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The history of palaeontological research and excavations at Monte San Giorgio 圣乔治山古生物研究和发掘史
IF 3 2区 地球科学 Q1 PALEONTOLOGY Pub Date : 2024-05-02 DOI: 10.1186/s13358-024-00314-9
Heinz Furrer
There is a long history of palaeontological excavations at Monte San Giorgio (Switzerland) and the adjoining Monte Pravello—Monte Orsa (Italy), aimed at finding well-preserved skeletons of Middle Triassic vertebrates. The first fossils were discovered in the mid-Nineteenth Century during mining of black shales (scisti bituminosi) near Besano, Italy, with further finds in the early Twentieth Century through industrial-scale mining. Studies of the material generated international interest and prompted formal palaeontological excavations on both sides of the border. The earliest excavations took place in 1863 and 1878, with the most extensive between 1924 and 1968. Systematic excavations have continued up to the present day, focusing on six distinct fossiliferous horizons: the Besano Formation and the overlying Meride Limestone with the Cava inferiore, Cava superiore, Cassina, Sceltrich and Kalkschieferzone beds. All these have provided material for study and display, with Monte San Giorgio itself recently designated a UNESCO World Heritage Site. The workers and organisations involved, locations excavated and material recovered are described herein.
在圣乔治山(瑞士)和毗邻的普拉韦洛山-蒙特奥尔萨(意大利)进行古生物发掘的历史悠久,目的是寻找保存完好的三叠纪中期脊椎动物骨骼。第一批化石是十九世纪中叶在意大利贝萨诺附近开采黑色页岩(scisti bituminosi)时发现的,二十世纪初通过工业规模的开采又有新的发现。对这些材料的研究引起了国际兴趣,并促使边境两侧进行了正式的古生物发掘。最早的发掘发生在 1863 年和 1878 年,最大规模的发掘发生在 1924 年至 1968 年。系统的发掘工作一直持续到今天,主要集中在六个不同的化石地层:贝萨诺地层和上覆的梅里德石灰岩层,以及卡瓦下层、卡瓦上层、卡西纳层、斯凯尔特里希层和卡克施费尔宗层。所有这些都为研究和展示提供了素材,圣乔治山本身最近也被联合国教科文组织列为世界遗产。本文介绍了参与其中的工作人员和组织、发掘地点和出土材料。
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引用次数: 0
The Middle Triassic palaeontomofauna of Monte San Giorgio with the description of Merithone laetitiae (†Permithonidae) gen. et sp. nov. Monte San Giorgio 的中三叠世古生物群以及对 Merithone laetitiae (†Permithonidae) gen.
IF 3 2区 地球科学 Q1 PALEONTOLOGY Pub Date : 2024-04-29 DOI: 10.1186/s13358-024-00317-6
Matteo Montagna, Giulia Magoga, Fabio Magnani
The Triassic is considered a crucial period for the establishment of the modern insect fauna and fossil records from this period are fundamental for understanding the real impact that the end Permian Mass Extinction events had on these animals. Here, we review the insect fossils from one of the main deposits of this period in the world, Monte San Giorgio, which is considered one of the nine main insect Fossillagerstätten. In this Lagerstätte, located on the border between Switzerland and Italy, a total of 273 fossil insects have been collected in five localities. The fossils found in Val Mara site D, one of the two richest insect fossils sites of Monte San Giorgio, present peculiar features, such as extraordinary sizes and phosphatisation of internal tissues revealing fine internal details. In contrast, the Val Mara site VM 12 fossil record (248 specimens) is dominated by small to medium size insects, usually almost intact, preserving details such as setae on wings and compound eyes. Besides these exceptional features, these fossil insects are of extreme evolutionary importance, since they represent the first or the last occurrence for their lineage. In this regard, their use to calibrate nodes in a phylogenomic dating analysis led to backdating the origin of many insect lineages, including Diptera and Heteroptera. Up to now, a total of five species from Monte San Giorgio have been formally described, belonging to the orders Archaeognatha (†Monura and Machilidae), Ephemeroptera, Hemiptera (Tingidae) and Coleoptera (Adephaga). A further species, Merithone laetitiae (†Permithonidae) gen. et sp. nov., whose fossil is included among the recent findings in Val Mara site VM 12, is described in the present work.
