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Hand and Foot Musculature of Anura: Structure, Homology, Terminology, and Synapomorphies for Major Clades Anura的手和脚肌肉:主要分支的结构、同源性、术语和同源性
IF 3.4 2区 环境科学与生态学 Q1 Agricultural and Biological Sciences Pub Date : 2020-11-06 DOI: 10.1206/0003-0090.443.1.1
Boris L. Blotto, M. Pereyra, Taran Grant, J. Faivovich
ABSTRACT Although studies of anuran hand and foot musculature began in the first half of the 19th century, all studies to date have been taxonomically or anatomically restricted in scope, and none has considered the diversity of autopodial myology in Anura as a whole. As a model for future comparisons, we thoroughly describe the hand and foot musculature of an arboreal species (the hylid Triprion petasatus), define the layers in which these muscles are arranged, and attribute presumed functions. On the basis of our myological analysis of 155 species representing 46 of the 54 currently recognized families and main clades of anurans, we describe 20 characters related to hand and foot muscles. Optimization of these characters on the most recent and inclusive phylogenetic hypothesis of Anura results in synapomorphies for several major clades (Bombinatoridae, Alytidae, Xenoanura + Acosmanura, Xenoanura, Pipidae, Acosmanura, Anomocoela, Scaphiopodidae, Pelodytidae + Pelobatidae + Megophryidae, Megophryidae, Neobatrachia, Heleophrynidae, Sooglossidae, Laurentobatrachia, Calyptocephalellidae, Myobatrachoidea, and Nobleobatrachia), including new, nonhomoplastic synapomorphies for clades previously supported only by molecular evidence and a few conflicting phenotypic characters (e.g., Acosmanura, Anomocoela, Neobatrachia). Additionally, we (1) address controversies regarding the homology of anuran and caudate muscles in the context of putative synapomorphies for Ascaphidae + Leiopelmatidae and its sister clade Lalagobatrachia; (2) evaluate a recently proposed terminology for anuran hand and foot musculature; (3) discuss the identities of several hand and foot muscles with problematic homologies; (4) establish a unified terminology for anuran hand and foot muscles, including a list of synonyms for all names employed in the literature; and (5) propose hypotheses for the origin of several myological novelties (neomorphs).
尽管对无尾动物手足肌肉组织的研究始于19世纪上半叶,但迄今为止所有研究的范围都受到分类学或解剖学的限制,而且没有一个研究将无尾动物自掌肌学的多样性作为一个整体加以考虑。作为未来比较的模型,我们详细描述了一种树栖物种(水螅Triprion petasatus)的手和脚肌肉组织,定义了这些肌肉排列的层,并赋予了假定的功能。在我们对目前已知的54科和主要分支中的46科155种的肌肉学分析的基础上,我们描述了20个与手和脚肌肉有关的特征。在最新的无尾动物系统发育假说上对这些特征进行优化,得到了几个主要分支(弹蝇科、Alytidae、Xenoanura + Acosmanura、Xenoanura、Pipidae、Acosmanura、Anomocoela、Scaphiopodidae、Pelodytidae + Pelobatidae + Megophryidae、Megophryidae、Neobatrachia、heleophridae、Sooglossidae、Laurentobatrachia、Calyptocephalellidae、Myobatrachoidea和Nobleobatrachia)的synapomorphies,包括新的、进化枝的非同质突触形态以前仅由分子证据和一些相互冲突的表型特征(例如,Acosmanura, Anomocoela, Neobatrachia)支持。此外,我们(1)解决了关于尾尾肌和尾尾肌同源性的争议,在假定的尾尾肌+ Leiopelmatidae及其姐妹分支Lalagobatrachia的突触形态的背景下;(2)评估最近提出的关于anuran手足肌肉组织的术语;(3)讨论了几个有问题的同系性的手足肌肉的身份;(4)为anuran手足肌肉建立统一的术语,包括文献中使用的所有名称的同义词列表;(5)对几种新形态(新形态)的起源提出假说。
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引用次数: 8
Systematic Revision of the Asian Forest Scorpions (Heterometrinae Simon, 1879), Revised Suprageneric Classification of Scorpionidae Latreille, 1802, and Revalidation of Rugodentidae Bastawade et al., 2005 《亚洲林蝎的系统修订》(Heterometrinae Simon,1879),《修订的蝎科超基因组分类》,1802年,《Rugodentidae Bastawade等人的再验证》,2005年
IF 3.4 2区 环境科学与生态学 Q1 Agricultural and Biological Sciences Pub Date : 2020-10-14 DOI: 10.1206/0003-0090.442.1.1
L. Prendini, Stephanie F. Loria
ABSTRACT The genera and species of the Asian forest scorpions (Scorpionidae Latreille, 1802) are revised based on a phylogenetic analysis of 186 morphological characters and 4188 base pairs of concatenated DNA sequence from three mitochondrial loci and two nuclear loci. Revision of the Asian scorpionids required a critical reappraisal of the suprageneric classification of Scorpionidae, on the basis of which the monotypic Indian scorpionoid genus, Rugodentus Bastawade et al., 2005, stat. rev., and its type species, Rugodentus keralaensis Bastawade et al., 2005, stat. rev., are revalidated, and subfamily Rugodentinae Bastawade et al., 2005, revalidated and elevated