增加,但人们对北半球中纬度到高纬度的气候变化知之甚少。主要障碍之一是这些地区缺乏地层上完整的最上始新世至渐新世序列。本研究的重点是1969年在北海盆地挪威部分钻探的Syracuse Oils Norge a/S 11/10-1井中厚的、几乎完整的卢比安序列中的甲藻囊肿(甲藻囊肿)。该井提供了中纬度恐龙囊肿组合的记录,提供了关键的生物地层学和古环境信息。所有分析都是对沟渠切割样本进行的。恐龙囊肿组合证实,该井穿透约600 m的卢比安沉积物,以及(与Nini-1井的相关性所支持的)最下面的卢比安(在Aresphaeridium diktyoplokum顶部或最后一次出现的下方)扩展。这些组合还表明存在两个断裂带:第一个断裂带从卢泰阶延伸到普里亚博阶(相当于D9nb–D12nb带),第二个断裂带跨越卢比安-查特阶边界(相当于D14nb亚带或NSO-5带)。尽管存在坍塌的风险,但恐龙囊肿组合支持现有的层序地层格架。组合反映了现场附近从远端沉积到近端沉积的明显过渡(穿过区域地震序列OSS-1-OSS,即渐新世地震序列-OSS-2)。OSS-2海退体系域(RST)的近端三角洲沉积物的特征是海面生产力高的脉冲和恐龙壳组合的显著变化,反映了受限海洋到边缘海洋环境中的高度动态环境。11/10-1井穿透的卢比阶序列产生了一个新物种,Areoligera?barskii sp.nov.,本文对此进行了详细描述。在所研究的序列中,耐冷水的Svalbardella cooksoniae恐龙囊肿以两个间隔存在。这些间隔与渐新世早期的冷却最大值(Oi-1a和Oi-2事件)有关。此外,这两个间隔与两个局部序列边界相关,表明它们很可能是冰川成因的。根据这些观测结果,我推测早期冰室气候在北海盆地渐新世序列的沉积发展中起着重要作用。尽管始新世-渐新世的过渡期并不完整(即卢特阶至普里亚博阶缺失或浓缩),11/10-1井值得对北海地区早期冰库气候进行高分辨率研究。尽管任何详细的研究理想情况下都应该在传统岩心而不是沟渠岩屑上进行,但该地区不存在跨越始新世-渐新世过渡期的此类样品。
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引用次数: 6
Reconstructing the Christian Malford ecosystem in the Oxford Clay Formation (Callovian, Jurassic) of Wiltshire: exceptional preservation, taphonomy, burial and compaction 重建威尔特郡牛津粘土组(卡洛维,侏罗纪)的克里斯蒂安·马尔福生态系统:特殊的保存、埋藏和压实
IF 2 3区 地球科学 Q1 PALEONTOLOGY Pub Date : 2019-07-19 DOI: 10.5194/JM-38-133-2019
M. Hart, K. Page, G. Price, C. Smart
Abstract. The Christian Malford lagerstätte in the Oxford Clay Formation of Wiltshire contains exceptionally well-preserved squid-like cephalopods, including Belemnotheutis antiquus (Pearce). Some of these fossils preserve muscle tissue, contents of ink sacks and other soft parts of the squid, including arms with hooks in situ and the head area with statoliths (ear bones) present in life position. The preservation of soft-tissue material is usually taken as an indication of anoxic or dysaerobic conditions on the sea floor and within the enclosing sediments. Interestingly, in the prepared residues of all these sediments there are both statoliths and arm hooks as well as abundant, species-rich, assemblages of both foraminifera and ostracods. Such occurrences appear to be incompatible with an interpretation of potential sea floor anoxia. The mudstones of the Oxford Clay Formation may have been compacted by 70 %–80 % during de-watering and burial, and in such a fine-grained lithology samples collected for microfossil examination probably represent several thousand years and, therefore, a significant number of foraminiferal life cycles. Such samples (even if only 1–2 cm thick) could, potentially, include several oxic–anoxic cycles and, if coupled with compaction, generate the apparent coincidence of well-preserved, soft-bodied, cephalopods and diverse assemblages of benthic foraminifera.