三叠纪被认为是建立现代昆虫动物群的关键时期,这一时期的化石记录对于了解二叠纪末大灭绝事件对这些动物的真正影响至关重要。在这里,我们回顾了这一时期世界主要矿床之一--圣乔治山(Monte San Giorgio)--的昆虫化石,该矿床被认为是世界九大昆虫化石矿床之一。在这个位于瑞士和意大利交界处的昆虫化石采集站,共在五个地方采集到 273 块昆虫化石。瓦尔马拉遗址 D 是圣乔治山昆虫化石最丰富的两个遗址之一,在这里发现的化石具有一些特殊的特征,如体积特别大,内部组织磷化,显示出内部的精细细节。相比之下,瓦尔马拉遗址 VM 12 的化石记录(248 个标本)以中小型昆虫为主,通常几乎完好无损,保留了翅膀上的刚毛和复眼等细节。除了这些特殊的特征外,这些昆虫化石在进化过程中也具有极其重要的意义,因为它们代表了其家族的首次或最后一次出现。在这方面,利用它们来校准系统发生学年代分析中的节点,可以追溯包括双翅目和杂翅目在内的许多昆虫谱系的起源。迄今为止,圣乔治山共有五个物种被正式描述,分别属于古生代目(†Monura 和 Machilidae)、蜉蝣目、半翅目(Tingidae)和鞘翅目(Adephaga)。本研究还描述了另一个物种 Merithone laetitiae (†Permithonidae) gen.
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引用次数: 0
First true mastodon from the Late Miocene of Iran 伊朗中新世晚期的第一只真正的乳齿象
IF 3 2区 地球科学 Q1 PALEONTOLOGY Pub Date : 2024-04-18 DOI: 10.1186/s13358-023-00300-7
Sadaf Yaghoubi, Ali Reza Ashouri, Majid Mirzaie Ataabadi, Abbas Ghaderi
A mammutid is described here for the first time from the Late Miocene (MN12 equivalent) deposits of Abkhareh village, Varzeghan region, in the North-Western part of Iran. It is identified as “Mammut” cf. obliquelophus and is represented by an isolated and moderately worn upper third molar with a zygodont crown pattern typical of mammutids. In addition, two upper incisors found associated with the molar and probably belonging to the same individual are assigned as Mammut. The studied material expands the geographic distribution of “Mammut” obliquelophus into Western Asia. There are some known localities bearing vertebrate fossil beds from Northwest of Iran like Maragheh (MN12, Mirzaie Ataabadi et al., 2013c), Ivand (MN12, Mirzaie Ataabadi et al, 2011a) and Kivi-1 (MN12, Jafarzadeh and Konidaris, 2020). However, some materials belonging to proboscideans were revealed from a village, Abkhareh (MN12), in Varzeghan area 150 km away from Maragheh. In this study, an upper third molar and a pair of upper tusks of an ancient elephant-like mammal were attributed to the mammutid “Mammut” cf. obliquelophus. This finding proves that this taxon inhabited Iran between 8–7.5 Ma, which falls in the Turolian stage of the Late Miocene. It marks the first record of this species in Iran and expands the known geographic distribution of “Mammut” obliquelophus into Western Asia.
