Pub Date : 2017-06-26DOI: 10.1206/0003-0090-415.1.1
Lauren A. Esposito, Humberto Y. Yamaguti, C. A. Souza, R. Pinto‐da‐Rocha, L. Prendini
ABSTRACT The Neotropical “club-tailed” scorpions of the genus Rhopalurus Thorell, 1876, and two related genera in family Buthidae C.L. Koch, 1837, i.e., Physoctonus Mello-Leitão, 1934, and Troglorhopalurus Lourenço et al., 2004, are revised, based on a simultaneous phylogenetic analysis of 90 morphological characters and 4260 aligned DNA nucleotides from three mitochondrial and two nuclear gene loci. The monophyletic New World buthid subfamily Rhopalurusinae Bücherl, 1971, to which these scorpions were originally assigned, is redefined, revised diagnoses and a key to identification of its genera and species (except for Centruroides Marx, 1890) provided, and their distributions mapped. The paraphyly of Rhopalurus Thorell, 1876, which comprises several monophyletic groups congruent with its disjunct distribution, justifies its relimitation and that of Troglorhopalurus Lourenço et al., 2004, the revalidation of Heteroctenus Pocock, 1893, and creation of Ischnotelson, gen. nov. (type species: Rhopalurus guanambiensis Lenarducci, Pinto-da-Rocha and Lucas, 2005) and Jaguajir, gen. nov. (type species: Rhopalurus agamemnon C.L. Koch, 1839). Ten new combinations are proposed: Heteroctenus abudi (Armas and Marcano Fondeur, 1987), comb. nov.; Heteroctenus bonettii (Armas, 1999), comb. nov.; Heteroctenus garridoi (Armas, 1974), comb. nov.; Heteroctenus gibarae (Teruel, 2006), comb. nov.; Heteroctenus princeps (Karsch, 1879), comb. nov.; Ischnotelson guanambiensis (Lenarducci, Pinto-da-Rocha and Lucas, 2005), comb. nov.; Jaguajir agamemnon (C.L. Koch, 1839), comb. nov.; Jaguajir pintoi (Mello-Leitão, 1932), comb. nov.; Jaguajir rochae (Borelli, 1910), comb. nov.; Troglorhopalurus lacrau (Lourenço and Pinto-da-Rocha, 1997), comb. nov. Three new species are described: Ischnotelson peruassu, sp. nov. (type locality: Parque Estadual do Peruassu, Minas Gerias, Brazil); Physoctonus striatus, sp. nov. (type locality: Castelo do Piauí, Piauí, Brazil); Rhopalurus ochoai, sp. nov. (type locality: San Agustín, Edo. Zulia, Venezuela). Fifteen new junior subjective synonyms are proposed: Rhopalurus acromelas Lutz and Mello, 1922, Rhopalurus melleipalpus Lutz and Mello, 1922, Rhopalurus iglesiasi Werner, 1927, Rhopalurus lambdophorus Mello-Leitão, 1932, Rhopalurus dorsomaculatus Prado, 1938, and Rhopalurus goiasensis Prado, 1940 = Jaguajir agamemnon (C.L. Koch, 1839); Rhopalurus pintoi kourouensis Lourenço, 2008 = Jaguajir pintoi (Mello-Leitão, 1932); Rhopalurus crassicauda Caporiacco, 1947, Rhopalurus amazonicus Lourenço, 1986, and Rhopalurus crassicauda paruensis Lourenço, 2008 = Rhopalurus laticauda Thorell, 1876; Rhopalurus melloleitaoi Teruel and Armas, 2006, and Rhopalurus aridicola (Teruel and Armas, 2012) = Heteroctenus junceus (Herbst, 1800); Rhopalurus granulimanus Teruel, 2006 = Heteroctenus gibarae (Teruel, 2006); Rhopalurus virkii Santiago-Blay, 2009 = Heteroctenus abudi (Armas and Marcano Fondeur, 1987); Rhopalurus brejo Lourenço, 2014 = Troglorhopalurus lacr
摘要对Thorell Rhopaluus属的新热带“俱乐部尾”蝎子,1876年,以及Buthidae C.L.Koch科的两个相关属,即Physoctonus Mello Leitão,1934年和Trogloroopaluus Lourenço等人,2004年进行了修订,基于对来自三个线粒体和两个核基因位点的90个形态特征和4260个排列的DNA核苷酸的同时系统发育分析。这些蝎子最初被分配到的新世界布希德亚科Rhopalurusinae Bücherl,1971年的单系,被重新定义、修订了诊断,并提供了识别其属和种的关键(除了Centruroides Marx,1890),并绘制了它们的分布图。Rhopalurus Thorell,1876,包括与其间断分布一致的几个单系群,证明了其重新局限性,Trogloropalurus Lourenço等人,2004,Heteroctenus Pocock的重新验证,1893,以及Ischnotelson,gen.nov.的创建(模式种:Rhopaluus guanabiensis Lenarducci,Pinto da Rocha和Lucas,2005)和Jaguajir,gen.nov.(模式种:龙舌兰C.L.Koch,1839)。提出了10个新的组合:阿布迪异ctenus abudi(Armas和Marcano Fondeur,1987),comb。十一月骨异ctenus bonettii(Armas,1999),梳。十一月加里多伊异ctenus garridoi(Armas,1974),梳。十一月吉氏异ctenus gibarae(Teruel,2006),梳。十一月王子异栉水母(Karsch,1879),梳。十一月Guanabiensis的Ischnotelson(Lenarducci,Pinto da Rocha和Lucas,2005),梳。十一月Jaguajir agamemnon(C.L.Koch,1839),梳。十一月Jaguajir pintoi(Mello Leitão,1932),梳。十一月Jaguajir rochae(Borelli,1910),梳。十一月Troglorhopalurus lacrau(Lourenço和Pinto da Rocha,1997),梳。nov.描述了三个新种:Ischnotelson peruassu,sp.nov.(类型地点:Parque Estadial do peruassu、Minas Gerias,巴西);条纹藻(Physoctonus striatus,sp.nov.)(类型产地:巴西皮亚伊Castelo do Piauí);Rhopaurus ochoai,sp.nov.(类型地点:委内瑞拉祖利亚埃多省圣阿古斯丁)。提出了15个新的初级主观同义词:Rhopaurus acromelas Lutz和Mello,1922,Rhopaururus melleipalus Lutz和Mello,1922;Rhopalurus pintoi kourouensis Lourenço,2008=Jaguajir pintoi(Mello Leitão,1932);粗尾Rhopaurus crassicauda Caporiacco,1947年,亚马逊Rhopauru amazonicus Lourenço,1986年,和粗尾Rhapaurus crasicauda paruensis Lourenço,2008=宽尾Rhopairus laticoda Thorell,1876年;Rhopaurus melllolitaoi Teruel和Armas,2006,以及Rhopauru aridicola(Teruel and Armas,2012)=异栉水母(Herbst,1800);麻粒Rhopaurus granlimanus Teruel,2006=吉氏异栉水母(Teruel)(2006);Rhopalurus virkii Santiago Blay,2009=阿布迪异栉水母(Armas和Marcano Fondeur,1987);Rhopalurus brejo Lourenço,2014=Trogloropalurus lacrau(Lourenço和Pinto da Rocha,1997)。
