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Systematic Revision of the Neotropical Club-Tailed Scorpions, Physoctonus, Rhopalurus, and Troglorhopalurus, Revalidation of Heteroctenus, and Descriptions of Two New Genera and Three New Species (Buthidae: Rhopalurusinae) 新热带棒尾蝎、棒尾蝎、棒尾蝎的系统修订,异尾蝎属的再确认,2新属3新种的描述(布甲科:棒尾蝎科)
IF 3.4 2区 环境科学与生态学 Q1 Agricultural and Biological Sciences Pub Date : 2017-06-26 DOI: 10.1206/0003-0090-415.1.1
Lauren A. Esposito, Humberto Y. Yamaguti, C. A. Souza, R. Pinto‐da‐Rocha, L. Prendini
ABSTRACT The Neotropical “club-tailed” scorpions of the genus Rhopalurus Thorell, 1876, and two related genera in family Buthidae C.L. Koch, 1837, i.e., Physoctonus Mello-Leitão, 1934, and Troglorhopalurus Lourenço et al., 2004, are revised, based on a simultaneous phylogenetic analysis of 90 morphological characters and 4260 aligned DNA nucleotides from three mitochondrial and two nuclear gene loci. The monophyletic New World buthid subfamily Rhopalurusinae Bücherl, 1971, to which these scorpions were originally assigned, is redefined, revised diagnoses and a key to identification of its genera and species (except for Centruroides Marx, 1890) provided, and their distributions mapped. The paraphyly of Rhopalurus Thorell, 1876, which comprises several monophyletic groups congruent with its disjunct distribution, justifies its relimitation and that of Troglorhopalurus Lourenço et al., 2004, the revalidation of Heteroctenus Pocock, 1893, and creation of Ischnotelson, gen. nov. (type species: Rhopalurus guanambiensis Lenarducci, Pinto-da-Rocha and Lucas, 2005) and Jaguajir, gen. nov. (type species: Rhopalurus agamemnon C.L. Koch, 1839). Ten new combinations are proposed: Heteroctenus abudi (Armas and Marcano Fondeur, 1987), comb. nov.; Heteroctenus bonettii (Armas, 1999), comb. nov.; Heteroctenus garridoi (Armas, 1974), comb. nov.; Heteroctenus gibarae (Teruel, 2006), comb. nov.; Heteroctenus princeps (Karsch, 1879), comb. nov.; Ischnotelson guanambiensis (Lenarducci, Pinto-da-Rocha and Lucas, 2005), comb. nov.; Jaguajir agamemnon (C.L. Koch, 1839), comb. nov.; Jaguajir pintoi (Mello-Leitão, 1932), comb. nov.; Jaguajir rochae (Borelli, 1910), comb. nov.; Troglorhopalurus lacrau (Lourenço and Pinto-da-Rocha, 1997), comb. nov. Three new species are described: Ischnotelson peruassu, sp. nov. (type locality: Parque Estadual do Peruassu, Minas Gerias, Brazil); Physoctonus striatus, sp. nov. (type locality: Castelo do Piauí, Piauí, Brazil); Rhopalurus ochoai, sp. nov. (type locality: San Agustín, Edo. Zulia, Venezuela). Fifteen new junior subjective synonyms are proposed: Rhopalurus acromelas Lutz and Mello, 1922, Rhopalurus melleipalpus Lutz and Mello, 1922, Rhopalurus iglesiasi Werner, 1927, Rhopalurus lambdophorus Mello-Leitão, 1932, Rhopalurus dorsomaculatus Prado, 1938, and Rhopalurus goiasensis Prado, 1940 = Jaguajir agamemnon (C.L. Koch, 1839); Rhopalurus pintoi kourouensis Lourenço, 2008 = Jaguajir pintoi (Mello-Leitão, 1932); Rhopalurus crassicauda Caporiacco, 1947, Rhopalurus amazonicus Lourenço, 1986, and Rhopalurus crassicauda paruensis Lourenço, 2008 = Rhopalurus laticauda Thorell, 1876; Rhopalurus melloleitaoi Teruel and Armas, 2006, and Rhopalurus aridicola (Teruel and Armas, 2012) = Heteroctenus junceus (Herbst, 1800); Rhopalurus granulimanus Teruel, 2006 = Heteroctenus gibarae (Teruel, 2006); Rhopalurus virkii Santiago-Blay, 2009 = Heteroctenus abudi (Armas and Marcano Fondeur, 1987); Rhopalurus brejo Lourenço, 2014 = Troglorhopalurus lacr