本文首次描述了伊朗西北部瓦尔泽甘地区 Abkhareh 村晚中新世(相当于 MN12)沉积层中的一种哺乳动物。它被鉴定为 "Mammut "cf.obliquelophus,其代表作是一颗孤立的、磨损程度适中的上第三臼齿,具有哺乳动物典型的颧齿冠纹。此外,发现的两颗与臼齿相关的上门齿可能属于同一个体,也被归类为猛犸象。所研究的材料将 "Mammut "obliquelophus的地理分布范围扩大到了西亚。伊朗西北部有一些已知的脊椎动物化石产地,如 Maragheh(MN12,Mirzaie Ataabadi 等人,2013c)、Ivand(MN12,Mirzaie Ataabadi 等人,2011a)和 Kivi-1(MN12,Jafarzadeh 和 Konidaris,2020)。不过,在距离马拉盖赫 150 公里的瓦尔泽甘地区的一个村庄 Abkhareh(MN12)发现了一些属于长鼻类的材料。在这项研究中,一种古象类哺乳动物的上第三臼齿和一对上獠牙被认为是长鼻类动物 "Mammut" cf. obliquelophus。这一发现证明,该类群曾在 8-7.5 Ma 之间的伊朗栖息,属于晚中新世的 Turolian 阶段。这标志着该物种在伊朗的首次记录,并将 "Mammut" obliquelophus 的已知地理分布范围扩大到了西亚。
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引用次数: 0
Orthoceratoid and coleoid cephalopods from the Middle Triassic of Switzerland with an updated taxonomic framework for Triassic Orthoceratoidea 瑞士中三叠世的正角类和鞘状头足类动物,以及更新的三叠世正角类动物分类框架
IF 3 2区 地球科学 Q1 PALEONTOLOGY Pub Date : 2024-04-03 DOI: 10.1186/s13358-024-00307-8
Alexander Pohle, Christian Klug
Orthoconic cephalopods are subordinate, but persistent, widespread and regionally abundant components of Triassic marine ecosystems. Here, we describe unpublished specimens from the Anisian (Middle Triassic) Besano Formation at Monte San Giorgio, Switzerland. They can be assigned to two major but unrelated lineages, the Coleoidea and the Orthoceratoidea. The orthoceratoids belong to Trematoceras elegans (Münster, 1841) and occur regularly within the Besano Formation, are uniform in size, and have few available morphological characters. In contrast, coleoids are more diverse and appear to be restricted to shorter intervals. A new coleoid is described as Ticinoteuthis chuchichaeschtli gen. et sp. nov. To better put the orthoceratoids of the Besano Formation into perspective, we also synthesise the current taxonomy of Triassic orthoceratoids on a global scale. The currently used scheme is largely outdated, with very little taxonomic progress in the past 100 years. Despite previous research showing the distinctness of Triassic orthoceratoids from Palaeozoic taxa, they are still commonly labelled as “Orthoceras” or “Michelinoceras”, which are confined to the Palaeozoic. We show that Triassic orthoceratoids probably belong to a single lineage, the Trematoceratidae, which can be assigned to the Pseudorthocerida based on the embryonic shell and endosiphuncular deposits. Many Triassic species can probably be assigned to Trematoceras, but there are at least two additional Triassic orthoceratoid genera, Paratrematoceras and Pseudotemperoceras. Finally, we review the palaeobiogeographic and stratigraphic distribution of the group and outline possible future research directions.
直齿头足类是三叠纪海洋生态系统中从属但持久、广泛和区域性的丰富成分。在这里,我们描述了来自瑞士圣乔治山安息(中三叠世)贝萨诺地层的未发表标本。这些标本可归属于两个主要但不相关的类群:鞘翅目(Coleoidea)和正戟目(Orthoceratoidea)。正角类属于 Trematoceras elegans(Münster,1841 年),经常出现在贝萨诺地层中,大小一致,几乎没有可用的形态特征。相比之下,鞘状石的种类更多,而且似乎仅限于较短的间隔。一种新的鞘状类被描述为 Ticinoteuthis chuchichaeschtli gen.为了更好地认识贝萨诺地层的正角类,我们还综合了目前全球范围内三叠纪正角类的分类方法。目前使用的分类方法在很大程度上已经过时,在过去的 100 年中分类工作进展甚微。尽管之前的研究表明三叠纪正长岩类群与古生代类群截然不同,但它们仍然通常被标记为 "Orthoceras "或 "Michelinoceras",仅限于古生代。我们的研究表明,三叠纪的直角戟类可能属于一个单系,即 Trematoceratidae,根据胚壳和内鞘沉积物,可将其归入假直角戟类。三叠纪的许多物种可能归属于Trematoceras,但至少还有另外两个三叠纪正角类属:Paratrematoceras和Pseudotemperoceras。最后,我们回顾了该类群的古生物地理和地层分布情况,并概述了未来可能的研究方向。
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引用次数: 0