{"title":"Systematic Revision of the Neotropical Club-Tailed Scorpions, Physoctonus, Rhopalurus, and Troglorhopalurus, Revalidation of Heteroctenus, and Descriptions of Two New Genera and Three New Species (Buthidae: Rhopalurusinae)","authors":"Lauren A. Esposito, Humberto Y. Yamaguti, C. A. Souza, R. Pinto‐da‐Rocha, L. Prendini","doi":"10.1206/0003-0090-415.1.1","DOIUrl":"https://doi.org/10.1206/0003-0090-415.1.1","url":null,"abstract":"ABSTRACT The Neotropical “club-tailed” scorpions of the genus Rhopalurus Thorell, 1876, and two related genera in family Buthidae C.L. Koch, 1837, i.e., Physoctonus Mello-Leitão, 1934, and Troglorhopalurus Lourenço et al., 2004, are revised, based on a simultaneous phylogenetic analysis of 90 morphological characters and 4260 aligned DNA nucleotides from three mitochondrial and two nuclear gene loci. The monophyletic New World buthid subfamily Rhopalurusinae Bücherl, 1971, to which these scorpions were originally assigned, is redefined, revised diagnoses and a key to identification of its genera and species (except for Centruroides Marx, 1890) provided, and their distributions mapped. The paraphyly of Rhopalurus Thorell, 1876, which comprises several monophyletic groups congruent with its disjunct distribution, justifies its relimitation and that of Troglorhopalurus Lourenço et al., 2004, the revalidation of Heteroctenus Pocock, 1893, and creation of Ischnotelson, gen. nov. (type species: Rhopalurus guanambiensis Lenarducci, Pinto-da-Rocha and Lucas, 2005) and Jaguajir, gen. nov. (type species: Rhopalurus agamemnon C.L. Koch, 1839). Ten new combinations are proposed: Heteroctenus abudi (Armas and Marcano Fondeur, 1987), comb. nov.; Heteroctenus bonettii (Armas, 1999), comb. nov.; Heteroctenus garridoi (Armas, 1974), comb. nov.; Heteroctenus gibarae (Teruel, 2006), comb. nov.; Heteroctenus princeps (Karsch, 1879), comb. nov.; Ischnotelson guanambiensis (Lenarducci, Pinto-da-Rocha and Lucas, 2005), comb. nov.; Jaguajir agamemnon (C.L. Koch, 1839), comb. nov.; Jaguajir pintoi (Mello-Leitão, 1932), comb. nov.; Jaguajir rochae (Borelli, 1910), comb. nov.; Troglorhopalurus lacrau (Lourenço and Pinto-da-Rocha, 1997), comb. nov. Three new species are described: Ischnotelson peruassu, sp. nov. (type locality: Parque Estadual do Peruassu, Minas Gerias, Brazil); Physoctonus striatus, sp. nov. (type locality: Castelo do Piauí, Piauí, Brazil); Rhopalurus ochoai, sp. nov. (type locality: San Agustín, Edo. Zulia, Venezuela). Fifteen new junior subjective synonyms are proposed: Rhopalurus acromelas Lutz and Mello, 1922, Rhopalurus melleipalpus Lutz and Mello, 1922, Rhopalurus iglesiasi Werner, 1927, Rhopalurus lambdophorus Mello-Leitão, 1932, Rhopalurus dorsomaculatus Prado, 1938, and Rhopalurus goiasensis Prado, 1940 = Jaguajir agamemnon (C.L. Koch, 1839); Rhopalurus pintoi kourouensis Lourenço, 2008 = Jaguajir pintoi (Mello-Leitão, 1932); Rhopalurus crassicauda Caporiacco, 1947, Rhopalurus amazonicus Lourenço, 1986, and Rhopalurus crassicauda paruensis Lourenço, 2008 = Rhopalurus laticauda Thorell, 1876; Rhopalurus melloleitaoi Teruel and Armas, 2006, and Rhopalurus aridicola (Teruel and Armas, 2012) = Heteroctenus junceus (Herbst, 1800); Rhopalurus granulimanus Teruel, 2006 = Heteroctenus gibarae (Teruel, 2006); Rhopalurus virkii Santiago-Blay, 2009 = Heteroctenus abudi (Armas and Marcano Fondeur, 1987); Rhopalurus brejo Lourenço, 2014 = Troglorhopalurus lacr","PeriodicalId":50721,"journal":{"name":"Bulletin of the American Museum of Natural History","volume":"1 1","pages":"1 - 136"},"PeriodicalIF":3.4,"publicationDate":"2017-06-26","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"https://sci-hub-pdf.com/10.1206/0003-0090-415.1.1","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"48064954","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":2,"RegionCategory":"环境科学与生态学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
Pub Date : 2017-06-26DOI: 10.1206/0003-0090-414.1.4
I. Walaszczyk, A. Plint, N. Landman
ABSTRACT Inoceramid bivalves are the dominant invertebrate fauna of the Coniacian and basal Santonian of the Western Canada Foreland Basin in western Alberta. In the upper lower Coniacian through to basal Santonian, six successive faunas are recognized, which provide the basis for corresponding, formally defined inoceramid zones. From bottom upward these are the zones of: Cremnoceramus crassus crassus /C. deformis deformis, Inoceramus gibbosus, Volviceramus koeneni, Volviceramus involutus, Sphenoceramus subcardissoides, and Sphenoceramus ex gr. pachti. Particular faunas represent assemblages known widely from the Euramerican biogeographic region, although they characterize mostly its northern, boreal area. The inoceramid-based biostratigraphic scheme allows correlation with other parts of the North American Western Interior and with parts of the Euramerican biogeographic region. The studied succession provides a good record of the Inoceramus gibbosus Zone, which characterizes the topmost lower Coniacian. This zone, first recognized from northern Germany, is usually absent, both in Europe and in North America, due to a stratigraphic gap resulting from a eustatic lowstand. The base of the middle Coniacian is marked by the abrupt appearance of the taxonomically variable Volviceramus fauna (V. koeneni (Müller), V. exogyroides (Meek and Hayden)), with associated Inoceramus undabundus Meek and Hayden and Volviceramus cardinalensis, newly described herein. Scaphites (Scaphites) ventricosus Meek and Hayden, the ammonite marker of the base of the middle Coniacian first appears in the late early Coniacian. The base of the upper Coniacian marks the first appearance of the characteristic northern inoceramid species Sphenoceramus subcardissoides (Schlüter), the appearance of which coincides with Scaphites (Scaphites) depressus Reeside, the ammonite marker of this boundary. Close to this boundary Volviceramus stotti also appears, which is newly described from the Canadian sections. The base of the Santonian corresponds to the abrupt appearance of Sphenoceramus ex gr. pachti (Arkhangelsky). The studied sections demonstrate that the appearance of new inoceramid faunas (lowest occurrence of Cremnoceramus crassus crassus (Petrascheck), of various species of Volviceramus, Sphenoceramus subcardissoides (Schlüter) and of S. ex gr. pachti) takes place immediately above major marine flooding surfaces, suggesting a close correspondence between evolutionary and/or migration events and episodes of relative sea-level rise.