摘要对Thorell Rhopaluus属的新热带“俱乐部尾”蝎子,1876年,以及Buthidae C.L.Koch科的两个相关属,即Physoctonus Mello Leitão,1934年和Trogloroopaluus Lourenço等人,2004年进行了修订,基于对来自三个线粒体和两个核基因位点的90个形态特征和4260个排列的DNA核苷酸的同时系统发育分析。这些蝎子最初被分配到的新世界布希德亚科Rhopalurusinae Bücherl,1971年的单系,被重新定义、修订了诊断,并提供了识别其属和种的关键(除了Centruroides Marx,1890),并绘制了它们的分布图。Rhopalurus Thorell,1876,包括与其间断分布一致的几个单系群,证明了其重新局限性,Trogloropalurus Lourenço等人,2004,Heteroctenus Pocock的重新验证,1893,以及Ischnotelson,gen.nov.的创建(模式种:Rhopaluus guanabiensis Lenarducci,Pinto da Rocha和Lucas,2005)和Jaguajir,gen.nov.(模式种:龙舌兰C.L.Koch,1839)。提出了10个新的组合:阿布迪异ctenus abudi(Armas和Marcano Fondeur,1987),comb。十一月骨异ctenus bonettii(Armas,1999),梳。十一月加里多伊异ctenus garridoi(Armas,1974),梳。十一月吉氏异ctenus gibarae(Teruel,2006),梳。十一月王子异栉水母(Karsch,1879),梳。十一月Guanabiensis的Ischnotelson(Lenarducci,Pinto da Rocha和Lucas,2005),梳。十一月Jaguajir agamemnon(C.L.Koch,1839),梳。十一月Jaguajir pintoi(Mello Leitão,1932),梳。十一月Jaguajir rochae(Borelli,1910),梳。十一月Troglorhopalurus lacrau(Lourenço和Pinto da Rocha,1997),梳。nov.描述了三个新种:Ischnotelson peruassu,sp.nov.(类型地点:Parque Estadial do peruassu、Minas Gerias,巴西);条纹藻(Physoctonus striatus,sp.nov.)(类型产地:巴西皮亚伊Castelo do Piauí);Rhopaurus ochoai,sp.nov.(类型地点:委内瑞拉祖利亚埃多省圣阿古斯丁)。提出了15个新的初级主观同义词:Rhopaurus acromelas Lutz和Mello,1922,Rhopaururus melleipalus Lutz和Mello,1922;Rhopalurus pintoi kourouensis Lourenço,2008=Jaguajir pintoi(Mello Leitão,1932);粗尾Rhopaurus crassicauda Caporiacco,1947年,亚马逊Rhopauru amazonicus Lourenço,1986年,和粗尾Rhapaurus crasicauda paruensis Lourenço,2008=宽尾Rhopairus laticoda Thorell,1876年;Rhopaurus melllolitaoi Teruel和Armas,2006,以及Rhopauru aridicola(Teruel and Armas,2012)=异栉水母(Herbst,1800);麻粒Rhopaurus granlimanus Teruel,2006=吉氏异栉水母(Teruel)(2006);Rhopalurus virkii Santiago Blay,2009=阿布迪异栉水母(Armas和Marcano Fondeur,1987);Rhopalurus brejo Lourenço,2014=Trogloropalurus lacrau(Lourenço和Pinto da Rocha,1997)。
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引用次数: 43
Chapter 2: Inoceramid Bivalves from the Coniacian and Basal Santonian (Upper Cretaceous) of the Western Canada Foreland Basin 第二章:加拿大西部前陆盆地上白垩纪Coniacian和基底三东统的双壳类
IF 3.4 2区 环境科学与生态学 Q1 Agricultural and Biological Sciences Pub Date : 2017-06-26 DOI: 10.1206/0003-0090-414.1.4
I. Walaszczyk, A. Plint, N. Landman
ABSTRACT Inoceramid bivalves are the dominant invertebrate fauna of the Coniacian and basal Santonian of the Western Canada Foreland Basin in western Alberta. In the upper lower Coniacian through to basal Santonian, six successive faunas are recognized, which provide the basis for corresponding, formally defined inoceramid zones. From bottom upward these are the zones of: Cremnoceramus crassus crassus /C. deformis deformis, Inoceramus gibbosus, Volviceramus koeneni, Volviceramus involutus, Sphenoceramus subcardissoides, and Sphenoceramus ex gr. pachti. Particular faunas represent assemblages known widely from the Euramerican biogeographic region, although they characterize mostly its northern, boreal area. The inoceramid-based biostratigraphic scheme allows correlation with other parts of the North American Western Interior and with parts of the Euramerican biogeographic region. The studied succession provides a good record of the Inoceramus gibbosus Zone, which characterizes the topmost lower Coniacian. This zone, first recognized from northern Germany, is usually absent, both in Europe and in North America, due to a stratigraphic gap resulting from a eustatic lowstand. The base of the middle Coniacian is marked by the abrupt appearance of the taxonomically variable Volviceramus fauna (V. koeneni (Müller), V. exogyroides (Meek and Hayden)), with associated Inoceramus undabundus Meek and Hayden and Volviceramus cardinalensis, newly described herein. Scaphites (Scaphites) ventricosus Meek and Hayden, the ammonite marker of the base of the middle Coniacian first appears in the late early Coniacian. The base of the upper Coniacian marks the first appearance of the characteristic northern inoceramid species Sphenoceramus subcardissoides (Schlüter), the appearance of which coincides with Scaphites (Scaphites) depressus Reeside, the ammonite marker of this boundary. Close to this boundary Volviceramus stotti also appears, which is newly described from the Canadian sections. The base of the Santonian corresponds to the abrupt appearance of Sphenoceramus ex gr. pachti (Arkhangelsky). The studied sections demonstrate that the appearance of new inoceramid faunas (lowest occurrence of Cremnoceramus crassus crassus (Petrascheck), of various species of Volviceramus, Sphenoceramus subcardissoides (Schlüter) and of S. ex gr. pachti) takes place immediately above major marine flooding surfaces, suggesting a close correspondence between evolutionary and/or migration events and episodes of relative sea-level rise.