A redescription of Trachelosaurus fischeri from the Buntsandstein (Middle Triassic) of Bernburg, Germany: the first European Dinocephalosaurus-like marine reptile and its systematic implications for long-necked early archosauromorphs 德国伯恩堡 Buntsandstein(中三叠世)Trachelosaurus fischeri 的重新描述:欧洲第一只类似恐龙的海洋爬行动物及其对长颈早期古龙类的系统影响
IF 3 2区 地球科学 Q1 PALEONTOLOGY Pub Date : 2024-03-15 DOI: 10.1186/s13358-024-00309-6
Stephan N. F. Spiekman, Martín D. Ezcurra, Adam Rytel, Wei Wang, Eudald Mujal, Michael Buchwitz, Rainer R. Schoch
Some of the earliest members of the archosaur-lineage (i.e., non-archosauriform archosauromorphs) are characterised by an extremely elongated neck. Recent fossil discoveries from the Guanling Formation (Middle Triassic) of southern China have revealed a dramatic increase in the known ecomorphological diversity of these extremely long-necked archosauromorphs, including the fully marine and viviparous Dinocephalosaurus orientalis. These recent discoveries merit a reinvestigation of enigmatic Triassic diapsid fossils from contemporaneous European deposits housed in historical collections. Here, we provide a redescription of Trachelosaurus fischeri, represented by a single, disarticulated specimen first described in 1918. Due to its unique morphology, which includes short, bifurcating cervical ribs, and a high presacral vertebral count, this taxon has been referred to either as a “protorosaurian” archosauromorph or a sauropterygian. Our revision clearly shows that Trachelosaurus represents the first unambiguous Dinocephalosaurus-like archosauromorph known from outside the Guanling Formation. Our finding has important systematic implications. Trachelosauridae Abel, 1919 represents the senior synonym for the recently identified Dinocephalosauridae Spiekman, Fraser and Scheyer, 2021. Based on our phylogenetic analyses, which employ two extensive datasets, we also corroborate previous findings that tanystropheids and trachelosaurids represent two families within a larger monophyletic group among non-crocopodan archosauromorphs, which is here named Tanysauria (clade nov.). Trachelosauridae is minimally composed of Trachelosaurus fischeri, Dinocephalosaurus orientalis, Pectodens zhenyuensis, and Austronaga minuta, but one of our analyses also found a probably taxonomically broader clade that may also include Gracilicollum latens and Fuyuansaurus acutirostris. Trachelosaurus fischeri considerably expands the known spatial and temporal range of Trachelosauridae to the earliest Anisian and the Central European Basin. Our findings add to the growing evidence for the presence of a diverse group of fully marine reptiles during the Middle Triassic among Tanysauria. These trachelosaurids possess flipper-like limbs, high vertebral counts, and elongate necks, thus superficially resembling long-necked Jurassic and Cretaceous plesiosaurs in some regards.
古龙类(即非古龙形态的古龙类)的一些最早成员具有极长的颈部。最近在中国南方关岭地层(中三叠世)发现的化石显示,这些极长颈的弓形龙类的已知非形态多样性急剧增加,其中包括完全生活在海洋中的东方恐龙。最近的这些发现值得我们重新研究历史收藏中来自同时代欧洲矿床的神秘的三叠纪双脊龙化石。在这里,我们对 Trachelosaurus fischeri 进行了重新描述,其代表化石是 1918 年首次描述的一个单独的解体标本。由于其独特的形态,包括短的、分叉的颈肋和较多的椎骨,该类群一直被称为 "原龙类 "古龙形目或长棘龙类。我们的修订清楚地表明,驯龙是关岭地层以外已知的第一个明确的类恐龙弓龙。我们的发现具有重要的系统学意义。Trachelosauridae Abel, 1919 是最近发现的恐龙头龙科 Spiekman, Fraser 和 Scheyer, 2021 的高级异名。基于我们的系统发育分析(采用了两个广泛的数据集),我们也证实了之前的发现,即栉龙科和三角龙科是非鳄龙类中一个更大的单系群中的两个科,在此命名为栉龙科(新支系)。栉龙科(Trachelosauridae)由栉龙(Trachelosaurus fischeri)、东方棘龙(Dinocephalosaurus orientalis)、振龙(Pectodens zhenyuensis)和小栉龙(Austronaga minuta)组成,但我们的一项分析还发现了一个可能在分类学上更为宽泛的支系,该支系可能还包括栉龙(Gracilicollum latens)和福源龙(Fuyuansaurus acutirostris)。栉龙(Trachelosaurus fischeri)大大扩展了栉龙科的已知时空范围,将其扩展到了最早的安息年和中欧盆地。我们的发现增加了越来越多的证据,证明在中三叠世湍龙科中存在着一个多样化的完全海洋性的爬行类群。这些栉龙类拥有鳍状肢、高椎骨数和长颈,因此在某些方面与侏罗纪和白垩纪的长颈长颈龙非常相似。