摘要威尔特郡牛津粘土地层中的克里斯蒂安·马尔福德lagerstätte包含保存异常完好的鱿鱼状头足类动物,包括Belemnotheutis antius(皮尔斯)。其中一些化石保存了肌肉组织、墨汁袋的内容物和鱿鱼的其他柔软部分,包括有钩子的手臂和有耳石(耳骨)的头部区域。软组织物质的保存通常被认为是海底和周围沉积物缺氧或缺氧条件的指示。有趣的是,在所有这些沉积物的制备残留物中,既有statolite和臂钩,也有丰富的有孔虫和介形虫组合。这种现象似乎与潜在的海底缺氧的解释不相容。牛津粘土组的泥岩可能在脱水和掩埋过程中被压实了70% - 80%,在这样一个细粒度的岩性样品中收集的微化石检查可能代表了几千年的有孔虫生命周期,因此,有孔虫生命周期的数量很大。这样的样本(即使只有1-2厘米厚)可能包含几个氧-缺氧循环,如果加上压实,就会产生保存完好的软体、头足类和各种底栖有孔虫组合的明显巧合。
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引用次数: 4
Calcification depth of deep-dwelling planktonic foraminifera from the eastern North Atlantic constrained by stable oxygen isotope ratios of shells from stratified plankton tows 北大西洋东部深海浮游有孔虫的钙化深度:受分层浮游生物拖带壳稳定氧同位素比值的约束
IF 2 3区 地球科学 Q1 PALEONTOLOGY Pub Date : 2019-07-12 DOI: 10.5194/JM-38-113-2019
A. Rebotim, A. Voelker, L. Jonkers, J. Waniek, M. Schulz, M. Kučera
Abstract. Stable oxygen isotopes (δ18O) of planktonic foraminifera areone of the most used tools to reconstruct environmental conditions of thewater column. Since different species live and calcify at different depthsin the water column, the δ18O of sedimentary foraminiferareflects to a large degree the vertical habitat and interspecies δ18O differences and can thus potentially provide information on thevertical structure of the water column. However, to fully unlock thepotential of foraminifera as recorders of past surface water properties, itis necessary to understand how and under what conditions the environmentalsignal is incorporated into the calcite shells of individual species.Deep-dwelling species play a particularly important role in this contextsince their calcification depth reaches below the surface mixed layer. Herewe report δ18O measurements made on four deep-dwellingGloborotalia species collected with stratified plankton tows in the eastern NorthAtlantic. Size and crust effects on the δ18O signal wereevaluated showing that a larger size increases the δ18O of G. inflata andG. hirsuta, and a crust effect is reflected in a higher δ18O signal in G. truncatulinoides. The greatmajority of the δ18O values can be explained without invokingdisequilibrium calcification. When interpreted in this way the data implydepth-integrated calcification with progressive addition of calcite withdepth to about 300 m for G. inflata and to about 500 m for G. hirsuta. In G. scitula, despite a strongsubsurface maximum in abundance, the vertical δ18O profile isflat and appears dominated by a surface layer signal. In G. truncatulinoides, the δ18O profile follows equilibrium for each depth, implying a constanthabitat during growth at each depth layer. The δ18O values aremore consistent with the predictions of the Shackleton (1974)palaeotemperature equation, except in G. scitula which shows values more consistentwith the Kim and O'Neil (1997) prediction. In all cases, we observe adifference between the level where most of the specimens were present andthe depth where most of their shell appears to calcify.