to the rank of family, Rugodentidae Bastawade et al., 2005, stat. nov. et stat. rev.; Heterometrinae Simon, 1879, stat. nov., and Opistophthalminae Rossi, 2016, stat. nov., are elevated to the rank of subfamily; Pandinopsis Vachon, 1974, stat. nov., and Pandipalpus Rossi, 2015, stat. nov., are elevated to the rank of genus, resulting in two new combinations: Pandinopsis dictator (Pocock, 1888), comb. nov., and Pandipalpus viatoris (Pocock, 1890), comb. nov.; and 10 new synonyms are presented: Pandinopsini Rossi, 2016 = Pandininae Thorell, 1876, syn. nov.; Protophthalmini Rossi, 2016 = Opistophthalminae Rossi, 2016, syn. nov.; Protophthalmus Lawrence, 1969 = Opistophthalmus C.L. Koch, 1837, syn. nov.; Pandinoides (Dunlopandinoides) Rossi, 2016 = Pandinoides Fet, 2000, syn. nov.; Pandinurus (Pandicaporiaccous) Rossi, 2015 = Pandiborellius Rossi, 2015, syn. nov.; Buthus defensor C.L. Koch, 1837 = Pandinurus gregoryi (Pocock, 1896), syn. nov.; Buthus heros C.L. Koch, 1837 = Pandinurus exitialis (Pocock, 1888), syn. nov.; Pandinus lowei Kovařík, 2012 = Pandipalpus viatoris (Pocock, 1890), syn. nov.; Pandinurus (Pandipalpus) pygmaeus Rossi, 2015 = Pandipalpus viatoris (Pocock, 1890), syn. nov.; Pandinus intermedius Borelli, 1919 = Pandinurus citernii (Borelli, 1919), syn. nov. The following revisions are implemented to the classification of the Asian forest scorpions (Heterometrinae). Three former subgenera of Heterometrus Ehrenberg, 1828 are revalidated and elevated to the rank of genus: Chersonesometrus Couzijn, 1978, stat. nov. et stat. rev.; Javanimetrus Couzijn, 1981, stat. nov. et stat. rev.; and Srilankametrus Couzijn, 1981, stat. nov. et stat. rev. One subgenus is elevated to the rank of genus: Gigantometrus Couzijn, 1978, stat. nov. Two new genera and eight new species are described: Deccanometrus, gen. nov.; Sahyadrimetrus, gen. nov.; Chersonesometrus bastawadei, sp. nov.; Chersonesometrus hendersoni, sp. nov.; Chersonesometrus nathanorum, sp. nov.; Chersonesometrus shivashankari, sp. nov.; Sahyadrimetrus mathewi, gen. et sp. nov.; Sahyadrimetrus tikaderi, gen. et sp. nov.; Srilankametrus couzijni, sp. nov.; Srilankametrus pococki, sp. nov. Heterometrus sensu stricto is restricted to eight species of the nominotypical subgenus, all other species, formerly placed in Heterometrus, are
;Palamnaeus liophysa Thorell,1888=长柄异齿龙(Herbst,1800)十一月Palamnaeus oatesii Pocock,1900=彼得西异齿龙(Thorell,1876)十一月Palamnaeus swammerdami flavimanus Pocock,1900=斯瓦默达米巨人(西蒙,1872)十一月马多氏异花介Kopstein,1921=蓝异花介(Thorell,1876),syn。十一月laevifrons Roewer,1943=蓝异花介(Thorell,1876)。十一月异Metrus(Chersonesometrus)granularanus Couzijn,1981=Srilantamus caesar(C.L.Koch,1841),syn。十一月Heterometrus(Heteromerus)liophysa separatus Couzijn,1981=Heterometrous glaucus(Thorell,1876),syn。十一月Heterometrus(Heteromerus)liophysa spartanicus Couzijn,1981=Heterometrous glaucus(Thorell,1876),syn。十一月bengkalitensis Couzijn,1981=长柄异头虫(Heterometrus)(Autumn,1800),syn。十一月异节病毒(异节病毒)longimanus marmoratus Couzijn,1981=异节病毒longimanu(Autumn,1800),syn。十一月Heterometrus(Heteromerus)petersii mindanensis Couzijn,1981=silenus Heterometrous(Simon,1884),syn。十一月异刺病毒(Heterometrus)spinifer solitarius Couzijn,1981=异刺病毒。十一月Heterometrus(Srilantamus)indus laevitensus Couzijn,1981=Srilantatus indus(DeGeer,1778),syn。十一月keralaensis Tikader和Bastawade,1983年=rugosus(Couzijn,1981),syn。十一月Heterometrus cimrmani Kovařík,2004=laevigatus(Thorell,1876)十一月mysorensis Kovařík,2004=Chersonesometrus tristis(Henderson,1919)十一月尼泊尔异核病毒Kovařík,2004=孟加拉异核病毒(Pocock,1900)十一月rolciki Kovařík,2004年=Sahyadrimetrus scaber(Thorell,1876)十一月Heterometrous sejnai Kovařík,2004=Javanimetrus cyaneus(C.L.Koch,1836)十一月tibetanus Lourenço等,2005=孟加拉Deccanometrus(Pocock,1900),syn。十一月liangi Heterometrus Zhu和Yang,2007=silenus Heteromerus(Simon,1884)十一月telanganaensis Javed等人,2010=Deccanometrus xanthopus(Pocock,1897),syn。十一月Atascorpius Mirza等人,2012=Chersonesometrus beccaloniae(Kovařík,2004)十一月牛头怪异头龙Plíšková等人,2016=laevigatus异头龙(Thorell,1876)十一月巴斯塔瓦代·罗西异Metrus bastawadei Rossi,2016=Rugodentus keralaensis Bastawade et al.,2005,syn。nov.在异子宫亚科共51个同义词中,又确认了前人的25个同义词。在适用的情况下,提供了带有对比图像的修订诊断、属和物种的关键分布图谱,以及关于其生态和保护状况的可用数据摘要。
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引用次数: 11