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引用次数: 0
Otoliths of the Gobiidae from the Neogene of tropical America 美洲热带新近纪戈壁鱼科的耳石
IF 3 2区 地球科学 Q1 PALEONTOLOGY Pub Date : 2024-03-06 DOI: 10.1186/s13358-023-00302-5
Werner W. Schwarzhans, Orangel A. Aguilera
Otoliths are common and diverse in the Neogene of tropical America. Following previous studies of Neogene tropical American otoliths of the lanternfishes (Myctophidae), marine catfishes (Ariidae), croakers (Sciaenidae), and cusk-eels (Ophidiiformes), we describe here the otoliths of the gobies (Gobiidae). The Gobiidae represent the richest marine fish family, with more than 2000 species worldwide and about 250 in America. In the fossil record too they are the species richest family in the Neogene of tropical America. We have investigated otoliths sampled from Ecuador, Pacific and Atlantic Panama, Atlantic Costa Rica, Dominican Republic, Venezuela, and Trinidad, ranging in age from late Early Miocene (late Burdigalian) to late Early Pleistocene (Calabrian). Most of the studied material originates from the collection expeditions of the Panama Paleontology Project (PPP). Our study represents the first comprehensive record of fossil gobies from America, and we recognize 107 species, of which 51 are new to science, 35 are in open nomenclature, and 19 represent species that also live in the region today. Previously, only two fossil otolith-based goby species have been described from the Neogene of tropical America. The dominant gobies in the fossil record of the region are from the Gobiosomatini, particularly of genera living over soft bottoms or in deeper water such as Bollmannia, Microgobius, Antilligobius, and Palatogobius. Another purpose of our study is to provide a first comprehensive account of otoliths of the extant Gobiidae of America, which we consider necessary for an adequate identification and interpretation of the Neogene otoliths. We studied otoliths of 130 extant American gobiid species and figured 106 of them for comparison. We also present a morphological analysis and characterization of the extant otoliths as a basis for the identification of fossil otoliths. Problems that commonly arise with the identification of fossil otoliths and specifically of fossil goby otoliths are addressed and discussed. A comparison of the history of the Gobiidae in tropical America reveals a high percentage of shared species between the Pacific and the Atlantic basins during the Late Miocene (Tortonian and Messinian) from at least 11 to 6 Ma. A recording gap on the Pacific side across the Pliocene allows a comparison again only in the late Early Pleistocene (Calabrian, 1.8 to 0.78 Ma), which shows a complete lack of shared species. These observations support the effective closure of the former Central American Seaway and emersion of the Isthmus of Panama in the intervening time. Groups that today only exist in the East Pacific were also identified in the Miocene and Pliocene of the West Atlantic, and there is also at least one instance of a genus now restricted to the West Atlantic having occurred in the East Pacific as late as the Pleistocene. The evolution of gobies in tropical America and the implications thereof are extensively discussed. Furthermore