摘要:在艾伯塔省西部加拿大西部前陆盆地的科尼亚阶和基底桑托阶,无神经酰胺双壳类是主要的无脊椎动物。在上下科尼亚阶到基底桑托阶,识别出六个连续的动物群,这为相应的、正式定义的无神经酰胺带提供了基础。从下到上是:Cremnoceramus crassus crassus/C的区域。deformis deformis、Inoceramus gibbosus、Volviceramus koeneni、Volvicera mus involutus、Sphenoceraus subcardisoides和Sphenocreamus ex gr.pachti。特定的动物群代表了在欧美生物地理区域广泛已知的群落,尽管它们主要是其北部北方地区的特征。基于无机神经酰胺的生物地层学方案允许与北美西部内陆的其他地区和欧美生物地理学区域的部分地区进行对比。所研究的演替提供了一个很好的记录Inoceramus gibbosus带,它是最上层的下科尼亚阶的特征。这一区域最早在德国北部被发现,在欧洲和北美通常都不存在,因为海平面低水位造成了地层间隙。中科尼亚纪的基底以分类可变的Volviceramus动物群(V.koeneni(Müller)、V.exogryoides(Meek和Hayden))的突然出现为标志,以及本文新描述的相关的undabundus Meek和Haiden和Volviceramus cardinadensis。Scaphites(Scaphites)ventricosus Meek和Hayden,中科尼亚纪基底的菊石标志物最早出现在科尼亚期晚期和早期。上科尼亚阶的基底标志着北方特有的无神经酰胺物种亚心形鞘氨醇酰胺(Schlüter)的首次出现,其出现与该边界的菊石标志物Scaphites(Scaphites)depressus Reeside重合。在这个边界附近也出现了Volviceramus stotti,这是加拿大部分最新描述的。Santonian的基底对应于Sphenoceramus ex gr.pachti(阿尔汉格尔斯基)的突然出现。所研究的剖面表明,新的无神经酰胺动物群的出现(Cremnoceramus crassus crassus(Petrascheck)、Volviceramus、Sphenoceramussubcardisoides(Schlüter)和S.ex gr.pachti的各种物种的出现率最低)发生在主要的海洋洪泛面上方,表明进化和/或迁徙事件与海平面相对上升事件之间存在密切对应关系。
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引用次数: 4
Chapter 3: Scaphitid Ammonites from the Upper Cretaceous (Coniacian-Santonian) Western Canada Foreland Basin 第三章:加拿大西部前陆盆地上白垩世(coniian - santonian)的舟形菊石
IF 3.4 2区 环境科学与生态学 Q1 Agricultural and Biological Sciences Pub Date : 2017-06-26 DOI: 10.1206/0003-0090-414.1.1
N. Landman, A. Plint, I. Walaszczyk
ABSTRACT The Upper Cretaceous (Coniacian-Santonian) of the Western Canada Foreland Basin, contains a rich record of scaphitid ammonites (scaphites). We describe four species: Scaphites (Scaphites) preventricosus Cobban, 1952, Scaphites (S.) ventricosus Meek and Hayden, 1862, Scaphites (S.) depressus Reeside, 1927, and Clioscaphites saxitonianus (McLearn, 1929). These are widespread index fossils that demarcate the upper lower-middle, middle, and upper Coniacian, and the lower Santonian, respectively. They occur in the lower part of the Wapiabi Formation, Alberta. The Coniacian part of the section has been divided into 24 informal allomembers based on the recognition of marine flooding surfaces, most of which can be traced through the >750 km extent of the study area. The most distinctive feature in the ontogenetic development of scaphites is the change in coiling during ontogeny. At the approach of maturity, the shell uncoils slightly, forming a shaft, which then recurves backward approaching the earlier secreted phragmocone. However, this sequence of scaphites shows an evolutionary trend toward recoiling, accompanied by an increase in size and degree of depression. These changes occurred against a background of changing environmental conditions resulting from the expansion of the Western Interior Seaway during the Niobrara transgression. This resulted in an increase in the area of offshore habitats, which may have promoted the appearance of larger species with more depressed whorl sections. Scaphites probably lived at depths of less than 100 m, and may have fed on small organisms in the water column.
摘要加拿大西部前陆盆地的上白垩纪(Coniacian-Santonian)有丰富的舟状菊石记录。我们描述了四个物种:防纹石斑石(Scaphites)preventricosus Cobban,1952年,心室石斑石和海登石斑石,1862年,凹陷石斑石Reeside,1927年,和沙氏石斑石。这些是广泛分布的索引化石,分别划分了上下中、中、上科尼亚阶和下桑托阶。它们产于阿尔伯塔省瓦皮亚比组的下部。根据对海洋洪泛面的识别,该剖面的科尼亚阶部分被划分为24个非正式的同种成员,其中大部分可以追溯到研究区域>750km的范围内。舟虫个体发育最显著的特征是在个体发育过程中卷曲的变化。成熟时,外壳轻微展开,形成轴,然后向后弯曲,接近早期分泌的隔膜。然而,这一序列的舟骨显示出向后退缩的进化趋势,同时伴随着体型和抑郁程度的增加。这些变化是在Niobrara海侵期间西部内部航道扩张导致环境条件变化的背景下发生的。这导致了近海栖息地面积的增加,这可能促进了具有更多凹陷轮生部分的较大物种的出现。Scaphites可能生活在不到100米的深度,可能以水柱中的小生物为食。
{"title":"Chapter 3: Scaphitid Ammonites from the Upper Cretaceous (Coniacian-Santonian) Western Canada Foreland Basin","authors":"N. Landman, A. Plint, I. Walaszczyk","doi":"10.1206/0003-0090-414.1.1","DOIUrl":"https://doi.org/10.1206/0003-0090-414.1.1","url":null,"abstract":"ABSTRACT The Upper Cretaceous (Coniacian-Santonian) of the Western Canada Foreland Basin, contains a rich record of scaphitid ammonites (scaphites). We describe four species: Scaphites (Scaphites) preventricosus Cobban, 1952, Scaphites (S.) ventricosus Meek and Hayden, 1862, Scaphites (S.) depressus Reeside, 1927, and Clioscaphites saxitonianus (McLearn, 1929). These are widespread index fossils that demarcate the upper lower-middle, middle, and upper Coniacian, and the lower Santonian, respectively. They occur in the lower part of the Wapiabi Formation, Alberta. The Coniacian part of the section has been divided into 24 informal allomembers based on the recognition of marine flooding surfaces, most of which can be traced through the >750 km extent of the study area. The most distinctive feature in the ontogenetic development of scaphites is the change in coiling during ontogeny. At the approach of maturity, the shell uncoils slightly, forming a shaft, which then recurves backward approaching the earlier secreted phragmocone. However, this sequence of scaphites shows an evolutionary trend toward recoiling, accompanied by an increase in size and degree of depression. These changes occurred against a background of changing environmental conditions resulting from the expansion of the Western Interior Seaway during the Niobrara transgression. This resulted in an increase in the area of offshore habitats, which may have promoted the appearance of larger species with more depressed whorl sections. Scaphites probably lived at depths of less than 100 m, and may have fed on small organisms in the water column.","PeriodicalId":50721,"journal":{"name":"Bulletin of the American Museum of Natural History","volume":null,"pages":null},"PeriodicalIF":3.4,"publicationDate":"2017-06-26","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"https://sci-hub-pdf.com/10.1206/0003-0090-414.1.1","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"45408924","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":2,"RegionCategory":"环境科学与生态学","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
引用次数: 7
Allostratigraphy and Biostratigraphy of the Upper Cretaceous (Coniacian-Santonian) Western Canada Foreland Basin 加拿大西部前陆盆地上白垩世(coniian - sanantonian)的异位地层学和生物地层学
IF 3.4 2区 环境科学与生态学 Q1 Agricultural and Biological Sciences Pub Date : 2017-06-26 DOI: 10.1206/0003-0090-414.1.2
N. Landman, A. Guy Plint, I. Walaszczyk
172 pages, 15 folded leaves of plates : illustrations (some color), maps (some color) ; 26 cm. chapter 1. Integrated, high-resolution allostratigraphic, biostratigraphic and carbon-isotope correlation of Coniacian strata (Upper Cretaceous), western Alberta and northern Montana / A. Guy Plint, Elizabeth A. Hooper, Meriem D. Grifi, Ireneusz Walaszczyk, Neil H. Landman, Darren R. Grocke, Joao P. Trabucho Alexandre, and Ian Jarvis -- chapter 2. Inoceramid bivalves from the Coniacian and basal Santonian (Upper Cretaceous) of the Western Canada Foreland Basin / Ireneusz Walaszczyk, A. Guy Plint, and Neil H. Landman -- chapter 3. Scaphitid ammonites from the Upper Cretaceous (Coniacian-Santonian) Western Canada Foreland Basin / Neil H. Landman, A. Guy Plint, and Ireneusz Walaszczyk.