Large Scaphitid Ammonites (Hoploscaphites) from the Upper Cretaceous (Upper Campanian–Lower Maastrichtian) of North America: Endless Variation on a Single Theme 北美洲上白垩纪(上坎潘阶-下马斯特里赫特阶)的大型Scaphitid Ammonites(Hoploscaphites):单一主题的无尽变化
IF 3.4 2区 环境科学与生态学 Q1 Agricultural and Biological Sciences Pub Date : 2020-09-09 DOI: 10.1206/0003-0090.441.1.1
N. Landman, W. Kennedy, Joyce C. Grier, Neal L. Larson, J. Grier, Tom Linn, L. Tackett, B. Jicha
ABSTRACT We describe three species of large scaphitid ammonites (Ammonoidea: Ancyloceratina) from the Upper Cretaceous (upper Campanian–lower Maastrichtian) of the Western Interior of North America. Each species occurs as two dimorphs, referred to as macroconch and microconch. All three species share a similar pattern of ornamentation consisting of long, thin, nonbifurcating ribs on the adoral part of the phragmocone, suggesting that they constitute a single monophyletic clade. Macroconchs of Hoploscaphites crassus (Coryell and Salmon, 1934) are characterized by a globose whorl section, with closely spaced ventrolateral tubercles on the body chamber, usually persisting to the aperture. Macroconchs of Hoploscaphites plenus (Meek and Hayden, 1860) differ from those of H. crassus in having a more subquadrate whorl section with flatter flanks, and fewer, larger, and more widely spaced ventrolateral tubercles. Macroconchs of Hoploscaphites peterseni, n. sp., closely resemble those of H. crassus, but differ in being nearly circular in side view with a more compressed whorl section. All three species lived at approximately the same time in the same general area and depositional environment. They are abundant in the Baculites baculus Zone but also occasionally occur in the B. eliasi Zone and possibly lower part of the B. grandis Zone. They are present in the Pierre Shale of east-central Montana and east-central Wyoming, the Lewis Shale of south-central Wyoming, and the Bearpaw Shale of northeast Montana. It is possible that these three species represent subspecies within a single species or a “flock” of very closely related species, similar to the “species flocks” observed in modern cichlid fishes.
摘要:我们描述了北美洲西部内陆上白垩纪(上坎潘阶-下马斯特里赫特阶)的三种大型舟状菊石(菊目:钩角亚目)。每个物种都以两个二形态出现,被称为macroconch和microconch。这三个物种都有着相似的纹饰模式,由膈腔朝拜部分的细长非分叉肋骨组成,这表明它们构成了一个单一的单系分支。粗腹蛙的大型海螺(Coryell和Salmon,1934)的特征是球形轮生部分,在体腔上有紧密间隔的腹外侧结节,通常持续到孔。Hoploscaphites plenus的大海螺(Meek和Hayden,1860)与粗海螺的不同之处在于,它们的轮生截面更接近方形,侧面更平坦,腹外侧结节更少、更大、间隔更宽。Hoploscaphites peterseni,n.sp.的大型海螺与粗糙海螺非常相似,但不同之处在于侧视时几乎是圆形的,轮生部分更为压缩。这三个物种几乎同时生活在相同的一般区域和沉积环境中。它们在Baculites baculus带丰富,但偶尔也会出现在B.eliasi带,可能是B.grandis带的下部。它们存在于蒙大拿州中东部和怀俄明州中东部的皮埃尔页岩、怀俄明州中南部的刘易斯页岩和蒙大拿州东北部的熊掌页岩中。这三个物种可能代表一个物种中的亚种,或是亲缘关系非常密切的物种的“群体”,类似于在现代慈鲷中观察到的“物种群体”。
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引用次数: 1
A Revision of the Didelphid Marsupial Genus MarmosaPart 2. Species of the Rapposa Group (Subgenus Micoureus) 迪德尔皮德有袋动物属Marmosa的修订(二)。Rapposa群的物种(Micoureus亚属)
IF 3.4 2区 环境科学与生态学 Q1 Agricultural and Biological Sciences Pub Date : 2020-06-01 DOI: 10.1206/0003-0090.439.1.1
R. Voss, Thomas C. Giarla, J. Díaz-Nieto, S. Jansa
ABSTRACT In this report, the second of a revisionary series on mouse opossums (Marmosa), we analyze cytochrome b sequence data from 166 specimens of the subgenus Micoureus and delimit putative species using the multirate Poisson Tree Processes (mPTP) method. That analysis identifies 21 putative species, many of which can be matched with available names, including alstoni, constantiae, demerarae, limae, germana, meridae, paraguayana, parda, perplexa, phaea, rapposa, and rutteri. However, some of these nominal taxa are not morphologically diagnosable, and in the absence of other corroborating evidence, we do not recommend that they all be recognized as valid. Phylogenetic analyses of a multigene dataset suggest that putative species of Micoureus belong to several well-supported clades, one of which (the “Rapposa Group”) is revised in this report. As defined herein, the Rapposa Group includes at least three valid species: M. rapposa Thomas, 1899 (including budini Thomas, 1920); M. parda Tate, 1931; and M. rutteri Thomas, 1924. Herein we document their ecogeographic distributions and diagnostic traits, comment on their taxonomic histories, and list the specimens we examined (including all relevant type material).