耳石在美洲热带新近纪十分常见,而且种类繁多。继之前对灯笼鱼科(Myctophidae)、海鲶科(Ariidae)、黄花鱼科(Sciaenidae)和黄鳝科(Ophidiiformes)的新元古代热带美洲耳石的研究之后,我们在此介绍虾虎鱼科(Gobiidae)的耳石。虾虎鱼科是最丰富的海洋鱼类家族,在全世界有 2000 多个物种,在美洲约有 250 个物种。在化石记录中,它们也是美洲热带新近纪物种最丰富的科。我们研究了从厄瓜多尔、太平洋和大西洋巴拿马、大西洋哥斯达黎加、多米尼加共和国、委内瑞拉和特立尼达取样的耳石,其年代从早中新世晚期(布迪加里安晚期)到早更新世晚期(卡拉布里亚)不等。研究的大部分材料来自巴拿马古生物学项目(PPP)的采集考察。我们的研究首次全面记录了美洲的虾虎鱼化石,我们确认了 107 个物种,其中 51 个是科学界的新物种,35 个属于开放命名,19 个代表了今天也生活在该地区的物种。在此之前,美洲热带新近纪只描述了两个基于耳石的虾虎鱼化石物种。该地区化石记录中最主要的虾虎鱼属于虾虎鱼科,尤其是生活在软底或深水中的虾虎鱼属,如 Bollmannia、Microgobius、Antilligobius 和 Palatogobius。我们研究的另一个目的是首次全面介绍美洲现生戈壁鱼科的耳石,我们认为这对于充分鉴定和解释新近纪耳石是必要的。我们研究了 130 种现存美洲戈壁鱼类的耳石,并对其中 106 种进行了图解,以便进行比较。我们还对现生耳石进行了形态分析和特征描述,以此作为鉴定耳石化石的基础。我们还讨论了在鉴定耳石化石,特别是虾虎鱼耳石化石时经常出现的问题。通过比较美洲热带戈壁鱼科的历史,发现在晚中新世(托尔托尼世和梅西尼亚世)期间,太平洋盆地和大西洋盆地至少在 11 至 6 Ma 之间有很高比例的共享物种。由于太平洋一侧在上新世的记录空白,因此只有在早更新世晚期(卡拉布里亚,1.8-0.78 Ma)才能再次进行比较,结果显示完全没有共享物种。这些观察结果支持了前中美洲海道的有效关闭和巴拿马地峡在这一时期的形成。在西大西洋的中新世和上新世也发现了今天只存在于东太平洋的鱼群,而且至少有一个现在仅限于西大西洋的鱼属早在更新世就出现在东太平洋。文章广泛讨论了虾虎鱼在美洲热带地区的演化及其影响。此外,还从古环境迹象和古生物地理学方面讨论了对该地区虾虎鱼化石的观察。
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引用次数: 0
Digital skull anatomy of the Oligocene North American tortoise Stylemys nebrascensis with taxonomic comments on the species and comparisons with extant testudinids of the Gopherus–Manouria clade 始新世北美陆龟 Stylemys nebrascensis 的数字头骨解剖图,附该物种的分类评论以及与现存地鳖-马努里亚科陆龟的比较
IF 3 2区 地球科学 Q1 PALEONTOLOGY Pub Date : 2024-03-05 DOI: 10.1186/s13358-024-00311-y
Serjoscha W. Evers, Zahra Al Iawati
The anatomy of North American tortoises is poorly understood, despite a rich fossil record from the Eocene and younger strata. Stylemys nebrascensis is a particularly noteworthy turtle in this regard, as hundreds of specimens are known from Oligocene deposits, and as this species is one of the earliest fossil turtles to have been described in the scientific literature. Since its initial description based on a shell, many specimens with more complete material have been referred to Stylemys nebrascensis. Here, we review and confirm the referral of an important historic specimen to Stylemys nebrascensis, which includes shell, non-shell postcranial, and skull material. This allows us to document unique skull features of Stylemys nebrascensis (e.g., an unusual ‘poststapedial canal’ that connects the posterior skull surface with the cavum acustico-jugulare) and to refer another well-preserved skull to the species. Based on computed-tomography scanning of these two skulls, we provide a detailed description of the cranial and mandibular osteology of Stylemys nebrascensis. Stylemys nebrascensis has a combination of plesiomorphic skull characteristics (e.g., retention of a medial jugal process) and derived traits shared with extant gopher tortoises (e.g., median premaxillary ridge) that suggest it may be a stem-representative of the gopher tortoise lineage. This supports the hypothesis that extant and fossil tortoises from North America form a geographically restricted clade that split from Asian relatives during the Paleogene.
