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A thin-skinned thrust model for ice-marginal glacitectonic detachment and emplacement of Carboniferous bedrock rafts at Kilcummin Head, NW Ireland 爱尔兰西北部 Kilcummin Head 冰缘冰蚀剥离和石炭纪基岩筏置换的薄层推力模型
IF 1.2 4区 地球科学 Q2 Earth and Planetary Sciences Pub Date : 2024-06-01 DOI: 10.1016/j.pgeola.2024.04.001
David P. Vaughan , Emrys Phillips , Jonathan R. Lee , Jane K. Hart

This paper presents the results of a detailed macro- and microscale structural study of the glacitectonised sequence at Kilcummin Head on the western side of Killala Bay, County Mayo, northwest Ireland. The sequence comprises two laterally extensive, thrust-stacked rafts of Carboniferous limestone and mudstone emplaced upon the in situ bedrock of the Ballina Limestone Formation. Restored cross-sections reveal that the glacitectonised sequence has undergone extensive deformation and shortening of 61 %. This accompanied the formation of a prominent thrust-moraine associated with a northward ice advance across Killala Bay during the Midlandian (Devensian). A five-stage model is proposed to describe the deformation. Stage 1 comprises detachment of the rafts, and the majority of the sequence shortening. North-directed transport of the rafts occurred on a major décollement surface located at the base of the rafted sequence, marked by a mudstone-rich glacitectonite. Stage 2 accommodated further shortening via a series of imbricate thrusts leading to the stacking of the bedrock rafts (Stages 3 and 4). Stage 5 saw final stacking of rafts within a prominent thrust-block moraine, followed by the deposition of a sequence of coarse-grained, ice-marginal glacigenic deposits.

本文介绍了对爱尔兰西北部梅奥郡基拉拉湾西侧基尔康明头冰蚀序列进行的详细宏观和微观结构研究的结果。该地层由两块横向扩展、推力堆积的石炭纪石灰岩和泥岩组成,位于巴利纳石灰岩地层的原位基岩上。复原的横截面显示,冰蚀序列经历了广泛的变形和 61% 的缩短。这伴随着一个突出的推力碛石的形成,该推力碛石与德文西元(Devensian)中叶穿越基拉拉湾的北向冰进有关。我们提出了一个五阶段模型来描述这种变形。第一阶段包括筏状构造的剥离和大部分序列的缩短。筏状岩层的北向搬运发生在位于筏状岩层底部的一个主要的解理面,该解理面以富含泥岩的冰花岩为标志。第 2 阶段通过一系列覆岩推力进一步缩短,导致基岩筏堆积(第 3 和第 4 阶段)。第 5 阶段,基岩筏最终堆积在一个突出的推力块冰碛中,随后沉积了一系列粗粒冰缘冰原沉积物。
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引用次数: 0
A re-evaluation of Scelidosaurus remains from Ireland and the importance of apomorphy-based identifications 重新评估爱尔兰的漩涡龙遗骸以及基于同构体鉴定的重要性
IF 1.2 4区 地球科学 Q2 Earth and Planetary Sciences Pub Date : 2024-06-01 DOI: 10.1016/j.pgeola.2024.03.002
Kieran G. Satchell

Scelidosaurus is a herbivorous dinosaur belonging to the ornithischian clade Thyreophora. Until recently, its distribution was limited to the cliffs of Charmouth, on the south coast of Dorset, United Kingdom. In 2021, a fragmentary specimen from Lower Jurassic strata in Northern Ireland (BELUM K3998) was referred tentatively to Scelidosaurus. This bone, the proximal end of a femur, was suggested to be an ornithischian based on several characteristics, including gross morphology and osteohistology. This study reassesses these characteristics, and finds no evidence to support their use for identifying this specimen as either an ornithischian or Scelidosaurus. With the most comprehensive studies on Scelidosaurus finding no autapomorphies of the proximal femur, this study regards BELUM K3998 as Dinosauria indet. Additionally, this study highlights the importance of apomorphy-based approaches in taxonomy and the benefits of such methodologies.