摘要浮游有孔虫的稳定氧同位素(δ18O)是重建水体环境条件最常用的工具之一。由于不同物种在水柱的不同深度生活和钙化,沉积有孔虫的δ18O在很大程度上反映了垂直生境和种间δ18O差异,因此可以潜在地提供水柱垂直结构的信息。然而,为了充分发掘有孔虫作为过去地表水性质记录者的潜力,有必要了解环境信号是如何以及在什么条件下被纳入单个物种的方解石壳中的。深栖物种在这种情况下起着特别重要的作用,因为它们的钙化深度达到地表混合层以下。本文报道了在北大西洋东部用分层浮游生物拖曳采集的四种深海生物的δ18O测量结果。分析了尺寸和地壳对δ18O信号的影响,结果表明,尺寸越大,G. inflata和G. bla的δ18O值越大。地壳效应反映在G. truncatulinoides较高的δ18O信号上。绝大多数δ18O值可以不用不平衡钙化来解释。当以这种方式解释时,数据暗示了深度积分钙化,随着深度的增加,方解石的逐渐增加,对G. inflata来说是300米,对G. hirsuta来说是500米。在G. scitula中,尽管有很强的次表层最大值,但垂直δ18O剖面是平坦的,并以表层信号为主。在G. truncatulinoides中,δ18O剖面在每个深度都遵循平衡,这意味着在每个深度层的生长过程中都有一个恒定的栖息地。除G. scitula的δ18O值与Kim和O'Neil(1997)的预测值更为一致外,其余δ18O值与Shackleton(1974)古温度方程的预测值较为一致。在所有情况下,我们观察到大多数标本存在的水平和它们的大部分外壳似乎钙化的深度之间存在差异。
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引用次数: 9
Holocene ostracod assemblages from the Co To Islands, northeastern Vietnam 越南东北部哥托群岛全新世介形类组合
IF 2 3区 地球科学 Q1 PALEONTOLOGY Pub Date : 2019-06-18 DOI: 10.5194/JM-38-97-2019
Sota Niiyama, G. Tanaka, T. Komatsu, H. D. Doan, H. B. Nguyen, H. T. Trinh, Minh T. Nguyen
Abstract. In this study, weinvestigated ostracod assemblages from the Co To Islands in northeasternVietnam. We identified 77 ostracod species belonging to 46 genera in ninesurface sediment samples and recognized three biofacies (I, II, and III)based on Q-mode cluster analysis. The dominant species of biofacies I and IIwere Aurila hataii, Loxoconcha japonica, andXestoleberis hanaii, which commonly occur in seaweed beds fromsouthern China to Japan. This is the first report on the ostracod assemblagefrom the open-sea area in northeastern Vietnam. We clarified that theostracod assemblages in the Gulf of Tongking, including northern Vietnam,have a strong biogeographical relationship with East Asia seas. A newspecies, Loxoconcha cotoensis sp. nov., was described herein fromthe Co To Islands(http://www.zoobank.org/NomenclaturalActs/41d3fb9f-ae17-4215-82c1-0874a8bf1a30, last access: 3 June 2019).
摘要在这项研究中,我们调查了来自越南东北部Co To群岛的介形虫组合。在9个表层沉积物样品中,鉴定出介形虫77种,隶属于46属,并通过q型聚类分析识别出3种生物相(I、II、III)。生物相ⅰ和生物相ⅲ的优势种为海螺(Aurila hataii)、日本Loxoconcha japonica和海南xestoleberis hanaii,它们常见于中国南方至日本的海藻床中。这是越南东北部外海地区首次报道介形类群落。研究结果表明,包括越南北部在内的东京湾的介形类群落与东亚海域具有很强的生物地理关系。本文描述了Co To群岛的一个新种Loxoconcha cotoensis sp. nov. (http://www.zoobank.org/NomenclaturalActs/41d3fb9f-ae17-4215-82c1-0874a8bf1a30,最后访问时间:2019年6月3日)。
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引用次数: 3
Chemotaxonomy of domesticated grasses: a pathway to understanding the origins of agriculture 驯化牧草的化学分类学:理解农业起源的途径
IF 2 3区 地球科学 Q1 PALEONTOLOGY Pub Date : 2019-06-07 DOI: 10.5194/JM-38-83-2019
P. Jardine, W. Gosling, B. Lomax, Adele C. M. Julier, W. Fraser
Abstract. The grass family (Poaceae) is one of the mosteconomically important plant groups in the world today. In particular manymajor food crops, including rice, wheat, maize, rye, barley, oats andmillet, are grasses that were domesticated from wild progenitors during theHolocene. Archaeological evidence has provided key information ondomestication pathways of different grass lineages through time and space.However, the most abundant empirical archive of floral change – the pollenrecord – has been underused for reconstructing grass domesticationpatterns because of the challenges of classifying grass pollen grains basedon their morphology alone. Here, we test the potential of a novel approachfor pollen classification based on the chemical signature of the pollengrains measured using Fourier transform infrared (FTIR) microspectroscopy.We use a dataset of eight domesticated and wild grass species, classifiedusing k-nearest neighbour classification coupled with leave-one-out crossvalidation. We demonstrate a 95 % classification success rate on trainingdata and an 82 % classification success rate on validation data. Thisresult shows that FTIR spectroscopy can provide enhanced taxonomicresolution enabling species level assignment from pollen. This will enablethe full testing of the timing and drivers of domestication and agriculturethrough the Holocene.