摘要在本报告中,我们分析了166个鼠尾亚属标本的细胞色素b序列数据,并使用多比率泊松树过程(mPTP)方法确定了推测的物种。该分析确定了21个推定物种,其中许多可以与现有名称相匹配,包括alstoni、constantiae、demaraae、limae、germana、meridae、paraguayana、parda、confusa、phaea、rapposa和rutteri。然而,其中一些命名分类群在形态学上是不可诊断的,在缺乏其他确凿证据的情况下,我们不建议将它们全部认定为有效。多基因数据集的系统发育分析表明,Micoureus的假定物种属于几个得到充分支持的分支,其中一个分支(“Rapposa群”)在本报告中进行了修订。如本文所定义的,Rapposa群包括至少三个有效物种:M.Rapposa-Thomas,1899(包括布迪尼·托马斯,1920);M.帕达·泰特,1931年;和M.rutteri Thomas,1924年。在此,我们记录了它们的生态地理分布和诊断特征,评论了它们的分类历史,并列出了我们检查的标本(包括所有相关的类型材料)。
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引用次数: 24
Taxonomy and Phylogenetics of Nanometinae and Other Australasian Orb-Weaving Spiders (Araneae: Tetragnathidae) 纳米蛛科及其他澳大利亚圆织蜘蛛的分类与系统发育(蜘蛛目:蛛科)
IF 3.4 2区 环境科学与生态学 Q1 Agricultural and Biological Sciences Pub Date : 2020-02-28 DOI: 10.1206/0003-0090.438.1.1
Fernando Álvarez-Padilla, R. Kallal, G. Hormiga
ABSTRACT The spider family Tetragnathidae Menge is a cosmopolitan, relatively well-studied spider clade with some members readily identifiable by their elongate chelicerae and/or their horizontal orb webs. It has four recognized subfamilies—Tetragnathinae, Metainae, Leucauginae, and the Australasian endemic Nanometinae—although many genera remain unassigned to subfamilial groups. Nanometinae alpha taxonomy is the least well understood of these lineages despite the inclusion of members of the subfamily in a number of phylogenetic analyses over the past decade. Here we describe 10 new species and revise seven additional tetragnathids from Australia, New Zealand, New Caledonia, and Papua New Guinea in the genera Nanometa, Taraire, gen. nov., Tawhai, gen. nov., Harlanethis, gen. nov., and Iamarra, gen. nov. These 17 species are: Nanometa gentilis Simon, 1908, N. trivittata (Keyserling, 1887), comb. nov., N. sarasini (Berland, 1924), comb. nov., N. lagenifera (Urquhart, 1888), comb. nov., N. purpurapunctata (Urquhart, 1889), comb. nov., N. fea, sp. nov., N. tasmaniensis, sp. nov., N. tetracaena, sp. nov., N. dimitrovi, sp. nov., N. dutrorum, sp. nov., N. forsteri, sp. nov., Taraire rufolineata (Urquhart, 1889), comb. nov., Taraire oculta, sp. nov., Tawhai arborea (Urquhart, 1891), comb. nov., Harlanethis lipscombae, sp. nov., H. weintrauborum, sp. nov., and Iamarra multitheca, sp. nov. We also synonymize Nediphya Marusik and Omelko, 2017, and the monotypic genus Eryciniolia Strand, 1912, with Nanometa, bringing the total number of species in the genus from one to 14. Using an expanded taxon sampling for prior studies based on six molecular markers—12S rRNA, 16S rRNA, 18S rRNA, 28S rRNA, cytochrome c oxidase subunit I, and histone H3—and both maximum likelihood and Bayesian methods, we place these taxa in the tetragnathid tree of life. Nanometinae and its constituent genera Nanometa and Pinkfloydia are reciprocally monophyletic. Harlanethis belongs to Leucauginae. The genera Taraire, Tawhai, and Iamarra defy robust phylogenetic placement and are not yet assigned to subfamily.
摘要:Menge四颚蛛科是一个世界性的、研究相对深入的蜘蛛分支,其一些成员很容易通过其细长的螯叶和/或水平的球形网来识别。它有四个公认的亚科——四颚亚科、Metainae亚科、Leucauginae亚科和澳大拉西亚特有的Nanometinae亚科——尽管许多属仍不属于亚科。尽管在过去十年的一些系统发育分析中包含了该亚科的成员,但纳米甲亚科α分类学是这些谱系中最不为人所知的。在这里,我们描述了来自澳大利亚、新西兰、新喀里多尼亚和巴布亚新几内亚的10个新物种,并修订了另外7个四颚虫属,分别为Nanometa属、Taraire属、Tawhai属、Harlanethis属和Iamarra属。这17个物种是:Nanometa gentilis Simon,1908,N.trivitta(Keyserling,1887),comb。nov.,N.sarasini(Berland,1924),梳。N.lagenifera(厄克特出版社,1888年),科姆。nov.,N.purpurapunctata(厄克特,1889),梳。nov.,N.fea,sp.nov.,N.tasmaniensis,sp.nova.,N.tetraaena,sp.nv.,N.dimitrovi,sp.nov,N.dutrorum,sp-nov.,N.forsteri,sp.nov.,Taraire rufolineta(Urquhart,1889),梳。nov.,Taraire oculta,sp.nov.,Tawhai arborea(Urquhart,1891),梳。nov.,Harlanethis lipscombae,sp.nov.,H.weintrauborum,sp.nova.和Iamarra multitheca,sp.nov。我们还将Nedifya Marusik和Omelko,2017,以及单型属Eryciniolia Strand,1912与Nanometa同义,使该属的物种总数从1种增加到14种。使用基于六个分子标记(12S rRNA、16S rNA、18S rRNA和28S rRNA,细胞色素c氧化酶亚基I和组蛋白H3)以及最大似然和贝叶斯方法的扩展分类单元抽样进行先前的研究,我们将这些分类单元放在四颚生命树中。Nanometinae属及其组成属Nanometa属和Pinkfloydia属是相互单系的。Harlanethis属于Leucauginae。Taraire属、Tawhai属和Iamarra属的系统发育位置不稳定,尚未被归入亚科。
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引用次数: 8
Methods of studying early theropod flight 研究早期兽脚亚目恐龙飞行的方法
IF 3.4 2区 环境科学与生态学 Q1 Agricultural and Biological Sciences Pub Date : 2020-01-01 DOI: 10.1206/0003-0090.440.1.1
Pittman, Ashley M. Heers, Francisco Serrano, D. Field, M. Habib, T. Dececchi, Thomas G. Kaye, H. Larsson
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引用次数: 9
Reassessment of a Historical Collection of Sauropod Dinosaurs from the Northern Morrison Formation of Wyoming, with Implications for Sauropod Biogeography 对怀俄明州莫里森组北部蜥脚类恐龙历史标本的重新评估及其对蜥脚类生物地理学的影响
IF 3.4 2区 环境科学与生态学 Q1 Agricultural and Biological Sciences Pub Date : 2019-11-04 DOI: 10.1206/0003-0090.437.1.1
E. Tschopp, S. Maidment, M. Lamanna, M. Norell