箭龙是一种食草恐龙,属于鸟臀目恐龙科。直到最近,它的分布范围还仅限于英国多塞特郡南海岸的查莫斯悬崖。2021 年,北爱尔兰下侏罗世地层中的一块残缺标本(BELUM K3998)被暂时归类为霹雳龙。这块骨骼是股骨的近端,根据其大体形态学和骨组织学等几个特征,被认为是兽脚类。本研究对这些特征进行了重新评估,发现没有证据支持将该标本鉴定为鸟龙或箭龙。关于箭龙的最全面的研究发现,其股骨近端没有自同形的特征,因此本研究将 BELUM K3998 视为恐龙亚目(Dinosauria indet)。此外,本研究还强调了基于同形异构的方法在分类学中的重要性以及这种方法的益处。
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引用次数: 0
The first Protosiren remains preserved in ornamental limestones, Middle Eocene, North Eastern Desert, Egypt 埃及东北部沙漠中始新世观赏灰岩中保存的第一具原生动物遗骸
IF 1.2 4区 地球科学 Q2 Earth and Planetary Sciences Pub Date : 2024-06-01 DOI: 10.1016/j.pgeola.2024.04.005
Ahmed Mamdouh , Ramadan M. El-Kahawy , Mohamed AbdelGawad , Gebely Abu El-Kheir

A recent revelation has come to our attention, revealing the remarkably well-preserved post-cranial remains belonging to Protosiren. These remains include vertebrae, ribs, and fragmentary bones, and were discovered from two different horizons located in the Mokattam Formation of El-Galala Plateau, North Eastern Desert of Egypt. The first specimen, GCU0101, is identified as Protosiren sp., whereas the GCU0201 specimen is recognised as Protosiren cf. P. fraasi due to certain distinguishing features. These include vertebrae with wide keyhole-shaped neural canals, reduced tuberculum on the ribs, lack of pachyostotic ribs, and the possession of rugose articular rib head. This discovery is remarkable, as it offers a unique opportunity to study the ancient marine mammal's morphology and osteology, shedding light on its evolutionary history and ecological niche. The study documents the first occurrence of Protosiren from an ornamental limestone of the Middle Eocene (Bartonian) Observatory Formation from El-Galala Plateau, Northeastern Desert, Egypt.

最近的一项发现引起了我们的注意,它揭示了属于原生动物的保存完好的后颅骨遗骸。这些遗骸包括脊椎骨、肋骨和碎骨,发现于埃及东北部沙漠加拉拉高原莫卡塔姆地层的两个不同地层。第一个标本(GCU0101)被认定为原蛇,而 GCU0201 标本则由于某些显著特征被认定为原蛇(Protosiren cf. P. fraasi)。这些特征包括椎骨具有宽大的锁孔状神经管、肋骨上的小瘤减少、缺乏椎骨畸形肋骨以及具有皱褶状关节肋骨头。这一发现意义非凡,它为研究这种古老海洋哺乳动物的形态学和骨学提供了一个独特的机会,揭示了它的进化史和生态位。该研究记录了埃及东北部沙漠加拉拉高原中始新世(巴顿期)天文台地层观赏石灰岩中首次出现的普罗托海龙。
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引用次数: 0
A Miocene (Tortonian, Menorca, Spain) benthic macrofauna preserved in a phosphatic hardground: A difficult but invaluable record of upwelling palaeoenvironments 保存在磷酸盐硬地中的中新世(西班牙梅诺卡岛托尔托尼世)底栖大型动物群:上涌古环境的艰难但宝贵的记录
IF 1.2 4区 地球科学 Q2 Earth and Planetary Sciences Pub Date : 2024-06-01 DOI: 10.1016/j.pgeola.2024.04.002
Robin I. Knight

This field study records that the Tortonian phosphatic hardground at S'Algar, Menorca has a diverse fossil macrobenthic fauna dominated by bivalves. These faunal elements occur in four preservational forms (A–D). Each taphonomic form within the hardground is characterised by different concentrations of Entobia isp. (clionaid sponge boring) and encrustation. The preservational forms have specific fossil macrofaunas associated with them, and the hardground represents a melange of bioclasts derived from different environments on the Menorcan Miocene shelf. These habitats are influenced by the upwelling event that provides the conditions for hardground mineralisation.

A Glycymeris-Crassatella-Clypeaster association (A) records part of a fauna that lived on the upper ramp slope that was periodically affected by high energy and low oxygen conditions due to upwelling waters. The smaller sized fauna (B), characterised by an arcid-carditid-venerid association is derived from the middle ramp that was affected by storms and a more persistent westward flowing current that intermittently drove upwelling. The fragmentary and disarticulated nature of both faunas indicates that they were mixed within the hardground via reworking processes driven by these seafloor energy regimes.