尽管始新世和更早的地层中有丰富的化石记录,但人们对北美陆龟的解剖结构知之甚少。在这方面,Stylemys nebrascensis 是一个特别值得注意的龟类,因为从渐新世的沉积中已知有数百个标本,而且该物种是科学文献中最早描述的龟类化石之一。自最初根据龟壳对其进行描述以来,许多更完整的标本都被归类为Stylemys nebrascensis。在这里,我们回顾并确认了将一个重要的历史标本归入Stylemys nebrascensis的观点,其中包括壳、非壳颅后和头骨材料。这使我们能够记录下Stylemys nebrascensis独特的头骨特征(例如,一个不寻常的 "后颅管",连接头骨后表面与凹腔),并将另一个保存完好的头骨归入该物种。基于对这两个头骨的计算机断层扫描,我们对Stylemys nebrascensis的头骨和下颌骨进行了详细的描述。Stylemys nebrascensis的头骨既具有同种异形的特征(如保留内侧颈突),又具有与现生地鼠陆龟共享的衍生特征(如前颌中脊),这表明它可能是地鼠陆龟的一个干系代表。这支持了这样的假说,即现存的北美陆龟和化石陆龟形成了一个地理上受到限制的支系,该支系在古近纪期间从亚洲亲缘种中分裂出来。
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引用次数: 0
The marine conservation deposits of Monte San Giorgio (Switzerland, Italy): the prototype of Triassic black shale Lagerstätten 圣乔治山(瑞士,意大利)的海洋保护矿床:三叠纪黑色页岩 Lagerstätten 的原型
IF 3 2区 地球科学 Q1 PALEONTOLOGY Pub Date : 2024-03-04 DOI: 10.1186/s13358-024-00308-7
Christian Klug, Stephan N. F. Spiekman, Dylan Bastiaans, Beat Scheffold, Torsten M. Scheyer
Marine conservation deposits (‘Konservat-Lagerstätten’) are characterized by their mode of fossil preservation, faunal composition and sedimentary facies. Here, we review these characteristics with respect to the famous conservation deposit of the Besano Formation (formerly Grenzbitumenzone; including the Anisian–Ladinian boundary), and the successively younger fossil-bearing units Cava inferiore, Cava superiore, Cassina beds and the Kalkschieferzone of Monte San Giorgio (Switzerland and Italy). We compare these units to a selection of important black shale-type Lagerstätten of the global Phanerozoic plus the Ediacaran in order to detect commonalities in their facies, genesis, and fossil content using principal component and hierarchical cluster analyses. Further, we put the Monte San Giorgio type Fossillagerstätten into the context of other comparable Triassic deposits worldwide based on their fossil content. The results of the principal component and cluster analyses allow a subdivision of the 45 analysed Lagerstätten into four groups, for which we suggest the use of the corresponding pioneering localities: Burgess type for the early Palaeozoic black shales, Monte San Giorgio type for the Triassic black shales, Holzmaden type for the pyrite-rich black shales and Solnhofen type for platy limestones.
海洋保护沉积("Konservat-Lagerstätten")的特点在于其化石保存方式、动物组成和沉积面貌。在此,我们回顾了著名的贝萨诺地层保护沉积(前格伦兹比图门岗,包括安尼西-拉迪尼边界)以及较年轻的含化石单元卡瓦下层、卡瓦上层、卡西纳床和圣乔治山(瑞士和意大利)的卡尔斯奇铁矿带的这些特征。我们将这些单元与全球新生代和埃迪卡拉纪的部分重要黑色页岩型Lagerstätten进行了比较,以便利用主成分和分层聚类分析发现它们在面貌、成因和化石含量方面的共性。此外,我们还根据化石含量,将圣乔治山类型化石矿床与全球其他可比的三叠纪矿床进行了比较。根据主成分分析和聚类分析的结果,我们将所分析的 45 个 Lagerstätten 分为四组,并建议使用相应的先驱地点:早古生代黑色页岩属于伯吉斯类型,三叠纪黑色页岩属于圣乔治山类型,富含黄铁矿的黑色页岩属于霍尔茨马登类型,板状灰岩属于索尔恩霍芬类型。
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引用次数: 0
The first Cyclida from the Triassic of Italy 意大利三叠纪最早的独眼巨人
IF 3 2区 地球科学 Q1 PALEONTOLOGY Pub Date : 2024-02-28 DOI: 10.1186/s13358-024-00306-9
Vittorio Pieroni
A well-preserved carapace of the crustacean Halicyne is here described. The finding comes from the Sostegno Basin (Piedmont, Italy). This is the first occurrence of a well-preserved arthropod from the Middle Triassic San Salvatore Formation of the Biellese area and the first report of a Triassic Cyclida from Italy. Cyclida often occurs in shallow marine environments with rapidly changing salinity conditions. The finding from the Sostegno area suggests a hypersaline paleoenvironment similar to that of Monte San Salvatore (Ticino, Switzerland) upper levels, where the same taxon was previously documented. These fossiliferous localities are shortly compared with that of Rasa di Varese (Lombardy, Italy).