ABSTRACT The Upper Jurassic Morrison Formation of the western United States preserves one of the best-known Mesozoic paleoecosystems worldwide. The formation crops out over an area from New Mexico and Oklahoma to Montana and Utah and encompasses a time span of approximately eight million years. Recent studies indicate a high diversity of gigantic, herbivorous sauropod dinosaurs, but the geographic and temporal distributions of species or even genera of these animals remain poorly understood. In particular, sauropod specimens from northern outcrops of the formation have rarely been studied in detail, and temporal relationships among sites are imprecise. Here, we reassess the taxonomic diversity of the sauropods from a historic Carnegie Museum locality in northern Wyoming. Previous referrals of material to the well-known diplodocid genera Apatosaurus and Diplodocus cannot be confidently confirmed; instead, all these specimens more likely represent elements from the recently recognized Galeamopus. Specimens previously assigned to Camarasaurus and Haplocanthosaurus could not be referred to these genera based on apomorphies, due to a lack of detailed knowledge concerning the genus- and species-level taxonomy of these sauropods. Our findings imply that many referrals of incomplete diplodocid skeletons to Apatosaurus and Diplodocus must be reassessed. These reassessments are particularly important with regard to specimens from northern localities of the Morrison Formation, as it is becoming increasingly evident that diplodocids from this area were distinct from better-known, more southerly taxa. This geographic segregation does not seem to apply to nondiplodocid sauropods; however, these taxa are also in need of systematic revision, which may reveal species-level patterns similar to those observed in Diplodocidae.
摘要美国西部的上侏罗纪Morrison组保存着世界上最著名的中生代古生态系统之一。该地层分布在从新墨西哥州、俄克拉何马州到蒙大拿州和犹他州的一个地区,时间跨度约为800万年。最近的研究表明,巨型草食性蜥脚类恐龙的多样性很高,但对这些动物的物种甚至属的地理和时间分布仍知之甚少。特别是,来自该地层北部露头的蜥脚类恐龙标本很少被详细研究,各地点之间的时间关系也不精确。在这里,我们重新评估了怀俄明州北部卡内基博物馆历史悠久的蜥脚类动物的分类多样性。以前将材料转介给著名的双足龙属Apatosaurus和Diplodocus的说法无法得到可靠的证实;相反,所有这些标本更可能代表了最近被认可的Galeamopus的元素。由于缺乏关于这些蜥脚类动物属和种级分类学的详细知识,以前被分配给卡马拉龙和单角龙的标本无法根据变形化石被称为这些属。我们的发现表明,许多不完整的双足类骨骼被转介给阿帕龙和双足类的情况必须重新评估。这些重新评估对Morrison组北部地区的标本尤为重要,因为越来越明显的是,该地区的双足类与更为知名、更为南方的分类群不同。这种地理隔离似乎不适用于非双足类蜥脚类;然而,这些分类群也需要系统的修订,这可能会揭示出与Diplodocidae中观察到的相似的物种水平模式。
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引用次数: 5
Catalog of the Staphylinidae (Insecta: Coleoptera). 1758 to the End of the Second Millennium.V. Staphylinine Group (Part 2)Staphylininae: Diochini, Maorothiini, Othiini, Platyprosopini, Staphylinini (Amblyopinina, Anisolinina, Hyptiomina, Philonthina) Staphylinidae目录(昆虫纲:鞘翅目)。1758年至第二个千年末。V.葡萄球菌群(下)葡萄球菌科:Diochini,Maorothiini,Othiini,Platyprosopini,葡萄球菌(Amblyopina,Anisolina,Hyptiomina,Philonthina)
IF 3.4 2区 环境科学与生态学 Q1 Agricultural and Biological Sciences Pub Date : 2019-07-18 DOI: 10.1206/0003-0090.265.1.5
L. Herman
This catalog (published in seven parts, all released on the same day) is based on only the published literature for the Staphylinidae. Of the 32 subfamilies, the following 28 are included herein: Apateticinae, Dasycerinae, Empelinae, Euaesthetinae, Glypholomatinae, Habrocerinae, Leptotyphlinae, Megalopsidiinae, Micropeplinae, Microsilphinae, Neophoninae, Olisthaerinae, Omaliinae, Osoriinae, Oxyporinae, Oxytelinae, Phloeocharinae, Piestinae, Protactinae†, Proteininae, Protopselaphinae, Pseudopsinae, Solieriinae, Staphylininae, Steninae, Tachyporinae, Trichophyinae, and Trigonurinae. The Aleocharinae, Paederinae, Pselphinae, and Scaphidiinae are excluded from this edition of the catalog. References to the original citation or description are given for available family-group, genus-group, and species-group names of both extant and extinct forms. The type genus is cited for each family-group name, the type species for each genus-group name, and the type locality for each species-group name. Where appropriate, all subgenera, subspecies, or synonyms are listed for each valid name. Annotated subsequent references are presented for all names. Distributional summaries are given for each valid taxon. Full bibliographic citations are in Part VII. A short historical review, coauthored with Aleš Smetana, follows the Introduction (Part I), with the main focus on biographical sketches that include many photographs. The goal of this catalog is to summarize the current state of knowledge of the family and to stimulate worldwide monographic studies. iv Address correspondence to: Lee H. Herman, Curator, Division of Invertebrates, American Museum of Natural History, Central Park West at 79th Street, New York, NY 10024-5192, USA (herman@amnh.org). Please send comments, corrections, or updates to either the postal or e-mail address. Reprints of new publications would be helpful and appreciated.