{"title":"Chapter 2: Inoceramid Bivalves from the Coniacian and Basal Santonian (Upper Cretaceous) of the Western Canada Foreland Basin","authors":"I. Walaszczyk, A. Plint, N. Landman","doi":"10.1206/0003-0090-414.1.4","DOIUrl":"https://doi.org/10.1206/0003-0090-414.1.4","url":null,"abstract":"ABSTRACT Inoceramid bivalves are the dominant invertebrate fauna of the Coniacian and basal Santonian of the Western Canada Foreland Basin in western Alberta. In the upper lower Coniacian through to basal Santonian, six successive faunas are recognized, which provide the basis for corresponding, formally defined inoceramid zones. From bottom upward these are the zones of: Cremnoceramus crassus crassus /C. deformis deformis, Inoceramus gibbosus, Volviceramus koeneni, Volviceramus involutus, Sphenoceramus subcardissoides, and Sphenoceramus ex gr. pachti. Particular faunas represent assemblages known widely from the Euramerican biogeographic region, although they characterize mostly its northern, boreal area. The inoceramid-based biostratigraphic scheme allows correlation with other parts of the North American Western Interior and with parts of the Euramerican biogeographic region. The studied succession provides a good record of the Inoceramus gibbosus Zone, which characterizes the topmost lower Coniacian. This zone, first recognized from northern Germany, is usually absent, both in Europe and in North America, due to a stratigraphic gap resulting from a eustatic lowstand. The base of the middle Coniacian is marked by the abrupt appearance of the taxonomically variable Volviceramus fauna (V. koeneni (Müller), V. exogyroides (Meek and Hayden)), with associated Inoceramus undabundus Meek and Hayden and Volviceramus cardinalensis, newly described herein. Scaphites (Scaphites) ventricosus Meek and Hayden, the ammonite marker of the base of the middle Coniacian first appears in the late early Coniacian. The base of the upper Coniacian marks the first appearance of the characteristic northern inoceramid species Sphenoceramus subcardissoides (Schlüter), the appearance of which coincides with Scaphites (Scaphites) depressus Reeside, the ammonite marker of this boundary. Close to this boundary Volviceramus stotti also appears, which is newly described from the Canadian sections. The base of the Santonian corresponds to the abrupt appearance of Sphenoceramus ex gr. pachti (Arkhangelsky). The studied sections demonstrate that the appearance of new inoceramid faunas (lowest occurrence of Cremnoceramus crassus crassus (Petrascheck), of various species of Volviceramus, Sphenoceramus subcardissoides (Schlüter) and of S. ex gr. pachti) takes place immediately above major marine flooding surfaces, suggesting a close correspondence between evolutionary and/or migration events and episodes of relative sea-level rise.","PeriodicalId":50721,"journal":{"name":"Bulletin of the American Museum of Natural History","volume":"1 1","pages":"103 - 53"},"PeriodicalIF":3.4,"publicationDate":"2017-06-26","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"https://sci-hub-pdf.com/10.1206/0003-0090-414.1.4","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"44485421","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":2,"RegionCategory":"环境科学与生态学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
Pub Date : 2017-06-26DOI: 10.1206/0003-0090-414.1.1
N. Landman, A. Plint, I. Walaszczyk
ABSTRACT The Upper Cretaceous (Coniacian-Santonian) of the Western Canada Foreland Basin, contains a rich record of scaphitid ammonites (scaphites). We describe four species: Scaphites (Scaphites) preventricosus Cobban, 1952, Scaphites (S.) ventricosus Meek and Hayden, 1862, Scaphites (S.) depressus Reeside, 1927, and Clioscaphites saxitonianus (McLearn, 1929). These are widespread index fossils that demarcate the upper lower-middle, middle, and upper Coniacian, and the lower Santonian, respectively. They occur in the lower part of the Wapiabi Formation, Alberta. The Coniacian part of the section has been divided into 24 informal allomembers based on the recognition of marine flooding surfaces, most of which can be traced through the >750 km extent of the study area. The most distinctive feature in the ontogenetic development of scaphites is the change in coiling during ontogeny. At the approach of maturity, the shell uncoils slightly, forming a shaft, which then recurves backward approaching the earlier secreted phragmocone. However, this sequence of scaphites shows an evolutionary trend toward recoiling, accompanied by an increase in size and degree of depression. These changes occurred against a background of changing environmental conditions resulting from the expansion of the Western Interior Seaway during the Niobrara transgression. This resulted in an increase in the area of offshore habitats, which may have promoted the appearance of larger species with more depressed whorl sections. Scaphites probably lived at depths of less than 100 m, and may have fed on small organisms in the water column.
{"title":"Chapter 3: Scaphitid Ammonites from the Upper Cretaceous (Coniacian-Santonian) Western Canada Foreland Basin","authors":"N. Landman, A. Plint, I. Walaszczyk","doi":"10.1206/0003-0090-414.1.1","DOIUrl":"https://doi.org/10.1206/0003-0090-414.1.1","url":null,"abstract":"ABSTRACT The Upper Cretaceous (Coniacian-Santonian) of the Western Canada Foreland Basin, contains a rich record of scaphitid ammonites (scaphites). We describe four species: Scaphites (Scaphites) preventricosus Cobban, 1952, Scaphites (S.) ventricosus Meek and Hayden, 1862, Scaphites (S.) depressus Reeside, 1927, and Clioscaphites saxitonianus (McLearn, 1929). These are widespread index fossils that demarcate the upper lower-middle, middle, and upper Coniacian, and the lower Santonian, respectively. They occur in the lower part of the Wapiabi Formation, Alberta. The Coniacian part of the section has been divided into 24 informal allomembers based on the recognition of marine flooding surfaces, most of which can be traced through the >750 km extent of the study area. The most distinctive feature in the ontogenetic development of scaphites is the change in coiling during ontogeny. At the approach of maturity, the shell uncoils slightly, forming a shaft, which then recurves backward approaching the earlier secreted phragmocone. However, this sequence of scaphites shows an evolutionary trend toward recoiling, accompanied by an increase in size and degree of depression. These changes occurred against a background of changing environmental conditions resulting from the expansion of the Western Interior Seaway during the Niobrara transgression. This resulted in an increase in the area of offshore habitats, which may have promoted the appearance of larger species with more depressed whorl sections. Scaphites probably lived at depths of less than 100 m, and may have fed on small organisms in the water column.","PeriodicalId":50721,"journal":{"name":"Bulletin of the American Museum of Natural History","volume":"1 1","pages":"105 - 172"},"PeriodicalIF":3.4,"publicationDate":"2017-06-26","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"https://sci-hub-pdf.com/10.1206/0003-0090-414.1.1","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"45408924","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":2,"RegionCategory":"环境科学与生态学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
Pub Date : 2017-06-26DOI: 10.1206/0003-0090-414.1.2
N. Landman, A. Guy Plint, I. Walaszczyk
172 pages, 15 folded leaves of plates : illustrations (some color), maps (some color) ; 26 cm. chapter 1. Integrated, high-resolution allostratigraphic, biostratigraphic and carbon-isotope correlation of Coniacian strata (Upper Cretaceous), western Alberta and northern Montana / A. Guy Plint, Elizabeth A. Hooper, Meriem D. Grifi, Ireneusz Walaszczyk, Neil H. Landman, Darren R. Grocke, Joao P. Trabucho Alexandre, and Ian Jarvis -- chapter 2. Inoceramid bivalves from the Coniacian and basal Santonian (Upper Cretaceous) of the Western Canada Foreland Basin / Ireneusz Walaszczyk, A. Guy Plint, and Neil H. Landman -- chapter 3. Scaphitid ammonites from the Upper Cretaceous (Coniacian-Santonian) Western Canada Foreland Basin / Neil H. Landman, A. Guy Plint, and Ireneusz Walaszczyk.