摘要禾本科(禾本科)是当今世界经济上最重要的植物类群之一。特别是许多主要的粮食作物,包括水稻、小麦、玉米、黑麦、大麦、燕麦和小米,都是在全新世从野生祖先驯化而来的草。考古证据为不同草系在时间和空间上的形成路径提供了关键信息。然而,由于仅根据草花粉粒的形态对其进行分类的挑战,最丰富的花变化经验档案——花粉记录——在重建草驯化模式方面没有得到充分利用。在这里,我们测试了一种新的花粉分类方法的潜力,该方法基于使用傅立叶变换红外(FTIR)显微光谱法测量的花粉内含物的化学特征。我们使用了一个由八种驯化和野生草种组成的数据集,使用k近邻分类和留一交叉验证进行分类。我们展示了95 % 训练数据分类成功率与an82 % 验证数据的分类成功率。这一结果表明,FTIR光谱可以提供增强的分类分辨率,从而能够从花粉中进行物种级分配。这将有助于对全新世驯化和农业的时间和驱动因素进行全面测试。
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引用次数: 5
Species of the diatom taxa Aulacodiscus and Trinacria with biostratigraphic utility in Palaeogene and Neogene North Sea sediments 古近系和新近系北海沉积物中具有生物地层应用价值的硅藻类群Aulacodiscus和Trinacria
IF 2 3区 地球科学 Q1 PALEONTOLOGY Pub Date : 2019-05-29 DOI: 10.5194/JM-38-67-2019
A. Mitlehner
Abstract. Species of Aulacodiscus and Trinacria, two important marine diatom genera with biostratigraphic utility in offshore North Sea exploration and onshore correlation, are identified, described and emended and the North Sea microfaunal zonation scheme is revised accordingly. Occurring mainly as pyritised diatom moulds or steinkerns, detailed scanning electron microscope (SEM) analysis of several specimens, formerly in open nomenclature, has allowed the correct taxonomic identification of pyritised morphologies found to belong to the genus Aulacodiscus, including A. allorgei, A. heterostictus, A. insignis, A. singilewskyanus, A. subexcavatus and A. suspectus. The important marker species Trinacria regina is emended. SEM studies, using specimens preserved in pyrite and original silica, have shed further light on the varying forms and frustule morphology of Trinacria regina so that valves and frustules formerly thought to represent separate species are now found to be grouped within this taxon; SEM studies have shown that many of these variations represent different valves within a chain, whilst others may signify ecophenotypic variants. Emendments are therefore made to clarify the taxonomic status of different variants within T. regina, important in the Palaeocene–Eocene boundary interval onshore and offshore such as the Sele and Balder formations and the Fur Formation diatomite of Jutland, Denmark. Species of taxa formerly in open nomenclature are now assigned to Aulacodiscus insignis, which are important offshore markers in offshore late Oligocene to early Miocene sediments in northwest Europe.