Some of the large gastropod taxa, dentition-up Glycymeris and Gastrochaena suggest that the hardground had its own fauna. Preservation A fauna and other opportunistic taxa inhabited an environment where the hardground was exposed on the Miocene seafloor as ‘islands’ surrounded by thin layers of looser sediment. Phosphatic bioclasts in the underlying limestone indicate that similar mineralised beds were formed elsewhere on the shelf prior to the formation of the studied outcrop.

这项实地研究表明,梅诺卡岛 S'Algar 的托尔托尼磷酸盐硬质岩中有以双壳类为主的多种大型底栖动物化石。这些动物以四种保存形式(A-D)出现。硬岩层中的每种岩石形态都以不同浓度的 Entobia isp.(海绵钻孔)和结壳。保存形式有与之相关的特定大型生物化石,而硬地则代表了来自梅诺卡中新世大陆架不同环境的生物碎屑的混合体。Glycymeris-Crassatella-Clypeaster 组合(A)记录了生活在上斜坡上的部分动物群落,该斜坡周期性地受到上涌水域造成的高能低氧条件的影响。规模较小的动物群(B)的特征是弧藻-心皮藻-瓣鳃藻联合体,来自受风暴和间歇性推动上升流的更持久的西流影响的斜坡中部。这两个动物群的零散性和分离性表明,它们是在这些海底能量机制的驱动下,通过再加工过程混入硬地的。保藏 A 动物群和其他机会类群栖息在这样一种环境中:硬地作为 "岛屿 "暴露在中新世海底,周围是薄层较松散的沉积物。下层石灰岩中的磷酸盐生物碎屑表明,在所研究的露头形成之前,大陆架上的其他地方也形成了类似的矿化床。
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引用次数: 0
Latest Jurassic–Early Cretaceous sedimentary cyclicity and events (Wessex Basin, southern England): A case of pulsed mantle convection? 最近的侏罗纪-早白垩世沉积周期和事件(英格兰南部韦塞克斯盆地):脉冲地幔对流?
IF 1.2 4区 地球科学 Q2 Earth and Planetary Sciences Pub Date : 2024-06-01 DOI: 10.1016/j.pgeola.2024.02.002
Jonathan D. Radley , Robert A. Coram

A pulsed mantle convection model has been proposed for regional third-order cyclic sea level changes during time intervals for which robust evidence of extensive polar ice caps is lacking. One such interval, the Late Jurassic–Early Cretaceous, is represented in southern England's Wessex Basin by the marine to non-marine Portland and Purbeck Limestone groups and Wealden Group and Supergroup. Third-order cyclicity in this succession is reviewed against the mantle convection model, involving pulsed growth of a regional uplift centred on the Cornubian Massif, bordering the Wessex Basin. Supporting evidence for the model is forthcoming from cyclicities in the Upper Tithonian marine Portland Group and Hauterivian–Lower Aptian alluvial–lacustrine–lagoonal Weald Clay Group. Third-order cycles in the Berriasian–Valanginian alluvial–lacustrine Hastings Group reflect fault reactivation along the Anglo-Brabant Massif, conventionally linked to North Atlantic opening. Credible cyclicities on comparable scales have not been identified in the Upper Tithonian–Lower Valanginian lagoonal–lacustrine Purbeck Limestone Group or the Valanginian–Barremian fluvial–alluvial Wessex Formation. Purbeck lithofacies are defined largely by climatic and local tectonic signals; cyclicities were potentially masked in the Wessex Formation by dynamic fluvial processes in proximity to the massif. Certain biotic events in the Purbeck Limestone Group and fluvial events in the Wessex Formation possibly reflect vertical movements of the Cornubian uplift.