{"title":"Allostratigraphy and Biostratigraphy of the Upper Cretaceous (Coniacian-Santonian) Western Canada Foreland Basin","authors":"N. Landman, A. Guy Plint, I. Walaszczyk","doi":"10.1206/0003-0090-414.1.2","DOIUrl":"https://doi.org/10.1206/0003-0090-414.1.2","url":null,"abstract":"172 pages, 15 folded leaves of plates : illustrations (some color), maps (some color) ; 26 cm. \u0000chapter 1. Integrated, high-resolution allostratigraphic, biostratigraphic and carbon-isotope correlation of Coniacian strata (Upper Cretaceous), western Alberta and northern Montana / A. Guy Plint, Elizabeth A. Hooper, Meriem D. Grifi, Ireneusz Walaszczyk, Neil H. Landman, Darren R. Grocke, Joao P. Trabucho Alexandre, and Ian Jarvis -- chapter 2. Inoceramid bivalves from the Coniacian and basal Santonian (Upper Cretaceous) of the Western Canada Foreland Basin / Ireneusz Walaszczyk, A. Guy Plint, and Neil H. Landman -- chapter 3. Scaphitid ammonites from the Upper Cretaceous (Coniacian-Santonian) Western Canada Foreland Basin / Neil H. Landman, A. Guy Plint, and Ireneusz Walaszczyk.","PeriodicalId":50721,"journal":{"name":"Bulletin of the American Museum of Natural History","volume":"1 1","pages":"1 - 172"},"PeriodicalIF":3.4,"publicationDate":"2017-06-26","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"43255611","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":2,"RegionCategory":"环境科学与生态学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
Pub Date : 2017-06-09DOI: 10.1206/0003-0090-413.1.1
B. Bai, Jin Meng, Yuanqing Wang, Haibing Wang, L. Holbrook
ABSTRACT The middle Eocene ceratomorph Hyrachyus has been considered a pivotal genus in ceratomorph evolution, either as a transitional form from tapiroids to rhinocerotoids, giving rise to all later rhinocerotoids, or else as the sister taxon to other rhinocerotoids. Thus, Hyrachyus has been commonly chosen as an outgroup in phylogenetic analyses of rhinocerotoids. However, little has been published on the osteology of Hyrachyus, even though well-preserved craniodental and postcranial specimens of this taxon have been in collections for decades. Here, we describe and illustrate the cranial and postcranial osteology of Hyrachyus modestus, based mainly on the exceptionally preserved specimens housed at the American Museum of Natural History, specifically AMNH FM 12664. Our bone-by-bone description provides detailed information on the osteological morphology of Hyrachyus, which should be useful for phylogenetic analyses of both rhinocerotoids and perissodactyls in general, because it provides one of the more complete and best-preserved examples of the skeleton of an earlier Eocene perissodactyl. The cranial morphology of Hyrachyus modestus shows a shallow narial notch, a lacrimal contacting the nasal, and a sphenorbital fissure closely situated to the anterior opening of the alisphenoid canal. In the basicranial region, there is a mastoid exposure of the petrosal between the occipital and the squamosal, and the posttympanic process and paracondylar process are partly fused. The postcranial morphology of Hyrachyus modestus includes the following features: The cervical region of the vertebral column is relatively short compared to the rest of the vertebral column. The lumbar vertebrae have concave-convex embracing prezygapophyses and postzygapophyses. The scapula has a distinct acromion process. The humerus has a greater tubercle that does not elevate above the head, and the deltoid tuberosity and deltopectoral crest are weak. The scaphoid and lunar facets of the radius are confluent. The olecranon of the ulna extends posteroproximally. The manus is functionally tetradactyl, with a complete fifth manual digit. The innominate bone has a long, narrow coxal tuberosity. The greater trochanter of the femur is elevated proximally above the head. The femur has a long, narrow, and symmetric trochlea. The patella has a moderately anteroposteriorly deep base. The intercondyloid eminences of the tibia are equal in height, and the extensor sulcus of the tibia is relatively deep. The fibula has a relatively slender shaft with expanded ends. The pes has three functional digits. The calcaneus does not contact the navicular, nor does the Mt III contact the cuboid. Comparisons between the skeleton of Hyrachyus modestus and those of the early tapiroid Heptodon, the hyracodontid Triplopus, the paraceratheriid Juxia, and the rhinocerotid Uintaceras were also investigated. These results indicate that Hyrachyus probably did not derive from Heptodon, but from a more basal
中始新世角鼻虫Hyrachyus被认为是角鼻虫进化过程中的一个关键属,它是角鼻虫从类貘到类鼻虫的过渡形式,产生了所有后来的类鼻虫,或者是其他类鼻虫的姐妹分类群。因此,在类犀牛的系统发育分析中,水螅类通常被选为外群。然而,尽管保存完好的颅齿和颅后标本已经被收集了几十年,但关于水龙的骨学研究却很少发表。在这里,我们主要基于美国自然历史博物馆保存的特殊标本,特别是AMNH FM 12664,描述和说明了水龙的颅骨和颅后骨学。我们对每块骨头的描述提供了水龙骨形态的详细信息,这对于类鼻兽和一般的异趾龙的系统发育分析都是有用的,因为它提供了始新世早期异趾龙骨骼的一个更完整和保存最好的例子。小圆鼻蛇的颅骨形态为浅鼻切迹,泪道与鼻腔相连,蝶眶裂紧靠翼蝶骨管的前开口。在颅底区,枕骨和鳞骨之间的岩骨有乳突暴露,鼓室后突和髁旁突部分融合。后颅形态包括以下特征:与脊柱的其余部分相比,脊柱的颈椎区域相对较短。腰椎呈凹凸状,包括肩关节前和肩关节后。肩胛骨有明显的肩峰突。肱骨有一个不高于头部的大结节,三角结节和三角胸峰较弱。桡骨的舟状面和月状面汇合。尺骨鹰嘴后近端延伸。手在功能上是四指形的,有完整的第五个手指。无名骨有一长而窄的尾结节。股骨大转子在头部上方近端升高。股骨有一个长、窄、对称的滑车。髌骨有一个中等正后方深的基底。胫骨髁间突高度相等,胫骨伸肌沟相对较深。腓骨有一个相对细长的轴,末端扩张。pes有三个功能性数字。跟骨不接触舟骨,第三股肌也不接触长方体。本文还比较了普通水龙与早期tapioid Heptodon、hyracodontid Triplopus、paraceroid Juxia和rhinocerotid untaceras的骨骼。这些结果表明Hyrachyus可能不是来自七齿兽,而是来自一个更基础的角兽类。此外,水龙与水龙科最早的代表——三足龙的骨骼差异表明,水龙不是水龙的后代。然而,类水螅类的祖先可能产生了其他非水螅类的类鼻。和其他始新世的异趾龙一样,普通蹄龙的后颅形态显示出适应性,这表明在异趾龙进化的早期就已经出现了移动运动。