摘要本文对Aulacodiscus和Trinacria这两个重要的海洋硅藻属进行了鉴定、描述和修正,并对北海微区系划分方案进行了相应的修订。主要以黄铁矿化硅藻霉菌或steinkerns的形式出现,对几个标本进行了详细的扫描电镜(SEM)分析,发现黄铁矿化形态属于Aulacodiscus属,包括A. alloorgei, A. heterostictus, A. insignis, A. singilewskyanus, A. subbexcavatus和A. susectus。对重要的标志种金银花进行了订正。利用保存在黄铁矿和原始二氧化硅中的标本进行的扫描电镜研究,进一步揭示了tracria regina的不同形态和截果形态,因此,以前被认为代表不同物种的阀和截果现在被发现归在这个分类单元中;扫描电镜研究表明,许多这些变异代表了链中的不同阀门,而其他变异可能意味着生态表型变异。因此,对T. regina中不同变体的分类地位进行了修正,T. regina在古新世-始新世边界段的陆上和海上都很重要,如丹麦日德兰半岛的Sele和Balder组以及Fur组硅藻土。在欧洲西北部晚渐新世至早中新世的近海沉积物中,原属开放命名的分类群种被划归为Aulacodiscus insignis。
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引用次数: 1
Where should we draw the lines between dinocyst “species”? Morphological continua in Black Sea dinocysts 我们应该如何区分恐龙“物种”?黑海恐龙囊的形态连续性
IF 2 3区 地球科学 Q1 PALEONTOLOGY Pub Date : 2019-04-26 DOI: 10.5194/JM-38-55-2019
T. M. Hoyle, Manuel Sala-Pérez, F. Sangiorgi
Abstract. The morphology of dinoflagellate cysts (dinocysts) is related not only to thegenetics of the motile dinoflagellate from which it derives, but is alsodependent on a range of environmental factors including salinity, temperatureand nutrient status. Although this knowledge improves our understanding ofthe drivers behind dinocyst morphological variations, it makes the taxonomygoverning their description somewhat complex. In basins such as the BlackSea, where environmental change can be extreme and occurs on relatively short(millennial) timescales, taxonomy becomes particularly challenging.Morphological continua can be observed between described forms, displaying alarge range of intermediate phenotypes that do not necessarily correspond toany genetic difference. As these morphological nuances may preserveinformation about palaeoenvironments, it is important to find a systematicmethod of characterising morphotypes. Here, we show a dinocyst matrix withinwhich dinocysts are described according to their similarity to (or differencefrom) described forms based on key descriptive parameters. In the example setout here, cyst shape and degree of process and/or ectophragmdevelopment are taken as two key parameters in Pyxidinopsis psilata and Spiniferites cruciformis, and can allow the description ofintermediate forms even though the definitions do not overlap. We review some frequently occurring morphotypes and propose that usingmatrices to show the gradual variation between endmember forms is the mostpragmatic approach until cyst–theca studies and genetic sequencing can beused to demonstrate relationships between genotypes and morphotypes. As priorstudies propose salinity to be a primary driver of intraspecific variability,the endmembers presented may represent salinity extremes within an overallbrackish environment. Although we cannot assign each morphotype to a value ora range of an environmental parameter (e.g. salinity) as the differentmorphotypes can occur in the same sample, using this matrix allowspreservation of information about morphological variability without creatingtaxonomic categories that are likely to require alteration if geneticevidence becomes available.
摘要鞭毛藻囊(dinocysts)的形态不仅与运动鞭毛藻的遗传有关,而且还取决于一系列环境因素,包括盐度、温度和营养状况。尽管这些知识提高了我们对恐龙囊形态变化背后驱动因素的理解,但它使支配它们描述的分类学变得有些复杂。在像黑海这样的盆地,环境变化可能是极端的,并且发生在相对较短的(千年)时间尺度上,分类学变得特别具有挑战性。在所描述的形式之间可以观察到形态连续性,显示出大范围的中间表型,这些表型不一定对应任何遗传差异。由于这些形态上的细微差别可能保存了有关古环境的信息,因此找到一种系统的方法来描述形态类型是很重要的。在这里,我们展示了一个dinocyst矩阵,在这个矩阵中,dinocyst是根据它们与基于关键描述参数的描述形式的相似性(或差异性)来描述的。在这里的例子中,在Pyxidinopsis psilata和spiferites crucformis中,囊肿的形状和过程和/或外膜发育的程度被作为两个关键参数,即使定义不重叠,也可以描述中间形式。我们回顾了一些经常发生的形态型,并提出使用矩阵来显示端元形式之间的逐渐变化是最实用的方法,直到囊膜研究和基因测序可以用来证明基因型和形态型之间的关系。由于先前的研究提出盐度是种内变异性的主要驱动因素,因此所呈现的端元可能代表了整体微咸环境中的极端盐度。虽然我们不能给每种形态分配一个值或一个环境参数的范围(例如盐度),因为不同的形态可能出现在同一个样本中,使用这个矩阵允许保留关于形态变异的信息,而不会创建可能需要改变的分类类别,如果遗传证据可用。