有人提出了一个脉冲地幔对流模型,用于解释在缺乏大面积极地冰盖有力证据的时间区间内的区域三阶周期性海平面变化。晚侏罗世-早白垩世就是这样一个时间段,英格兰南部威塞克斯盆地的海洋-非海洋波特兰和普贝克石灰岩群以及瓦尔登群和超群就是其中的代表。根据地幔对流模型对这一演替的三阶周期性进行了审查,其中涉及以威塞克斯盆地边界的康努比山丘为中心的区域隆升的脉冲式增长。上提香海相波特兰组和豪特里维世-下安普世冲积-湖积-泻湖维尔德粘土组的周期性变化为该模型提供了支持性证据。白垩纪-瓦朗基尼冲积-湖积-泻湖维尔德粘土组的三阶周期反映了盎格鲁-布拉班特地块沿线的断层再活化,这通常与北大西洋开口有关。在上提香年-下瓦朗基年泻湖-湖积-普贝克石灰岩组或瓦朗基年-巴里米亚冲积-冲积威塞克斯地层中,尚未发现规模相当的可信周期。普贝克岩性主要是由气候和当地构造信号确定的;威塞克斯地层中的周期性可能被地块附近的动态河流过程所掩盖。普贝克石灰岩组中的某些生物事件和威塞克斯地层中的河流事件可能反映了科努比亚隆起的垂直运动。
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引用次数: 0
Was there a low-altitude Younger Dryas Stadial glacier in south-east Wales? Re-interpretation of landforms and palaeo-climatic inferences 威尔士东南部是否存在低海拔杨干世晚期冰川?重新解释地貌和古气候推断
IF 1.2 4区 地球科学 Q2 Earth and Planetary Sciences Pub Date : 2024-06-01 DOI: 10.1016/j.pgeola.2024.04.003
Richard A. Shakesby , Stephen L. Cornford , John F. Hiemstra

A glacial origin for cirque-like hollows cut into the western escarpment of the Usk valley near Abergavenny, South Wales has become widely accepted. Associated supposed extensive moraine ‘festoons’ have been depicted merging and contemporaneous with Last Glacial Maximum (LGM) deposits formed by ice occupying the adjacent Usk valley. We re-interpret these festoons as the product mainly of rock slope failures (RSFs) emanating from the hollows. A cirque glacier origin is preferred to account for a compact double-ridge feature in one of the hollows. The equilibrium-line altitude (ELA) of the reconstructed glacier (357 m) is > 60 m lower than all similarly small, presumed Younger Dryas Stadial (YDS; c., 12.9–11.7 ka) glaciers elsewhere in South Wales. To test whether this glacier nevertheless might date from the YDS, we apply three approaches to reconstruct annual palaeo-precipitation amounts at the ELA, two based on relationships between accumulation and ablation for modern glaciers and the third on a simple degree-day model (DDM) using likely climatic characteristics for this event. The DDM can be tailored to represent the recognised large-amplitude YDS annual temperature range rather than the much smaller one experienced by modern glaciers, making it our preferred approach. Although conditions along the Usk valley escarpment during the LGM would have been well suited to cirque glacier formation, the DDM approach, using the large-amplitude annual temperature ranges, suggests that a YDS age might also be possible. The results have implications for re-assessing the likely ages of some former small glaciers in South Wales.

在南威尔士阿伯加文尼附近的乌斯克山谷西部悬崖上开凿出的环形凹地的冰川起源已被广泛接受。相关的假定大面积冰碛 "节理 "已被描绘成与邻近乌斯克山谷冰川形成的末次冰川极盛时期(LGM)沉积物相融合、同时发生的现象。我们将这些冰碛带重新解释为主要是来自山谷的岩坡崩塌(RSF)的产物。为了解释其中一个山谷中紧凑的双脊特征,我们倾向于采用盘旋冰川的起源。重建冰川的平衡线海拔(ELA)(357 米)比南威尔士其他地方所有类似的小冰川都要低 60 米。为了测试该冰川是否可以追溯到 YDS 时期,我们采用了三种方法来重建 ELA 的年古降水量,其中两种方法基于现代冰川的积聚与消融之间的关系,第三种方法则基于一个简单的度日模型(DDM),该模型使用了该事件可能的气候特征。DDM 可以根据公认的大振幅 YDS 年温度范围进行调整,而不是现代冰川所经历的小得多的温度范围,因此是我们的首选方法。虽然乌斯克山谷悬崖沿线在全新纪期间的条件非常适合形成盘旋冰川,但使用大振幅年温度范围的 DDM 方法表明,YDS 年龄也是可能的。这些结果对重新评估南威尔士一些前小型冰川的可能年龄具有重要意义。
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引用次数: 0
Scratch circles and circular purported ammoglyphs: Novel observations from the Cape south coast of South Africa 划痕圆圈和圆形所谓的阿摩格文字:南非开普南海岸的新发现
IF 1.2 4区 地球科学 Q2 Earth and Planetary Sciences Pub Date : 2024-06-01 DOI: 10.1016/j.pgeola.2024.03.004
Charles W. Helm , Robin M. Catchpole , Hayley C. Cawthra , Richard M. Cowling , Jan C. De Vynck , Mark G. Dixon , Renée Rust , Willo Stear , Guy H.H. Thesen