本文描述了一种保存完好的甲壳类动物 Halicyne 的甲壳。该发现来自索斯特格诺盆地(意大利皮埃蒙特)。这是首次在比埃利塞地区三叠纪中期的圣萨尔瓦多地层中发现保存完好的节肢动物,也是意大利首次报道三叠纪的Cyclida。Cyclida通常出现在盐度条件快速变化的浅海环境中。索斯特格诺地区的这一发现表明,这里的古环境与圣萨尔瓦多山(瑞士提契诺州)上层类似,之前在那里也发现过相同的分类群。这些化石产地不久将与 Rasa di Varese(意大利伦巴第)的化石产地进行比较。
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引用次数: 0
Bernhard Peyer and his discoveries of Triassic vertebrates in Switzerland Bernhard Peyer 和他在瑞士发现的三叠纪脊椎动物
IF 3 2区 地球科学 Q1 PALEONTOLOGY Pub Date : 2024-02-28 DOI: 10.1186/s13358-024-00310-z
Hans-Dieter Sues
Bernhard Peyer (1885–1963) was a prominent Swiss vertebrate palaeontologist and anatomist at the University of Zurich. In 1919, he discovered the occurrence of Middle Triassic (Anisian–Ladinian) ichthyosaurs, sauropterygians, and other reptiles at Monte San Giorgio on the border between Switzerland and Italy. Starting in 1924, Peyer, and later his student and successor Emil Kuhn-Schnyder, worked tirelessly to excavate and collect material from fossil-bearing deposits in this region. The discovery of an articulated skeleton of a long-necked reptile in 1929 allowed Peyer to solve the riddle of unusually elongate bones from the German Muschelkalk, which, he demonstrated, were cervical vertebrae of the extraordinarily long-necked reptile Tanystropheus. As a student in 1919 and later in 1942, Peyer explored a Rhaetian bonebed close to his native town of Schaffhausen. Processing the sediment, he recovered numerous small bones and teeth of vertebrates, among which teeth of diverse stem-mammals are of particular importance.
Bernhard Peyer(1885-1963 年)是瑞士著名的古脊椎动物学家和苏黎世大学解剖学家。1919 年,他在瑞士和意大利交界处的圣乔治山发现了三叠纪中期(阿尼斯期-拉迪尼期)的鱼龙、长鼻龙和其他爬行动物。从 1924 年开始,佩耶和他后来的学生和继任者埃米尔-库恩-施奈德(Emil Kuhn-Schnyder)孜孜不倦地在这一地区的化石矿床中挖掘和收集材料。1929 年,佩耶发现了一具长颈爬行动物的关节骨架,从而解开了德国 Muschelkalk 地区异常细长骨骼的谜团,他证明这些骨骼是超长颈爬行动物 Tanystropheus 的颈椎骨。1919 年和 1942 年,作为一名学生,佩耶在他的家乡沙夫豪森附近勘探了一个雷提亚骨床。在处理沉积物时,他发现了许多脊椎动物的小骨头和牙齿,其中各种茎哺乳动物的牙齿尤为重要。
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引用次数: 0
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Swiss Journal of Palaeontology
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