172页,15页折页:插图(有些颜色)、地图(有些颜色的);26厘米。第1章。Coniacian地层(上白垩纪)、阿尔伯塔省西部和蒙大拿州北部/A的综合高分辨率异地层、生物地层学和碳同位素对比。Guy Plint、Elizabeth A.Hooper、Meriem D.Grifi、Ireneusz Walaszczyk、Neil H.Landman、Darren R.Grocke、Joao P.Trabucho Alexandre和Ian Jarvis——第2章。加拿大西部Foreland盆地/Ireneusz-Walaszczyk、A.Guy Plint和Neil H.Landman的Coniacian和基底Santonian(上白垩纪)的无神经酰胺双壳类——第3章。加拿大西部前陆盆地上白垩纪(Coniacian-Santonian)的Scaphitid菊石/Neil H.Landman、A.Guy Plint和Ireneusz-Walaszczyk。
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引用次数: 0
Osteology of The Middle Eocene Ceratomorph Hyrachyus modestus (Mammalia, Perissodactyla) 中始新世角化兽(哺乳目,准足趾目)的骨学
IF 3.4 2区 环境科学与生态学 Q1 Agricultural and Biological Sciences Pub Date : 2017-06-09 DOI: 10.1206/0003-0090-413.1.1
B. Bai, Jin Meng, Yuanqing Wang, Haibing Wang, L. Holbrook
ABSTRACT The middle Eocene ceratomorph Hyrachyus has been considered a pivotal genus in ceratomorph evolution, either as a transitional form from tapiroids to rhinocerotoids, giving rise to all later rhinocerotoids, or else as the sister taxon to other rhinocerotoids. Thus, Hyrachyus has been commonly chosen as an outgroup in phylogenetic analyses of rhinocerotoids. However, little has been published on the osteology of Hyrachyus, even though well-preserved craniodental and postcranial specimens of this taxon have been in collections for decades. Here, we describe and illustrate the cranial and postcranial osteology of Hyrachyus modestus, based mainly on the exceptionally preserved specimens housed at the American Museum of Natural History, specifically AMNH FM 12664. Our bone-by-bone description provides detailed information on the osteological morphology of Hyrachyus, which should be useful for phylogenetic analyses of both rhinocerotoids and perissodactyls in general, because it provides one of the more complete and best-preserved examples of the skeleton of an earlier Eocene perissodactyl. The cranial morphology of Hyrachyus modestus shows a shallow narial notch, a lacrimal contacting the nasal, and a sphenorbital fissure closely situated to the anterior opening of the alisphenoid canal. In the basicranial region, there is a mastoid exposure of the petrosal between the occipital and the squamosal, and the posttympanic process and paracondylar process are partly fused. The postcranial morphology of Hyrachyus modestus includes the following features: The cervical region of the vertebral column is relatively short compared to the rest of the vertebral column. The lumbar vertebrae have concave-convex embracing prezygapophyses and postzygapophyses. The scapula has a distinct acromion process. The humerus has a greater tubercle that does not elevate above the head, and the deltoid tuberosity and deltopectoral crest are weak. The scaphoid and lunar facets of the radius are confluent. The olecranon of the ulna extends posteroproximally. The manus is functionally tetradactyl, with a complete fifth manual digit. The innominate bone has a long, narrow coxal tuberosity. The greater trochanter of the femur is elevated proximally above the head. The femur has a long, narrow, and symmetric trochlea. The patella has a moderately anteroposteriorly deep base. The intercondyloid eminences of the tibia are equal in height, and the extensor sulcus of the tibia is relatively deep. The fibula has a relatively slender shaft with expanded ends. The pes has three functional digits. The calcaneus does not contact the navicular, nor does the Mt III contact the cuboid. Comparisons between the skeleton of Hyrachyus modestus and those of the early tapiroid Heptodon, the hyracodontid Triplopus, the paraceratheriid Juxia, and the rhinocerotid Uintaceras were also investigated. These results indicate that Hyrachyus probably did not derive from Heptodon, but from a more basal
中始新世角鼻虫Hyrachyus被认为是角鼻虫进化过程中的一个关键属,它是角鼻虫从类貘到类鼻虫的过渡形式,产生了所有后来的类鼻虫,或者是其他类鼻虫的姐妹分类群。因此,在类犀牛的系统发育分析中,水螅类通常被选为外群。然而,尽管保存完好的颅齿和颅后标本已经被收集了几十年,但关于水龙的骨学研究却很少发表。在这里,我们主要基于美国自然历史博物馆保存的特殊标本,特别是AMNH FM 12664,描述和说明了水龙的颅骨和颅后骨学。我们对每块骨头的描述提供了水龙骨形态的详细信息,这对于类鼻兽和一般的异趾龙的系统发育分析都是有用的,因为它提供了始新世早期异趾龙骨骼的一个更完整和保存最好的例子。小圆鼻蛇的颅骨形态为浅鼻切迹,泪道与鼻腔相连,蝶眶裂紧靠翼蝶骨管的前开口。在颅底区,枕骨和鳞骨之间的岩骨有乳突暴露,鼓室后突和髁旁突部分融合。后颅形态包括以下特征:与脊柱的其余部分相比,脊柱的颈椎区域相对较短。腰椎呈凹凸状,包括肩关节前和肩关节后。肩胛骨有明显的肩峰突。肱骨有一个不高于头部的大结节,三角结节和三角胸峰较弱。桡骨的舟状面和月状面汇合。尺骨鹰嘴后近端延伸。手在功能上是四指形的,有完整的第五个手指。无名骨有一长而窄的尾结节。股骨大转子在头部上方近端升高。股骨有一个长、窄、对称的滑车。髌骨有一个中等正后方深的基底。胫骨髁间突高度相等,胫骨伸肌沟相对较深。腓骨有一个相对细长的轴,末端扩张。pes有三个功能性数字。跟骨不接触舟骨,第三股肌也不接触长方体。本文还比较了普通水龙与早期tapioid Heptodon、hyracodontid Triplopus、paraceroid Juxia和rhinocerotid untaceras的骨骼。这些结果表明Hyrachyus可能不是来自七齿兽,而是来自一个更基础的角兽类。此外,水龙与水龙科最早的代表——三足龙的骨骼差异表明,水龙不是水龙的后代。然而,类水螅类的祖先可能产生了其他非水螅类的类鼻。和其他始新世的异趾龙一样,普通蹄龙的后颅形态显示出适应性,这表明在异趾龙进化的早期就已经出现了移动运动。