Scratch circles, previously defined as ‘bedding plane parallel sedimentary structures formed by the passive rotation of a tethered organism into the surrounding sediment’, have hitherto been identified in the geological record from the Ediacaran to the Paleocene, as well as in modern settings. They have not met the definition of trace fossils, being passively registered by a part of a plant. Several variations of scratch circle morphology have been identified in or on Pliocene and Pleistocene deposits on the Cape south coast of South Africa, allowing for an expansion of the scratch circle temporal range. Furthermore, these novel forms require a redefinition of the term ‘scratch circle’. Anthropogenic origins need to be considered for occurrences in the Pleistocene; guidelines to assist in distinguishing such causes from typical scratch circles are presented. Scratch circles may conceivably have inspired the creation of circular patterns (palaeo-art) in sand by Middle Stone Age hominins. Evaluation of scratch circles in snow allows for subtle features to be determined. A re-evaluation of what the term ‘trace fossil’ constitutes could be considered, in order to include the term palaeo-ichnobotany.

划痕圈以前被定义为 "系留生物被动旋转到周围沉积物中形成的垫层平面平行沉积结构",迄今为止,从埃迪卡拉纪到古新世的地质记录以及现代环境中都发现了划痕圈。它们不符合痕量化石的定义,因为它们是由植物的一部分被动记录下来的。在南非开普南海岸的上新世和更新世沉积物中或沉积物上发现了几种不同形态的划痕圈,从而扩大了划痕圈的时间范围。此外,这些新形式要求重新定义 "划痕圈 "一词。对于更新世出现的划痕圈,需要考虑其人类活动的起源;本文提出了有助于将这些原因与典型划痕圈区分开来的指导原则。可以想象,划痕圈可能是受中石器时代人类在沙地上创造圆形图案(古美术)的启发。对雪地划痕圈的评估可以确定其细微特征。可以考虑重新评估 "痕迹化石 "一词的含义,以便将古生物植物学纳入其中。
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引用次数: 0
The late Quaternary glacial depositional environment at Filey Bay, eastern England: Accretionary mechanisms for thick sequences of tills and stratified diamictons 英格兰东部菲里湾的第四纪晚期冰川沉积环境:厚层堆积物和分层二迭石的堆积机制
IF 1.2 4区 地球科学 Q2 Earth and Planetary Sciences Pub Date : 2024-06-01 DOI: 10.1016/j.pgeola.2024.01.003
David J.A. Evans , David H. Roberts , Emrys Phillips

Construction of the Holderness/Flamborough Head moraine belt on the East Yorkshire coast, England, records the oscillatory onshore flow of the North Sea Lobe of the British-Irish Ice Sheet from ~ 25.8 to ~ 19.7 ka BP, during which time a thick sequence of multiple diamictons and associated stratified sediments were emplaced. The sedimentology of a > 40 m thick stratigraphy through the moraine belt at Filey Bay, in combination with local borehole records, is used here to reconstruct the depositional processes associated with glacier ice moving ca., 12 km onshore and damming the mouth of the Vale of Pickering, which resulted in the accumulation of an unusually thick and complex sequence of deposits traditionally classified as the “Filey till”. The base of the sedimentary sequence comprises stratified diamictons, which are interpreted as glacilacustrine deposits emplaced predominantly by sediment gravity flows in an ice-contact ‘mud apron’ on the distal slope of a subaqueous push ridge constructed in the earliest proglacial lake in Filey Bay; a vertical increase in coarse-grained lithofacies records increasing glacier proximity. Glacier overriding of the mud apron is recorded by a stacked sequence of tills that interdigitate with lake sediments inland. A zone of till-lake sediment interdigitation migrated first westward during North Sea Lobe advance and then eastward during its retreat, into and out of Glacial Lake Pickering, respectively. Multiple tills and intra-till stratified beds and lenses at the top of the sequence at Filey represent alternating deforming bed-sliding bed facies (subglacial traction tills and subglacial canal fills) associated with the construction of inset push moraines, constructed by sub-marginal incremental thickening or punctuated aggradation. This depositional scenario addresses the problems arising from genetic classifications of substantial accumulations of glacigenic diamictons as ‘till’ when modern analogues indicate only modest thicknesses of subglacial traction till beneath glaciers. Onshore thickening of glacigenic deposits through subaqueous push moraine construction and mud apron progradation is compatible with glacier surging behaviour, but not necessarily solely diagnostic of a surging North Sea Lobe during the last glaciation.