{"title":"Osteology of The Middle Eocene Ceratomorph Hyrachyus modestus (Mammalia, Perissodactyla)","authors":"B. Bai, Jin Meng, Yuanqing Wang, Haibing Wang, L. Holbrook","doi":"10.1206/0003-0090-413.1.1","DOIUrl":"https://doi.org/10.1206/0003-0090-413.1.1","url":null,"abstract":"ABSTRACT The middle Eocene ceratomorph Hyrachyus has been considered a pivotal genus in ceratomorph evolution, either as a transitional form from tapiroids to rhinocerotoids, giving rise to all later rhinocerotoids, or else as the sister taxon to other rhinocerotoids. Thus, Hyrachyus has been commonly chosen as an outgroup in phylogenetic analyses of rhinocerotoids. However, little has been published on the osteology of Hyrachyus, even though well-preserved craniodental and postcranial specimens of this taxon have been in collections for decades. Here, we describe and illustrate the cranial and postcranial osteology of Hyrachyus modestus, based mainly on the exceptionally preserved specimens housed at the American Museum of Natural History, specifically AMNH FM 12664. Our bone-by-bone description provides detailed information on the osteological morphology of Hyrachyus, which should be useful for phylogenetic analyses of both rhinocerotoids and perissodactyls in general, because it provides one of the more complete and best-preserved examples of the skeleton of an earlier Eocene perissodactyl. The cranial morphology of Hyrachyus modestus shows a shallow narial notch, a lacrimal contacting the nasal, and a sphenorbital fissure closely situated to the anterior opening of the alisphenoid canal. In the basicranial region, there is a mastoid exposure of the petrosal between the occipital and the squamosal, and the posttympanic process and paracondylar process are partly fused. The postcranial morphology of Hyrachyus modestus includes the following features: The cervical region of the vertebral column is relatively short compared to the rest of the vertebral column. The lumbar vertebrae have concave-convex embracing prezygapophyses and postzygapophyses. The scapula has a distinct acromion process. The humerus has a greater tubercle that does not elevate above the head, and the deltoid tuberosity and deltopectoral crest are weak. The scaphoid and lunar facets of the radius are confluent. The olecranon of the ulna extends posteroproximally. The manus is functionally tetradactyl, with a complete fifth manual digit. The innominate bone has a long, narrow coxal tuberosity. The greater trochanter of the femur is elevated proximally above the head. The femur has a long, narrow, and symmetric trochlea. The patella has a moderately anteroposteriorly deep base. The intercondyloid eminences of the tibia are equal in height, and the extensor sulcus of the tibia is relatively deep. The fibula has a relatively slender shaft with expanded ends. The pes has three functional digits. The calcaneus does not contact the navicular, nor does the Mt III contact the cuboid. Comparisons between the skeleton of Hyrachyus modestus and those of the early tapiroid Heptodon, the hyracodontid Triplopus, the paraceratheriid Juxia, and the rhinocerotid Uintaceras were also investigated. These results indicate that Hyrachyus probably did not derive from Heptodon, but from a more basal ","PeriodicalId":50721,"journal":{"name":"Bulletin of the American Museum of Natural History","volume":"30 1","pages":"1 - 70"},"PeriodicalIF":3.4,"publicationDate":"2017-06-09","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"https://sci-hub-pdf.com/10.1206/0003-0090-413.1.1","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"66160857","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":2,"RegionCategory":"环境科学与生态学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
Pub Date : 2017-04-13DOI: 10.1206/0003-0090-412.1.1
V. González, Allan H. Smith-Pardo, M. Engel
ABSTRACT We establish a new genus for an unusual species of Peruvian calliopsine bees (Panurginae: Calliopsini) that was initially reported in the literature as an undescribed species of Spinoliella Ashmead that purportedly expanded the range of the latter genus beyond Argentina and Chile. Although the new genus superficially resembles Spinoliella, it is easily distinguished by a unique combination of characters in both sexes but particularly in the male hidden metasomal sterna and genitalia. A cladistic analysis of 82 adult external morphological characters including all species of Spinoliella, as well as species of the remaining genera of Calliopsini, suggests that this group is sister to a clade consisting of Spinoliella and Callonychium Brèthes. We describe and illustrate Xeranthrena imponticula Gonzalez and Engel, new genus and species, from males and females collected in xeric areas along the Pacific slopes of the Peruvian Andes. In addition, the phylogenetic study suggests two well-defined clades within Spinoliella and corresponding to the previously recognized subgenera, although we do not advocate for their reinstatement. We briefly discuss new putative synapomorphies for Spinoliella and, building upon prior revisionary work, we describe and figure five new species: Spinoliella aidae Gonzalez, Smith-Pardo, and Engel, new species; S. confusa Gonzalez and Engel, new species; S. propinqua Gonzalez and Engel, new species; S. packeri Gonzalez and Engel, new species; and S. polita Gonzalez and Engel, new species. In addition, we synonymize S. karhadra Rodríguez, Toro, and Ruz under S. rufiventris Toro and Ruz (new synonymy). We provide new geographical and floral records, an identification keys to all 17 recognized species of Spinoliella, and updated key to the genera of Calliopsini.