本目录(分七部分出版,均在同一天发布)仅基于已发表的葡萄球菌科文献。在32个亚科中,以下28个亚科包括在本文中:Apateticinae、Dassycerinae、Empelinae、Euaesthetinae、Glypolomatinae、Habrocerinae、Leptotyphlinae、Megalopidinae、Microeplinae、Microsilphinae、Neophoninae、Olistherinae、Omaliinae、Osoriinae、Oxyponinae、Oxytinae、Phloeocharinae、Piestinae、Protactinae†、Proteininae、Protopslapinae、Pseudopsinae、Solierinae、Staphylininae、,Steninae、Tachyporinae、Trichophyinae和Trigonurinae。Aleocharinae、Paederinae、Pselphinae和Scaphidiinae被排除在本版目录之外。原始引文或描述的参考文献是现有和已灭绝形式的科群、属群和种群名称。每个科群名称都引用模式属,每个属群名称引用模式种,每个物种群名称引用类型位置。在适当的情况下,列出每个有效名称的所有亚属、亚种或同义词。所有名称都附有注释的后续参考文献。给出了每个有效分类单元的分布摘要。完整的书目引文见第七部分。在引言(第一部分)之后,与AlešSmetana合著了一篇简短的历史综述,主要关注传记草图,其中包括许多照片。该目录的目标是总结家庭知识的现状,并促进世界范围内的专题研究。iv通讯地址:Lee H.Herman,美国自然历史博物馆无脊椎动物部馆长,美国纽约州纽约市第79街中央公园西10024-5192(herman@amnh.org)。请将评论、更正或更新发送到邮寄或电子邮件地址。重印新出版物将是有益的,并表示赞赏。
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引用次数: 7
Stratigraphy and Paleobiology of the Upper Cretaceous-Lower Paleogene Sediments from the Trans-Saharan Seaway in Mali 马里跨撒哈拉航道上白垩纪-下古近系沉积物的地层学和古生物学
IF 3.4 2区 环境科学与生态学 Q1 Agricultural and Biological Sciences Pub Date : 2019-06-28 DOI: 10.1206/0003-0090.436.1.1
M. O'Leary, M. Bouaré, Kerin M. Claeson, Kelly Heilbronn, Robert V. Hill, J. A. Mccartney, J. Sessa, F. Sissoko, L. Tapanila, E. Wheeler, E. Roberts
An epicontinental sea bisected West Africa periodically from the Late Cretaceous to the early Eocene, in dramatic contrast to the current Sahara Desert that dominates the same region today. Known as the Trans-Saharan Seaway, this warm and shallow ocean was a manifestation of globally elevated sea level associated with the rapid break-up of the supercontinent Gondwana in the late Mesozoic. Although it varied in size through time, the Trans-Saharan Seaway is estimated to have covered as much as 3000 km2 of the African continent and was approximately 50 m deep. The edges of the sea were defined in part by the high topography of the Precambrian cratons and mobile belts of West Africa. Over its approximately 50 million year episodic existence, through six major periods of transgression and regression, the Trans-Saharan Seaway left behind extensive nearshore marine sedimentary strata with abundant fossils. The waters that yielded these deposits supported and preserved the remains of numerous vertebrate, invertebrate, plant, and microbial species that are now extinct. These species document a regional picture of ancient tropical life that spanned two major Earth events: the Cretaceous-Paleogene (K-Pg) boundary and the Paleocene-Eocene Thermal Maximum (PETM). Whereas extensive epeiric seas flooded the interior portions of most continents during these intervals, the emerging multicontinental narrative has often overlooked the Trans-Saharan Seaway, in part because fundamental research, including the naming of geological formations and the primary description and analysis of fossil species, remained to be done. We provide such synthesis here based on two decades of fieldwork and analyses of sedimentary deposits in the Republic of Mali. Northern parts of the Republic of Mali today include some of the farthest inland reaches of the ancient sea. We bring together and expand on our prior geological and paleontological publications and provide new information on ancient sedimentary rocks and fossils that document paleoequatorial life of the past. Ours is the first formal description of and nomenclature for the Upper Cretaceous and lower Paleogene geological formations of this region and we tie these names to regional correlations over multiple modern territorial boundaries. The ancient seaway left intriguing and previously unclassified phosphate deposits that, quite possibly, represent the most extensive vertebrate macrofossil bone beds known from anywhere on Earth. These bone beds, and the paper shales and carbonates associated with them, have preserved a diverse assemblage of fossils, including a variety of new species of invertebrates and vertebrates, rare mammals, and trace fossils. The shallow marine waters included a wide range of paleoenvironments from delta systems, to hypersaline embayments, protected lagoons, and carbonate shoals. Our overarching goal has been to collect vertebrate fossils tied to a K-Pg stratigraphic section in Africa. We provid