英格兰东约克郡海岸霍尔德内斯/弗兰博罗海德冰碛带的建造记录了英-爱尔兰冰盖北海裂片在公元前约 25.8 至约 19.7 千年期间的陆上振荡流动,在此期间形成了厚厚的多层二迭系地层和相关的分层沉积物。通过菲里湾冰碛带厚度大于 40 米地层的沉积学研究,并结合当地的钻孔记录,我们在此重建了与冰川冰层向海岸移动 12 千米并在皮克林谷口筑坝有关的沉积过程,这导致了传统上被归类为 "菲里冰碛 "的异常厚而复杂的沉积序列的堆积。沉积序列的底部由分层的二迭石组成,被解释为主要由沉积物重力流在菲里湾最早的冰期湖泊中形成的水下推挤脊远端斜坡上与冰接触的 "泥坪 "中堆积而成的冰川岩屑沉积物;粗粒度岩性的垂直增加记录了冰川距离的增加。冰川对泥质地表的覆盖表现为与内陆湖泊沉积物相互交错的堆积层。在北海裂片推进过程中,褶曲与湖泊沉积物相互交错的区域首先向西迁移,然后在其后退过程中向东迁移,分别进入和离开皮克林冰川湖。菲里(Filey)地层顶端的多层堆积层和堆积层内分层床及透镜体代表了交替变形的床面-滑动床面(亚冰川牵引堆积层和亚冰川运河填充物),这些床面与嵌入式推移冰碛的建造有关,是由亚边缘增量加厚或点状加层建造而成的。这种沉积情况解决了将大量冰原二迭石堆积物归类为 "冰碛 "的遗传学问题,而现代类似研究表明冰川下的亚冰川牵引冰碛厚度不大。冰原沉积物通过水下推移碛构造和泥坪渐变而在近岸增厚,这与冰川涌动行为是一致的,但并不一定是最后一次冰川期北海裂谷涌动的唯一特征。
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引用次数: 0
IF 1.2 4区 地球科学 Q2 Earth and Planetary Sciences Pub Date : 2024-06-01 DOI: 10.1016/j.pgeola.2024.03.005
Dmitry A. Ruban
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引用次数: 0
A ‘giant’ pterodactyloid pterosaur from the British Jurassic 英国侏罗纪的 "巨型 "翼龙
IF 1.2 4区 地球科学 Q2 Earth and Planetary Sciences Pub Date : 2024-06-01 DOI: 10.1016/j.pgeola.2024.05.002
James L. Etienne , Roy E. Smith , David M. Unwin , Robert S.H. Smyth , David M. Martill

The fossil remains of a pterodactyloid pterosaur from the Kimmeridge Clay Formation (Jurassic: Tithonian) of Abingdon, Oxfordshire, central England are identified as a partial left first wing finger phalanx. The elongation of the phalanx and distinctive morphology of the proximal articular region, in particular the square outline of the extensor tendon process, permit the specimen to be assigned to Ctenochasmatoidea. Although fragmentary, it is sufficiently well preserved to determine accurately its dimensions when complete. Morphometric analysis reveals the specimen to represent one of the largest known examples of a Jurassic pterosaur, with an estimated wingspan of at least 3 m, and is one of the first pterodactyloids to be reported from the Jurassic of the United Kingdom.

来自英格兰中部牛津郡阿宾顿金默里奇粘土层(侏罗纪:提顿时期)的翼龙化石遗骸被鉴定为左侧第一翼指的部分趾骨。指骨的伸长和近端关节区的独特形态,特别是伸肌腱突的方形轮廓,使该标本被归类为栉水母纲(Ctenochasmatoidea)。该标本虽然残缺不全,但保存完好,可以准确确定完整标本的尺寸。形态分析表明,该标本是已知最大的侏罗纪翼龙之一,翼展估计至少有 3 米,是英国侏罗纪最早报道的翼龙之一。
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引用次数: 0
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