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引用次数: 13
Phylogenetic Relationships of a New Genus of Calliopsine Bees from Peru, with a Review of Spinoliella Ashmead (Hymenoptera: Andrenidae) 秘鲁小蜂一新属的系统发育关系——兼论小蜂属(膜翅目:蜂科)
IF 3.4 2区 环境科学与生态学 Q1 Agricultural and Biological Sciences Pub Date : 2017-04-13 DOI: 10.1206/0003-0090-412.1.1
V. González, Allan H. Smith-Pardo, M. Engel
ABSTRACT We establish a new genus for an unusual species of Peruvian calliopsine bees (Panurginae: Calliopsini) that was initially reported in the literature as an undescribed species of Spinoliella Ashmead that purportedly expanded the range of the latter genus beyond Argentina and Chile. Although the new genus superficially resembles Spinoliella, it is easily distinguished by a unique combination of characters in both sexes but particularly in the male hidden metasomal sterna and genitalia. A cladistic analysis of 82 adult external morphological characters including all species of Spinoliella, as well as species of the remaining genera of Calliopsini, suggests that this group is sister to a clade consisting of Spinoliella and Callonychium Brèthes. We describe and illustrate Xeranthrena imponticula Gonzalez and Engel, new genus and species, from males and females collected in xeric areas along the Pacific slopes of the Peruvian Andes. In addition, the phylogenetic study suggests two well-defined clades within Spinoliella and corresponding to the previously recognized subgenera, although we do not advocate for their reinstatement. We briefly discuss new putative synapomorphies for Spinoliella and, building upon prior revisionary work, we describe and figure five new species: Spinoliella aidae Gonzalez, Smith-Pardo, and Engel, new species; S. confusa Gonzalez and Engel, new species; S. propinqua Gonzalez and Engel, new species; S. packeri Gonzalez and Engel, new species; and S. polita Gonzalez and Engel, new species. In addition, we synonymize S. karhadra Rodríguez, Toro, and Ruz under S. rufiventris Toro and Ruz (new synonymy). We provide new geographical and floral records, an identification keys to all 17 recognized species of Spinoliella, and updated key to the genera of Calliopsini.
摘要:我们为一种罕见的秘鲁Calliopsini蜜蜂(Panurginae: Calliopsini)建立了一个新属,该属最初在文献中被报道为Spinoliella Ashmead的一个未被描述的物种,据称将后者的范围扩大到阿根廷和智利以外。虽然新属表面上类似于Spinoliella,但它很容易通过两性的独特特征组合来区分,特别是在雄性隐藏的跖骨和生殖器上。通过对Spinoliella所有种和Calliopsini剩余属的82个成虫外部形态特征的分支分析,认为该类群与Spinoliella和Callonychium br组成的分支是姊妹关系。本文描述并举例说明了在秘鲁安第斯山脉太平洋斜坡的干旱地区采集的雄性和雌性Xeranthrena imponticula Gonzalez和Engel这两个新属和新种。此外,系统发育研究表明,Spinoliella中有两个明确的分支,对应于先前公认的亚属,尽管我们不主张恢复它们。我们简要地讨论了Spinoliella的新推测的突触形态,并在先前修订工作的基础上,我们描述和绘制了五个新种:Spinoliella aidae Gonzalez, Smith-Pardo和Engel,新种;conusa Gonzalez和Engel,新种;S. propinqua Gonzalez和Engel,新种;S. packeri Gonzalez和Engel,新种;S. polita Gonzalez和Engel是新种。此外,我们将S. karhadra Rodríguez, Toro和Ruz同义化在S. rufiventris Toro和Ruz(新同义词)下。我们提供了新的地理和植物记录、Spinoliella所有17个已知种的鉴定密钥和Calliopsini属的更新密钥。
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引用次数: 3
New Specimens of Anchiornis huxleyi (Theropoda: Paraves) from the Late Jurassic of Northeastern China 中国东北晚侏罗世赫胥黎近鸟龙新种(兽足目:兽足目)
IF 3.4 2区 环境科学与生态学 Q1 Agricultural and Biological Sciences Pub Date : 2017-04-13 DOI: 10.1206/0003-0090-411.1.1
R. Pei, Quanguo Li, Qingjin Meng, M. Norell, K. Gao
ABSTRACT Four new specimens of Anchiornis huxleyi (PKUP V1068, BMNHC PH804, BMNHC PH822, and BMNHC PH823) were recently recovered from the Late Jurassic fossil beds of the Tiaojishan Formation in northeastern China. These new specimens are almost completely preserved with cranial and postcranial skeletons. Morphological features of Anchiornis huxleyi have implications for paravian character evolution and provide insights into the relationships of major paravian lineages. Anchiornis huxleyi shares derived features with avialans, such as a straight nasal process of the premaxilla and the absence of an external mandibular fenestra in lateral view. However, Anchiornis huxleyi lacks several derived deinonychosaurian features, including a laterally exposed splenial and a specialized raptorial pedal digit II. Morphological comparisons strongly suggest Anchiornis is more closely related to avialans than to deinonychosaurians or troodontids. Anchiornis huxleyi exhibits many conservative paravian features, and closely resembles Archaeopteryx and other Jurassic paravians from Jianchang County, such as Xiaotingia and Eosinopteryx. The other Jianchang paravian, Aurornis xui, is likely a junior synonym of Anchiornis huxleyi.