{"title":"Phylogenetic Relationships of a New Genus of Calliopsine Bees from Peru, with a Review of Spinoliella Ashmead (Hymenoptera: Andrenidae)","authors":"V. González, Allan H. Smith-Pardo, M. Engel","doi":"10.1206/0003-0090-412.1.1","DOIUrl":"https://doi.org/10.1206/0003-0090-412.1.1","url":null,"abstract":"ABSTRACT We establish a new genus for an unusual species of Peruvian calliopsine bees (Panurginae: Calliopsini) that was initially reported in the literature as an undescribed species of Spinoliella Ashmead that purportedly expanded the range of the latter genus beyond Argentina and Chile. Although the new genus superficially resembles Spinoliella, it is easily distinguished by a unique combination of characters in both sexes but particularly in the male hidden metasomal sterna and genitalia. A cladistic analysis of 82 adult external morphological characters including all species of Spinoliella, as well as species of the remaining genera of Calliopsini, suggests that this group is sister to a clade consisting of Spinoliella and Callonychium Brèthes. We describe and illustrate Xeranthrena imponticula Gonzalez and Engel, new genus and species, from males and females collected in xeric areas along the Pacific slopes of the Peruvian Andes. In addition, the phylogenetic study suggests two well-defined clades within Spinoliella and corresponding to the previously recognized subgenera, although we do not advocate for their reinstatement. We briefly discuss new putative synapomorphies for Spinoliella and, building upon prior revisionary work, we describe and figure five new species: Spinoliella aidae Gonzalez, Smith-Pardo, and Engel, new species; S. confusa Gonzalez and Engel, new species; S. propinqua Gonzalez and Engel, new species; S. packeri Gonzalez and Engel, new species; and S. polita Gonzalez and Engel, new species. In addition, we synonymize S. karhadra Rodríguez, Toro, and Ruz under S. rufiventris Toro and Ruz (new synonymy). We provide new geographical and floral records, an identification keys to all 17 recognized species of Spinoliella, and updated key to the genera of Calliopsini.","PeriodicalId":50721,"journal":{"name":"Bulletin of the American Museum of Natural History","volume":"1 1","pages":"1 - 71"},"PeriodicalIF":3.4,"publicationDate":"2017-04-13","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"https://sci-hub-pdf.com/10.1206/0003-0090-412.1.1","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"45329321","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":2,"RegionCategory":"环境科学与生态学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
Pub Date : 2017-04-13DOI: 10.1206/0003-0090-411.1.1
R. Pei, Quanguo Li, Qingjin Meng, M. Norell, K. Gao
ABSTRACT Four new specimens of Anchiornis huxleyi (PKUP V1068, BMNHC PH804, BMNHC PH822, and BMNHC PH823) were recently recovered from the Late Jurassic fossil beds of the Tiaojishan Formation in northeastern China. These new specimens are almost completely preserved with cranial and postcranial skeletons. Morphological features of Anchiornis huxleyi have implications for paravian character evolution and provide insights into the relationships of major paravian lineages. Anchiornis huxleyi shares derived features with avialans, such as a straight nasal process of the premaxilla and the absence of an external mandibular fenestra in lateral view. However, Anchiornis huxleyi lacks several derived deinonychosaurian features, including a laterally exposed splenial and a specialized raptorial pedal digit II. Morphological comparisons strongly suggest Anchiornis is more closely related to avialans than to deinonychosaurians or troodontids. Anchiornis huxleyi exhibits many conservative paravian features, and closely resembles Archaeopteryx and other Jurassic paravians from Jianchang County, such as Xiaotingia and Eosinopteryx. The other Jianchang paravian, Aurornis xui, is likely a junior synonym of Anchiornis huxleyi.
{"title":"New Specimens of Anchiornis huxleyi (Theropoda: Paraves) from the Late Jurassic of Northeastern China","authors":"R. Pei, Quanguo Li, Qingjin Meng, M. Norell, K. Gao","doi":"10.1206/0003-0090-411.1.1","DOIUrl":"https://doi.org/10.1206/0003-0090-411.1.1","url":null,"abstract":"ABSTRACT Four new specimens of Anchiornis huxleyi (PKUP V1068, BMNHC PH804, BMNHC PH822, and BMNHC PH823) were recently recovered from the Late Jurassic fossil beds of the Tiaojishan Formation in northeastern China. These new specimens are almost completely preserved with cranial and postcranial skeletons. Morphological features of Anchiornis huxleyi have implications for paravian character evolution and provide insights into the relationships of major paravian lineages. Anchiornis huxleyi shares derived features with avialans, such as a straight nasal process of the premaxilla and the absence of an external mandibular fenestra in lateral view. However, Anchiornis huxleyi lacks several derived deinonychosaurian features, including a laterally exposed splenial and a specialized raptorial pedal digit II. Morphological comparisons strongly suggest Anchiornis is more closely related to avialans than to deinonychosaurians or troodontids. Anchiornis huxleyi exhibits many conservative paravian features, and closely resembles Archaeopteryx and other Jurassic paravians from Jianchang County, such as Xiaotingia and Eosinopteryx. The other Jianchang paravian, Aurornis xui, is likely a junior synonym of Anchiornis huxleyi.","PeriodicalId":50721,"journal":{"name":"Bulletin of the American Museum of Natural History","volume":"1 1","pages":"1 - 67"},"PeriodicalIF":3.4,"publicationDate":"2017-04-13","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"https://sci-hub-pdf.com/10.1206/0003-0090-411.1.1","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"45827080","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":2,"RegionCategory":"环境科学与生态学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
Pub Date : 2017-02-22DOI: 10.1206/0003-0090-410.1.1
M. Izquierdo, M. Ramírez
ABSTRACT The genus Orchestina Simon is distributed worldwide and is characterized by having an enlarged fourth femur with which these species are capable of jumping. The genus is also characterized by having a well-sclerotized sperm duct, a near H-shaped arrangement of the eye group, a 4-4-3-3 pattern of raised receptors on the tarsal organs of the legs I–IV, respectively, and by lacking spines on all the legs. All these characters, together with molecular evidence, support the monophyly of the genus, as well as its placement as sister group of Oonopinae. Most American species of Orchestina inhabit the forest foliage and canopy, but in unforested areas they also occur in shrubs and grasses. In this work, we revise the American representatives of Orchestina in a comprehensive context for the first time. In the past, six species were known from the Americas: five from the United States and only one from South America, described from Venezuela. After the study of the principal collections of the world and several field trips to several South American countries, we describe 85 new species and redescribe all previously known species. Matching sexes was occasionally problematic; while females are very homogeneous in somatic traits, males may have modifications on different parts of the body, making the matching very difficult. Therefore, in this review 56 of the species are described from only one sex, whereas 20 unmatched species are informally described as morphospecies, pending the discovery of conspecific sexes. Two species, O. pavesiiformis Saaristo and O. dentifera Simon, originally known from Israel and Sri Lanka, respectively, are here reported as introduced in several countries in the Americas and other continents. O. justini Saaristo described from the Seychelles is here considered a synonym of O. dentifera. One species, O. truncata Wunderlich, previously known as a subfossil spider from Colombian Copal is here tentatively redescribed based on recent material from Costa Rica, Colombia, and Ecuador; the female is also described for the first time. The species list by country is as follows (numbers refer to records, independently of the locality of the type material): United States (9 species: O. utahana Chamberlin and Ivie, O. moaba Chamberlin and Ivie, O. obscura Chamberlin and Ivie, O. saltitans Banks, O. nadleri Chickering, the introduced O. pavesiiformis Saaristo, and three new species, O. quasimodo, O. kamehameha, and O. auburndalensis); Mexico (3 species: O. utahana Chamberlin and Ivie, and two new species, O. nahuatl and O. chaparrita); Guatemala (1 new species: O. guatemala); Costa Rica (3 new species: O. laselva, O. griswoldi, and O. chiriqui; and the previously known O. truncata Wunderlich); Panama (5 new species: O. chiriqui, O. labarquei, O. pan, O. campana, and O. galapagos); Jamaica (2 species, the introduced O. dentifera Simon and O. galapagos); Haiti and Dominican Republic (only the introduced O. dentifera Simon); Colombia (6 n