从白垩纪晚期到始新世早期,一个陆表海周期性地将西非一分为二,与今天统治同一地区的撒哈拉沙漠形成鲜明对比。这个温暖的浅海被称为跨撒哈拉海道,是全球海平面上升的表现,与中生代晚期冈瓦纳超大陆的快速分裂有关。尽管随着时间的推移,它的大小有所不同,但据估计,撒哈拉沙漠外的海道覆盖了非洲大陆3000平方公里的面积,深度约为50米。海的边缘部分是由前寒武纪克拉通的高地形和西非的活动带确定的。在其大约5000万年的幕式存在中,经历了六个主要的海侵和退行期,跨撒哈拉海道留下了广泛的近岸海相沉积地层和丰富的化石。产生这些沉积物的水域支持并保存了许多现已灭绝的脊椎动物、无脊椎动物、植物和微生物物种的遗骸。这些物种记录了古代热带生命的区域图景,跨越了两个主要的地球事件:白垩纪-古近纪(K-Pg)边界和古新世-始新世热最大值(PETM)。尽管在这段时间内,大部分大陆的内陆地区都被广阔的外海淹没,但新兴的多大陆叙事往往忽略了跨撒哈拉海道,部分原因是基础研究,包括地质构造的命名和化石物种的初步描述和分析,仍有待完成。我们在此根据二十年来对马里共和国沉积矿床的实地考察和分析提供这样的综合。今天马里共和国的北部地区包括古海的一些最远的内陆地区。我们汇集并扩展了我们之前的地质和古生物学出版物,并提供了关于古代沉积岩和化石的新信息,这些信息记录了过去的古赤道生活。我们的研究是对该地区上白垩纪和下古近纪地质构造的第一个正式描述和命名,我们将这些名称与多个现代领土边界的区域相关性联系起来。古老的海道留下了有趣的、以前未分类的磷酸盐沉积物,很可能代表了地球上已知的最广泛的脊椎动物大化石骨床。这些骨床,以及与之相关的纸页岩和碳酸盐,保存了各种各样的化石组合,包括各种无脊椎动物和脊椎动物的新种,稀有哺乳动物和痕迹化石。浅海水域包括从三角洲系统到高盐河口、受保护的泻湖和碳酸盐浅滩的各种古环境。我们的首要目标是收集与非洲K-Pg地层剖面相关的脊椎动物化石。我们提供了这样一个剖面,与先前的观点一致,表明马里岩石在古新世早期存在沉积间隙,西非其他地方也提出了这种不整合。我们对跨越K-Pg边界的几种脊椎动物进化枝的系统发育分析已经阐明了多个分类群的进化枝-进化枝物种水平的生存和范围扩展。根据目前的证据,很少有跨撒哈拉海道的大型化石物种在K-Pg边界或PETM表现出明显的变化,尽管结果非常初步。在我们早期关于西非古新世钻岩双壳类首次记录的报告的基础上,我们在这里进一步描述了白垩纪和古近纪以现代热带特征分类群为主的软体动物动物群。在新发现的骨鱼目化石中,巨齿螈和一种新的始新世淡水鲶鱼的体型都很大,这是在非洲发现的最大的鲶鱼化石之一。我们新的古生态和动物重建显示了一个常绿阔叶林,其中包括一些已知的最古老的红树林。古代马里生态系统中有许多顶级捕食者,包括鳄鱼类、蛇类和阿米达科,其中一些是其分支中最大的物种。跨撒哈拉海道与主要海洋表现出间歇性的隔绝。这种环境变量可能创造了水生特有中心,刺激了对巨大的选择,就像以前在陆地岛屿上观察到的物种一样。
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引用次数: 15
Systematics of the Short-Tailed Whipscorpion Genus Stenochrus Chamberlin, 1922 (Schizomida: Hubbardiidae), with Descriptions of Six New Genera and Five New Species 短尾鞭虫属Stenochrus Chamberlin的系统学,1922(裂殖虫目:Hubbardiidae),附6个新属和5个新种的描述
IF 3.4 2区 环境科学与生态学 Q1 Agricultural and Biological Sciences Pub Date : 2019-06-20 DOI: 10.1206/0003-0090.435.1.1
Rodrigo Monjaraz-Ruedas, L. Prendini, O. Francke
. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .5 Introduction. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .5 Material and Methods . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .9 Systematics . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .16 Key to Identification of the North American Genera of Hubbardiidae (Schizomida) . . . .16 Family Hubbardiidae Cook, 1899 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .18 Subfamily Hubbardiinae Cook, 1899. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .18 Ambulantactus, gen. nov. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .18 Ambulantactus aquismon, sp. nov. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .22 Ambulantactus davisi (Gertsch, 1940), comb. nov. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .24 Ambulantactus montielae, sp. nov. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .24 Baalrog, gen. nov. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .27 Baalrog firstmani (Rowland, 1973), comb. nov. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .32 Baalrog magico (Monjaraz-Ruedas and Francke, 2018), comb. nov. . . . . . . . . . . . . . . . . . . .33 Baalrog sbordonii (Brignoli, 1973), comb. nov. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .34 Baalrog yacato sp. nov. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .37 Harveyus, gen. nov.. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .37 Harveyus contrerasi, sp. nov. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .39 Harveyus mexicanus (Rowland, 1971), comb. nov. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .40 Harveyus mulaiki (Gertsch, 1940), comb. nov. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .41 Harveyus reddelli (Rowland, 1971), comb. nov.. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .41 Heteroschizomus Rowland, 1973, stat. rev. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .44 Heteroschizomus goodnightorum Rowland, 1973 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
5引言。5材料和方法。9系统学。16 Hubbardiidae(Schizomida)北美属鉴定的关键。16 Hubbardiidae Cook家族,1899年。18 Hubbardiinae Cook家族,1899年。18 Ambulatactus,gen.nov。11月24日………….月24日Ambulatacus montielae,sp.nov。11月…………..32 Baarog magico(Monjaraz Ruedas和Francke,2018),comb。11月…………33日Baarog sbordonii(Brignoli,1973),梳。11月…………..34 Baarog yacato sp,sp.nov。11月40日Harveyus mulaiki(Gertsch,1940),梳。11月41日Harveyus redelli(罗兰,1971),梳。11月…………..41罗兰异裂殖吸虫,1973年,统计修订版。47 kekchi异裂殖吸虫,sp.nov。11月………….49异源裂殖吸虫(Rowland and Reddell,1977),梳。11月…………..51 Nahual,gen.11月…..52 Nahual bokmai,sp.11月54 Nahual caballero(Monjaraz Ruedas和Francke,2018),comb。11月56日Nahual lanceolatus(罗兰,1975),梳。11月56日Nahual pallidus(罗兰,1975),梳。11月57日Olmacazomus,化名。11月58日Pacal Reddell和Cokendolpher,1995年。