摘要最近在中国东北晚侏罗世条集山组化石层中发现了4个新的赫胥黎近鸟龙标本(PKUP V1068、BMNHC PH804、BMNHC PH822和BMNHC PH823)。这些新标本几乎完整地保存了颅骨和颅骨后骨骼。赫胥黎近鸟龙的形态特征对类人猿的特征演化具有重要意义,并对类人猿主要世系的关系提供了新的认识。赫胥黎近鸟龙与鸟类具有相同的衍生特征,如上颌骨前鼻突直,侧面无下颌外窗。然而,赫胥黎近鸟龙缺乏几个衍生的恐爪龙特征,包括外侧暴露的脾脏和专门的猛禽趾II。形态学比较强烈地表明,近鸟龙与鸟类的亲缘关系比与恐爪龙或troodontids的亲缘关系更近。赫胥黎近鸟鸟表现出许多保守的类鸟特征,与始祖鸟和建昌县的其他侏罗纪类鸟(如晓廷鸟和伊红鸟鸟)非常相似。另一种剑昌类鸟,许氏Aurornis xuui,可能是近鸟龙赫胥黎的低级同义词。
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引用次数: 41
Taxonomic Revision of the Jumping Goblin Spiders of the Genus Orchestina Simon, 1882, in the Americas (Araneae: Oonopidae) 美洲1882年Orchestina Simon属跳蛛的分类学修订(蜘蛛目:卵蛛科)
IF 3.4 2区 环境科学与生态学 Q1 Agricultural and Biological Sciences Pub Date : 2017-02-22 DOI: 10.1206/0003-0090-410.1.1
M. Izquierdo, M. Ramírez
ABSTRACT The genus Orchestina Simon is distributed worldwide and is characterized by having an enlarged fourth femur with which these species are capable of jumping. The genus is also characterized by having a well-sclerotized sperm duct, a near H-shaped arrangement of the eye group, a 4-4-3-3 pattern of raised receptors on the tarsal organs of the legs I–IV, respectively, and by lacking spines on all the legs. All these characters, together with molecular evidence, support the monophyly of the genus, as well as its placement as sister group of Oonopinae. Most American species of Orchestina inhabit the forest foliage and canopy, but in unforested areas they also occur in shrubs and grasses. In this work, we revise the American representatives of Orchestina in a comprehensive context for the first time. In the past, six species were known from the Americas: five from the United States and only one from South America, described from Venezuela. After the study of the principal collections of the world and several field trips to several South American countries, we describe 85 new species and redescribe all previously known species. Matching sexes was occasionally problematic; while females are very homogeneous in somatic traits, males may have modifications on different parts of the body, making the matching very difficult. Therefore, in this review 56 of the species are described from only one sex, whereas 20 unmatched species are informally described as morphospecies, pending the discovery of conspecific sexes. Two species, O. pavesiiformis Saaristo and O. dentifera Simon, originally known from Israel and Sri Lanka, respectively, are here reported as introduced in several countries in the Americas and other continents. O. justini Saaristo described from the Seychelles is here considered a synonym of O. dentifera. One species, O. truncata Wunderlich, previously known as a subfossil spider from Colombian Copal is here tentatively redescribed based on recent material from Costa Rica, Colombia, and Ecuador; the female is also described for the first time. The species list by country is as follows (numbers refer to records, independently of the locality of the type material): United States (9 species: O. utahana Chamberlin and Ivie, O. moaba Chamberlin and Ivie, O. obscura Chamberlin and Ivie, O. saltitans Banks, O. nadleri Chickering, the introduced O. pavesiiformis Saaristo, and three new species, O. quasimodo, O. kamehameha, and O. auburndalensis); Mexico (3 species: O. utahana Chamberlin and Ivie, and two new species, O. nahuatl and O. chaparrita); Guatemala (1 new species: O. guatemala); Costa Rica (3 new species: O. laselva, O. griswoldi, and O. chiriqui; and the previously known O. truncata Wunderlich); Panama (5 new species: O. chiriqui, O. labarquei, O. pan, O. campana, and O. galapagos); Jamaica (2 species, the introduced O. dentifera Simon and O. galapagos); Haiti and Dominican Republic (only the introduced O. dentifera Simon); Colombia (6 n