{"title":"Taxonomic Revision of the Jumping Goblin Spiders of the Genus Orchestina Simon, 1882, in the Americas (Araneae: Oonopidae)","authors":"M. Izquierdo, M. Ramírez","doi":"10.1206/0003-0090-410.1.1","DOIUrl":"https://doi.org/10.1206/0003-0090-410.1.1","url":null,"abstract":"ABSTRACT The genus Orchestina Simon is distributed worldwide and is characterized by having an enlarged fourth femur with which these species are capable of jumping. The genus is also characterized by having a well-sclerotized sperm duct, a near H-shaped arrangement of the eye group, a 4-4-3-3 pattern of raised receptors on the tarsal organs of the legs I–IV, respectively, and by lacking spines on all the legs. All these characters, together with molecular evidence, support the monophyly of the genus, as well as its placement as sister group of Oonopinae. Most American species of Orchestina inhabit the forest foliage and canopy, but in unforested areas they also occur in shrubs and grasses. In this work, we revise the American representatives of Orchestina in a comprehensive context for the first time. In the past, six species were known from the Americas: five from the United States and only one from South America, described from Venezuela. After the study of the principal collections of the world and several field trips to several South American countries, we describe 85 new species and redescribe all previously known species. Matching sexes was occasionally problematic; while females are very homogeneous in somatic traits, males may have modifications on different parts of the body, making the matching very difficult. Therefore, in this review 56 of the species are described from only one sex, whereas 20 unmatched species are informally described as morphospecies, pending the discovery of conspecific sexes. Two species, O. pavesiiformis Saaristo and O. dentifera Simon, originally known from Israel and Sri Lanka, respectively, are here reported as introduced in several countries in the Americas and other continents. O. justini Saaristo described from the Seychelles is here considered a synonym of O. dentifera. One species, O. truncata Wunderlich, previously known as a subfossil spider from Colombian Copal is here tentatively redescribed based on recent material from Costa Rica, Colombia, and Ecuador; the female is also described for the first time. The species list by country is as follows (numbers refer to records, independently of the locality of the type material): United States (9 species: O. utahana Chamberlin and Ivie, O. moaba Chamberlin and Ivie, O. obscura Chamberlin and Ivie, O. saltitans Banks, O. nadleri Chickering, the introduced O. pavesiiformis Saaristo, and three new species, O. quasimodo, O. kamehameha, and O. auburndalensis); Mexico (3 species: O. utahana Chamberlin and Ivie, and two new species, O. nahuatl and O. chaparrita); Guatemala (1 new species: O. guatemala); Costa Rica (3 new species: O. laselva, O. griswoldi, and O. chiriqui; and the previously known O. truncata Wunderlich); Panama (5 new species: O. chiriqui, O. labarquei, O. pan, O. campana, and O. galapagos); Jamaica (2 species, the introduced O. dentifera Simon and O. galapagos); Haiti and Dominican Republic (only the introduced O. dentifera Simon); Colombia (6 n","PeriodicalId":50721,"journal":{"name":"Bulletin of the American Museum of Natural History","volume":"1 1","pages":"1 - 362"},"PeriodicalIF":3.4,"publicationDate":"2017-02-22","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"https://sci-hub-pdf.com/10.1206/0003-0090-410.1.1","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"66160820","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":2,"RegionCategory":"环境科学与生态学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
Pub Date : 2016-10-13DOI: 10.1206/0003-0090-408.1.1
D. Grimaldi
ABSTRACT A remarkable diversity of new nonempidoid orthorrhaphan flies from the mid-Cretaceous of Myanmar (Late Albian–Early Cenomanian, ca. 99 Ma) is presented, including 28 species (all but one new) in 22 genera (13 new), and at least 12 families. Two families are new; three genera are unplaced in Tabanomorpha and one unplaced within Brachycera. Comparisons are presented between the amber taxa and extensive lithified taxa from the Jurassic and Cretaceous of eastern Laurasia. In Stratiomyomorpha: A new species of Zhangsolvidae has color patterns and body shape that apparently mimic Vespidae or other stinging aculeate wasps. Diverse new Xylomyidae and Stratiomyidae are described, the latter with male terminalia preserved in detail. In Tabanomorpha: The genus Athericites Mostovski et al. is synonymized with Palaepangonius Ren, and a new species of Galloatherix Nel is described in which the female proboscis is much longer than that of the male. All three genera were attributed to Athericidae, but the amber species reveal they are more basal tabanomorphs. Described are a new genus of stem-group Tabanoidea and a new species of Cratotabanus Martins-Neto and Kucero-Santos (Tabanidae), previously known from the Cretaceous of Brazil and New Jersey. In Nemestrinoidea: Three species of Hirmoneura Meigen are the first Nemestrinidae known from amber; one species has long cerci typical of Recent species of the genus. A new species of the Mesozoic family Rhagionemestriidae reveals this family is closely related to Acroceridae, not Xylophagidae as previously proposed. In Archisargoidea: A new species of Tethepomyiidae is described, the family known only in Cretaceous amber from New Jersey, Spain, and Myanmar. In Asiloidea: Three new species and genera of Bombyliidae are described, two of them with abdominal setal “baskets” distinctive to females of higher bombyliids. A fourth new asiloid genus is a probable stem-group bombyliid. The recently described Pseudorhagio Zhang et al., is transferred from Tabanomorpha to Bombyliidae. The male of a new species of Burmapsilocephala Gaimari and Mostovski (Apsilocephalidae) has terminalia very similar to that of Evocoa (Evocoidae: Recent, Chile), corroborating close relationship of the two families. In Families incertae sedis: A new species of Hilarimorphites Grimaldi and Cumming (Apystomyiidae) is described, the family known only in New Jersey and Burmese amber and the Recent fauna of California, and a sister group to either Cyclorrhapha or Eremoneura. Mysteromyiidae and Eucaudomyiidae, new families, are described, based on highly modified species with vestigial mouthparts, reduced venation, and unique specializations. Discussion is presented on some morphological features in fossil and Recent Brachycera that have apparent phylogenetic significance but are rarely discussed. This study adds further data to the phenotypic diversity, geological history, and biogeography of a major radiation.