59精神分裂症,新一代。11月…………..63卢肯西精神分裂症(罗兰,1973),梳。11月………….………..63 Stenochrus Chamberlin,1922年。64 Stenochrus alcalai Monjaraz Ruedas和Francke,2018。68 Stenochrus chimalapas Monjaraz Ruedas和Francke,2018。 5引言。5材料和方法。9系统学。16 Hubbardiidae(Schizomida)北美属鉴定的关键。16 Hubbardiidae Cook家族,1899年。18 Hubbardiinae Cook家族,1899年。18 Ambulatactus,gen.nov。11月24日………….月24日Ambulatacus montielae,sp.nov。11月…………..32 Baarog magico(Monjaraz Ruedas和Francke,2018),comb。11月…………33日Baarog sbordonii(Brignoli,1973),梳。11月…………..34 Baarog yacato sp,sp.nov。11月40日Harveyus mulaiki(Gertsch,1940),梳。11月41日Harveyus redelli(罗兰,1971),梳。11月…………..41罗兰异裂殖吸虫,1973年,统计修订版。47 kekchi异裂殖吸虫,sp.nov。11月………….49异源裂殖吸虫(Rowland and Reddell,1977),梳。11月…………..51 Nahual,gen.11月…..52 Nahual bokmai,sp.11月54 Nahual caballero(Monjaraz Ruedas和Francke,2018),comb。11月56日Nahual lanceolatus(罗兰,1975),梳。11月56日Nahual pallidus(罗兰,1975),梳。11月57日Olmacazomus,化名。11月58日Pacal Reddell和Cokendolpher,1995年。59精神分裂症,新一代。11月…………..63卢肯西精神分裂症(罗兰,1973),梳。11月………….………..63 Stenochrus Chamberlin,1922年。64 Stenochrus alcalai Monjaraz Ruedas和Francke,2018。68 Stenochrus chimalapas Monjaraz Ruedas和Francke,2018。 68 Stenochrus gruta Monjaraz Ruedas和Francke,2018。68 Stenochrus pecki(罗兰,1973)。69张伯林,1922年。69 4美国自然历史博物馆公告编号:435 4 Troglostnechrus,gen.nov。11月…………75日Troglostnechrus valdezi(Monjaraz Ruedas,2012),梳。11月75日讨论。75鸣谢。78参考文献。79附录1。用于系统发育分析的短尾鞭蝎属Stenochrus Chamberlin,1922,及其相关属(裂殖动物门:Hubbardiidae)的形态学特征。88附录2。组织样本的凭证号和DNA序列的GenBank登录码。90
{"title":"Systematics of the Short-Tailed Whipscorpion Genus Stenochrus Chamberlin, 1922 (Schizomida: Hubbardiidae), with Descriptions of Six New Genera and Five New Species","authors":"Rodrigo Monjaraz-Ruedas, L. Prendini, O. Francke","doi":"10.1206/0003-0090.435.1.1","DOIUrl":"https://doi.org/10.1206/0003-0090.435.1.1","url":null,"abstract":". . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .5 Introduction. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .5 Material and Methods . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .9 Systematics . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .16 Key to Identification of the North American Genera of Hubbardiidae (Schizomida) . . . .16 Family Hubbardiidae Cook, 1899 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .18 Subfamily Hubbardiinae Cook, 1899. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .18 Ambulantactus, gen. nov. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .18 Ambulantactus aquismon, sp. nov. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .22 Ambulantactus davisi (Gertsch, 1940), comb. nov. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .24 Ambulantactus montielae, sp. nov. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .24 Baalrog, gen. nov. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .27 Baalrog firstmani (Rowland, 1973), comb. nov. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .32 Baalrog magico (Monjaraz-Ruedas and Francke, 2018), comb. nov. . . . . . . . . . . . . . . . . . . .33 Baalrog sbordonii (Brignoli, 1973), comb. nov. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .34 Baalrog yacato sp. nov. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .37 Harveyus, gen. nov.. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .37 Harveyus contrerasi, sp. nov. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .39 Harveyus mexicanus (Rowland, 1971), comb. nov. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .40 Harveyus mulaiki (Gertsch, 1940), comb. nov. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .41 Harveyus reddelli (Rowland, 1971), comb. nov.. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .41 Heteroschizomus Rowland, 1973, stat. rev. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .44 Heteroschizomus goodnightorum Rowland, 1973 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . ","PeriodicalId":50721,"journal":{"name":"Bulletin of the American Museum of Natural History","volume":null,"pages":null},"PeriodicalIF":3.4,"publicationDate":"2019-06-20","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"42502419","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":2,"RegionCategory":"环境科学与生态学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
引用次数: 7
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Bulletin of the American Museum of Natural History
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