西蒙管弦乐队属分布在世界各地,其特点是具有扩大的第四股骨,这些物种能够跳跃。该属的另一个特点是精子管硬化良好,眼群呈近h形排列,在腿I-IV的跗骨器官上分别有4-4-3-3型凸起的受体,所有的腿上都没有刺。所有这些特征,连同分子证据,都支持该属的单系性,以及它作为乌纳科姐妹类群的位置。大多数美洲的兰花栖息在森林的叶子和树冠上,但在非森林地区,它们也出现在灌木和草中。在这项工作中,我们第一次在一个全面的背景下修订了美国的管弦乐队代表。过去,美洲已知的有6种:5种来自美国,只有1种来自南美,来自委内瑞拉。经过对世界主要物种的研究和对几个南美国家的实地考察,我们描述了85个新种,并重新描述了所有已知的物种。性别匹配有时会出现问题;虽然雌性在身体特征上非常相似,但雄性在身体的不同部位可能会有变化,这使得匹配非常困难。因此,在这篇综述中,56个物种被描述为只有一个性别,而20个不匹配的物种被非正式地描述为形态种,等待发现同种性别。两个物种,O. pavesiiformis Saaristo和O. dentifera Simon,最初分别来自以色列和斯里兰卡,在这里被报道为在美洲和其他大陆的几个国家引进。从塞舌尔描述的O. justini Saaristo在这里被认为是O. dentifera的同义词。其中一种,O. truncata Wunderlich,以前被认为是哥伦比亚Copal的亚化石蜘蛛,现在根据哥斯达黎加、哥伦比亚和厄瓜多尔的最新资料进行了初步的重新描述;雌性也首次被描述。美国(9种:O. utahana Chamberlin和Ivie, O. moaba Chamberlin和Ivie, O. obscura Chamberlin和Ivie, O. saltitans Banks, O. nadleri Chickering,引进的O. pavesiiformis Saaristo, 3种新种:O. quasimodo, O. kamehameha和O. auburndalensis);墨西哥(3种:O. utahana Chamberlin和Ivie, 2新种:O. nahuatl和O. chaparrita);危地马拉(1新种:O. Guatemala);哥斯达黎加(3新种:O. laselva、O. griswoldi和O. chiriqui;以及之前已知的O. truncata Wunderlich);巴拿马(5新种:O. chiriqui、O. labarquei、O. pan、O. campana和O. galapagos);牙买加(2种,引进的O. dentifera Simon和O. galapagos);海地和多米尼加共和国(只引进了O. dentifera Simon);哥伦比亚(6新种:O. filandia、O. zingara、O. arboleda、O. cali、O. platnicki、O. pakitza;和O. truncata Wunderlich,加上形态种OMI020和OMI038);委内瑞拉(7种:O. saltabunda;新增6个(O. venezuela, O. aragua, O. bolivar, O. maracay, O. ranchogrande和O. neblina);特立尼达和多巴哥(1新种:O. kairi);圭亚那(1种形态:OMI026);厄瓜多尔(18个新种:O. galapagos、O. fernandina、O. erwini、O. ecuatoriensis、O. sotoi、O. magna、O. shuar、O. golem、O. waorani、O. tzantza、O. predator、O. goblin、O. yanayacu、O. otonga、O. santodomingo、O. quijos、O. mayo、O. laselva和O. truncata Wunderlich,以及形态种OMI020、OMI021、OMI022、OMI023、OMI024、OMI025、OMI026、OMI027、OMI029、OMI036和OMI038);秘鲁(9新种:O. cajamarca、O. comaina、O. atocongo、O. mancocapac、O. silvae、O. madrededios、O. pakitza、O. losamigos、O. golem,形态种OMI020、OMI023、OMI026、OMI030);巴西(25个新种;O. cristinae、O. coari、O. moura、O. valquiria、O. aproeste、O. caxiuana、O. para、O. taruma、O. retiro、O. divisor、O. juruti、O. platnicki、O. iemanja、O. bonaldoi、O. rapaz、O. itapety、O. catarina、O. leon、O. saudade、O. sarava、O. goblin、O. sotoi、O. golem、O. waorani、O. ucumar、引进的O. pavesiiformis Saaristo和O. dentifera Simon,以及形态物种OMI021、OMI023、OMI024、OMI025、OMI026、OMI027、OMI029、OMI032、OMI033、OMI034、OMI035、OMI036、OMI037、OMI039,和OMI040);玻利维亚(3新种:O. moyuchi、O. grismadoi和O. ucumar);智利(12个新种:O. pandeazucar、O. caleta、O. totoralillo、O. pizarroi、O. molles、O. granizo、O. quenies、O. curico、O. cachai、O. nahuelbuta、O. jaiba和O. osorno);阿根廷(6个新种:O. ucumar、O. andianavarroi、O. luispi、O. cristinae、O. platnicki、O. jaiba和引进的O. pavesiiformis Saaristo)。 虽然该属的内部关系仍不清楚,但我们根据形态特征和地理分布提出了一些种群。来自美国西部的物种都有网状的颜色图案,在一些物种中,雄性的甲壳或第一腿也有变化。这一组雌性的特点是存在一个薄的,通常长和扭曲的前托和外部口袋在腹壁区域。这个群体居住在一个地理区域,在白垩纪晚期仍然是孤立的,并且仍然受到地理和气候条件的隔离。考虑到南美洲成员的广泛多样性,它们的物种群更难建立。然而,有些种类是独特的,在雌性生殖器内有口袋,在雄性交配球上有突起。智利种可能是单系群;雌虫的上腹部有外袋和脊状突起,与其他雌虫区分开来,而雄虫的交媾球形态非常相似,非常简单,有长栓子,有时有额外的棘状突。这一群体可能在安第斯山脉上升期间和之后经历了与世隔绝的事件。和这个科的其他成员一样,这个属的有趣之处在于雄性的第二性征的存在,而雌性的体细胞特征实际上是同质的。然而
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引用次数: 13
Diverse Orthorrhaphan Flies (Insecta: Diptera: Brachycera) in Amber From the Cretaceous of Myanmar: Brachycera in Cretaceous Amber, Part VII 缅甸白垩纪琥珀中的各种直蝇(昆虫纲:双翅目:短肢目):白垩纪琥珀中的短肢目,第七部分
IF 3.4 2区 环境科学与生态学 Q1 Agricultural and Biological Sciences Pub Date : 2016-10-13 DOI: 10.1206/0003-0090-408.1.1
D. Grimaldi
ABSTRACT A remarkable diversity of new nonempidoid orthorrhaphan flies from the mid-Cretaceous of Myanmar (Late Albian–Early Cenomanian, ca. 99 Ma) is presented, including 28 species (all but one new) in 22 genera (13 new), and at least 12 families. Two families are new; three genera are unplaced in Tabanomorpha and one unplaced within Brachycera. Comparisons are presented between the amber taxa and extensive lithified taxa from the Jurassic and Cretaceous of eastern Laurasia. In Stratiomyomorpha: A new species of Zhangsolvidae has color patterns and body shape that apparently mimic Vespidae or other stinging aculeate wasps. Diverse new Xylomyidae and Stratiomyidae are described, the latter with male terminalia preserved in detail. In Tabanomorpha: The genus Athericites Mostovski et