{"title":"Diverse Orthorrhaphan Flies (Insecta: Diptera: Brachycera) in Amber From the Cretaceous of Myanmar: Brachycera in Cretaceous Amber, Part VII","authors":"D. Grimaldi","doi":"10.1206/0003-0090-408.1.1","DOIUrl":"https://doi.org/10.1206/0003-0090-408.1.1","url":null,"abstract":"ABSTRACT A remarkable diversity of new nonempidoid orthorrhaphan flies from the mid-Cretaceous of Myanmar (Late Albian–Early Cenomanian, ca. 99 Ma) is presented, including 28 species (all but one new) in 22 genera (13 new), and at least 12 families. Two families are new; three genera are unplaced in Tabanomorpha and one unplaced within Brachycera. Comparisons are presented between the amber taxa and extensive lithified taxa from the Jurassic and Cretaceous of eastern Laurasia. In Stratiomyomorpha: A new species of Zhangsolvidae has color patterns and body shape that apparently mimic Vespidae or other stinging aculeate wasps. Diverse new Xylomyidae and Stratiomyidae are described, the latter with male terminalia preserved in detail. In Tabanomorpha: The genus Athericites Mostovski et al. is synonymized with Palaepangonius Ren, and a new species of Galloatherix Nel is described in which the female proboscis is much longer than that of the male. All three genera were attributed to Athericidae, but the amber species reveal they are more basal tabanomorphs. Described are a new genus of stem-group Tabanoidea and a new species of Cratotabanus Martins-Neto and Kucero-Santos (Tabanidae), previously known from the Cretaceous of Brazil and New Jersey. In Nemestrinoidea: Three species of Hirmoneura Meigen are the first Nemestrinidae known from amber; one species has long cerci typical of Recent species of the genus. A new species of the Mesozoic family Rhagionemestriidae reveals this family is closely related to Acroceridae, not Xylophagidae as previously proposed. In Archisargoidea: A new species of Tethepomyiidae is described, the family known only in Cretaceous amber from New Jersey, Spain, and Myanmar. In Asiloidea: Three new species and genera of Bombyliidae are described, two of them with abdominal setal “baskets” distinctive to females of higher bombyliids. A fourth new asiloid genus is a probable stem-group bombyliid. The recently described Pseudorhagio Zhang et al., is transferred from Tabanomorpha to Bombyliidae. The male of a new species of Burmapsilocephala Gaimari and Mostovski (Apsilocephalidae) has terminalia very similar to that of Evocoa (Evocoidae: Recent, Chile), corroborating close relationship of the two families. In Families incertae sedis: A new species of Hilarimorphites Grimaldi and Cumming (Apystomyiidae) is described, the family known only in New Jersey and Burmese amber and the Recent fauna of California, and a sister group to either Cyclorrhapha or Eremoneura. Mysteromyiidae and Eucaudomyiidae, new families, are described, based on highly modified species with vestigial mouthparts, reduced venation, and unique specializations. Discussion is presented on some morphological features in fossil and Recent Brachycera that have apparent phylogenetic significance but are rarely discussed. This study adds further data to the phenotypic diversity, geological history, and biogeography of a major radiation.","PeriodicalId":50721,"journal":{"name":"Bulletin of the American Museum of Natural History","volume":"408 1","pages":"1 - 131"},"PeriodicalIF":3.4,"publicationDate":"2016-10-13","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"https://sci-hub-pdf.com/10.1206/0003-0090-408.1.1","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"66160797","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":2,"RegionCategory":"环境科学与生态学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
Pub Date : 2016-10-07DOI: 10.1206/0003-0090-409.1.1
M. O'Leary
ABSTRACT Comparative data from the ear region has played an important role in recent combined-data phylogenetic analyses of the relationships of living and extinct Artiodactyla and the position of that clade among Euungulata. These studies have also been important for establishing the phylogenetic position of Cetacea and for understanding the relationships of a diversity of euungulate species to their fossil relatives. Detailed and standardized descriptive reference works of the basicranium for a range of living artiodactylans are not, however, readily available. Here I describe exemplar species from the four major extant terrestrial and semiaquatic artiodactylan clades (Hippopotamidae, Ruminantia, Suina, and Camelidae) and illustrate the anatomy of the ear region with the auditory bulla both in place and removed. Terrestrial artiodactyls exhibit varying degrees of expansion of the bony external acoustic meatus laterally relative to the mediolateral dimensions of the rounded, medial aspect of the auditory bulla, a characteristic that is least developed in Tragulidae. A relatively elongate external acoustic meatus has previously been described as entirely absent in living and fossil cetaceans and in some fossil species such as Diacodexis pakistanensis. Variation also exists in the proximity of the petrosal-bullar complex to midline basicranial bones. Isolation of these bones from other basicranial structures has been previously interpreted as functionally important for underwater hearing in Cetacea. Many artiodactylans have contact between the auditory bulla and the basioccipital but no contact between the deeper pars cochlearis of the petrosal bone and the basioccipital/basisphenoid. Exceptions are species of Hippopotamidae in which both the bulla and the petrosal are separated from midline bones. The functional interpretation of this separation has previously been linked to aquatic hearing, but this association may be more complex than originally thought. Other features observed in the basicrania of terrestrial artiodactylans described here are a general coalescence of basicranial foramina (i.e., the basicapsular fissure, carotid foramen, piriform fenestra, and sometimes the foramen ovale), the development of large and ornate styliform processes in species of Ruminantia, and widespread contact between the auditory bulla and the paracondylar process of the exoccipital.
{"title":"Comparative Basicranial Anatomy of Extant Terrestrial and Semiaquatic Artiodactyla","authors":"M. O'Leary","doi":"10.1206/0003-0090-409.1.1","DOIUrl":"https://doi.org/10.1206/0003-0090-409.1.1","url":null,"abstract":"ABSTRACT Comparative data from the ear region has played an important role in recent combined-data phylogenetic analyses of the relationships of living and extinct Artiodactyla and the position of that clade among Euungulata. These studies have also been important for establishing the phylogenetic position of Cetacea and for understanding the relationships of a diversity of euungulate species to their fossil relatives. Detailed and standardized descriptive reference works of the basicranium for a range of living artiodactylans are not, however, readily available. Here I describe exemplar species from the four major extant terrestrial and semiaquatic artiodactylan clades (Hippopotamidae, Ruminantia, Suina, and Camelidae) and illustrate the anatomy of the ear region with the auditory bulla both in place and removed. Terrestrial artiodactyls exhibit varying degrees of expansion of the bony external acoustic meatus laterally relative to the mediolateral dimensions of the rounded, medial aspect of the auditory bulla, a characteristic that is least developed in Tragulidae. A relatively elongate external acoustic meatus has previously been described as entirely absent in living and fossil cetaceans and in some fossil species such as Diacodexis pakistanensis. Variation also exists in the proximity of the petrosal-bullar complex to midline basicranial bones. Isolation of these bones from other basicranial structures has been previously interpreted as functionally important for underwater hearing in Cetacea. Many artiodactylans have contact between the auditory bulla and the basioccipital but no contact between the deeper pars cochlearis of the petrosal bone and the basioccipital/basisphenoid. Exceptions are species of Hippopotamidae in which both the bulla and the petrosal are separated from midline bones. The functional interpretation of this separation has previously been linked to aquatic hearing, but this association may be more complex than originally thought. Other features observed in the basicrania of terrestrial artiodactylans described here are a general coalescence of basicranial foramina (i.e., the basicapsular fissure, carotid foramen, piriform fenestra, and sometimes the foramen ovale), the development of large and ornate styliform processes in species of Ruminantia, and widespread contact between the auditory bulla and the paracondylar process of the exoccipital.","PeriodicalId":50721,"journal":{"name":"Bulletin of the American Museum of Natural History","volume":"409 1","pages":"1 - 55"},"PeriodicalIF":3.4,"publicationDate":"2016-10-07","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"https://sci-hub-pdf.com/10.1206/0003-0090-409.1.1","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"66160803","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":2,"RegionCategory":"环境科学与生态学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}