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引用次数: 10
Calcareous nannofossil assemblages of the Late Cretaceous Fiqa Formation, north Oman 阿曼北部晚白垩世Fiqa组钙质纳米化石组合
IF 2 3区 地球科学 Q1 PALEONTOLOGY Pub Date : 2019-02-28 DOI: 10.5194/JM-38-25-2019
Zainab Al Rawahi, T. Dunkley Jones
Abstract. This study presents thefirst detailed calcareous nannofossil assemblage data from the LateCretaceous succession of the subsurface Aruma Basin, north Oman. Thetaxonomic description and documentation of assemblage composition are basedon extensive quantitative analysis of ditch cuttings and side-wall samplesfrom eight hydrocarbon exploration wells across north Oman. The samplesstudied from those wells cover the Coniacian to earliest Maastrichtian deepmarine shales and marls of the subsurface Fiqa Formation. These fine-grainedsiliciclastic deposits often yield moderately to well-preserved nannofossilassemblages, especially in the Campanian intervals. Consequently, diverseassemblages have been recorded from the Fiqa Formation, with a totaldiversity of ∼200 species, including two new species,Staurolithites ormae sp. nov. and Chiastozygus fahudensissp. nov., which are illustrated and described. Extensive imaging of thisdiversity is provided here, as are stratigraphic distributions of the maincomponents from a key reference well in north Oman, W-4. Poorly describedgroups such as Staurolithites are closely investigated and theirutility for stratigraphic applications is highlighted. Relative abundances ofnannofossil taxa with strong palaeoenvironmental preferences have revealednew insights into the palaeo-productivity of the basin. High-fertilityspecies like Discorhabdus ignotus, Biscutum constans andZeugrhabdotus erectus show substantial variations in abundancethroughout the Fiqa Formation, reflecting long-term shifts in theproductivity conditions of the basin. This is supported by abundance patternsof Watznaueria barnesiae and Micula staurophora that show abroadly inverse correlation with the high-fertility species. The FiqaFormation represents a key regional seal unit for the Cretaceous hydrocarbonreservoirs of Oman, as well as being a productive unit elsewhere in theArabian Peninsula. Beyond the Aruma Basin of Oman, this study will provide akey reference point for future biostratigraphic or palaeoenvironmentalanalyses of the Late Cretaceous calcareous nannofossil assemblages across theMiddle East and other southern Tethyan areas.
摘要本研究首次提供了来自阿曼北部阿鲁马盆地地下晚白垩世序列的详细钙质超微化石组合数据。组合组成的分类学描述和文献记录是基于对阿曼北部八口油气探井的沟渠岩屑和侧壁样本的广泛定量分析。从这些井中研究的样本覆盖了地下Fiqa组的Coniacian至最早的Maastrichtian深海页岩和泥灰岩。这些细粒硅化岩矿床通常产出中等至保存良好的超微化石,尤其是在Campanian层段。因此,Fiqa组记录了多种组合,总多样性约为200种,包括两个新种,Staurolithites ormae sp.nov.和Chiastozygus fahudensis ssp。nov.,对其进行了说明和描述。这里提供了这种多样性的广泛成像,以及阿曼北部一口关键参考井W-4的主要成分的地层分布。对描述较差的组群,如铁云母进行了密切调查,并强调了它们在地层应用中的实用性。具有强烈古环境偏好的非化石类群的相对丰度为盆地的古生产力提供了新的见解。Discorhabdus ignatus、Biscutum constans和Zeugrhabdotus直立等高肥力物种在整个Fiqa组的丰度都有很大变化,反映了盆地生产力条件的长期变化。这一点得到了藤壶菌(Watznaueria barnesiae)和尾壶菌(Micula staurophora)丰度模式的支持,它们和高肥力物种呈广泛的负相关。FiqaFormation是阿曼白垩纪碳氢化合物储层的一个关键区域封闭单元,也是阿拉伯半岛其他地区的一个生产单元。在阿曼阿鲁马盆地之外,这项研究将为中东和其他特提斯南部地区晚白垩世钙质超微化石组合的未来生物地层学或古环境分析提供一个重要的参考点。
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引用次数: 2
期刊
Journal of Micropalaeontology
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