al. is synonymized with Palaepangonius Ren, and a new species of Galloatherix Nel is described in which the female proboscis is much longer than that of the male. All three genera were attributed to Athericidae, but the amber species reveal they are more basal tabanomorphs. Described are a new genus of stem-group Tabanoidea and a new species of Cratotabanus Martins-Neto and Kucero-Santos (Tabanidae), previously known from the Cretaceous of Brazil and New Jersey. In Nemestrinoidea: Three species of Hirmoneura Meigen are the first Nemestrinidae known from amber; one species has long cerci typical of Recent species of the genus. A new species of the Mesozoic family Rhagionemestriidae reveals this family is closely related to Acroceridae, not Xylophagidae as previously proposed. In Archisargoidea: A new species of Tethepomyiidae is described, the family known only in Cretaceous amber from New Jersey, Spain, and Myanmar. In Asiloidea: Three new species and genera of Bombyliidae are described, two of them with abdominal setal “baskets” distinctive to females of higher bombyliids. A fourth new asiloid genus is a probable stem-group bombyliid. The recently described Pseudorhagio Zhang et al., is transferred from Tabanomorpha to Bombyliidae. The male of a new species of Burmapsilocephala Gaimari and Mostovski (Apsilocephalidae) has terminalia very similar to that of Evocoa (Evocoidae: Recent, Chile), corroborating close relationship of the two families. In Families incertae sedis: A new species of Hilarimorphites Grimaldi and Cumming (Apystomyiidae) is described, the family known only in New Jersey and Burmese amber and the Recent fauna of California, and a sister group to either Cyclorrhapha or Eremoneura. Mysteromyiidae and Eucaudomyiidae, new families, are described, based on highly modified species with vestigial mouthparts, reduced venation, and unique specializations. Discussion is presented on some morphological features in fossil and Recent Brachycera that have apparent phylogenetic significance but are rarely discussed. This study adds further data to the phenotypic diversity, geological history, and biogeography of a major radiation.
摘要:本文报道了缅甸中白垩纪(晚阿尔及利亚-早塞诺曼尼亚,约99 Ma)的非拟正翅目蝇的新多样性,包括22属(13新种)的28种(除1新种外),至少12科。两个家庭是新的;在Tabanomorpha中有三个属未被安置,在Brachycera中有一个属未被安置。对月桂岛东部侏罗纪和白垩纪的琥珀分类群与广泛的石化分类群进行了比较。层肌蜂科:掌形蜂科的一新种,其颜色图案和体型明显模仿胡蜂科或其他有刺刺的胡蜂。描述了多种新的木蝇科和层蝇科,其中层蝇科的雄性终端被详细保存。在Tabanomorpha中:Athericites Mostovski等人将其属与Palaepangonius Ren同义,并描述了Galloatherix Nel的一个新种,其中雌性的喙比雄性的长得多。这三个属都属于Athericidae,但琥珀类显示它们是更基本的tabanomorpha。描述了茎群Tabanoidea的一个新属和Cratotabanus Martins-Neto和Kucero-Santos (Tabanidae)的一个新种,以前在巴西和新泽西州的白垩纪已知。在nemestrindae中:三种Meigen Hirmoneura是最早从琥珀中发现的Nemestrinidae;一种有长尾,是本属最近种的典型。中生代Rhagionemestriidae科新种揭示了该科与Acroceridae的亲缘关系,而不是先前提出的Xylophagidae。在Archisargoidea中:描述了一种新的Tethepomyiidae,该科仅在新泽西州,西班牙和缅甸的白垩纪琥珀中发现。纲:描述了家蚕科3个新种和新属,其中2个具有较高级家蚕雌性特有的腹尾“筐”。第四个新的拟虫属可能是茎群拟虫。最近描述的Pseudorhagio Zhang等人,从Tabanomorpha转移到Bombyliidae。一新种Burmapsilocephala Gaimari和Mostovski (apsilcephalidae)的雄性终端与Evocoa (evocoae:最近,智利)非常相似,证实了两科的亲缘关系。在菊科中:描述了一种新的Hilarimorphites Grimaldi和Cumming (apystomiidae),该科仅在新泽西和缅甸琥珀以及加利福尼亚的近期动物群中已知,是cyclophala或Eremoneura的姐妹群。根据口器退化、脉脉减少和独特特化的高度修饰的物种,描述了神秘蝇科和桉蝇科这两个新科。讨论了化石类和近代类腕足类中具有明显系统发育意义但很少被讨论的形态学特征。本研究为主要辐射的表型多样性、地质历史和生物地理学提供了进一步的数据。
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引用次数: 44
Comparative Basicranial Anatomy of Extant Terrestrial and Semiaquatic Artiodactyla 现存陆生和半水生偶蹄动物的比较颅底解剖
IF 3.4 2区 环境科学与生态学 Q1 Agricultural and Biological Sciences Pub Date : 2016-10-07 DOI: 10.1206/0003-0090-409.1.1
M. O'Leary
ABSTRACT Comparative data from the ear region has played an important role in recent combined-data phylogenetic analyses of the relationships of living and extinct Artiodactyla and the position of that clade among Euungulata. These studies have also been important for establishing the phylogenetic position of Cetacea and for understanding the relationships of a diversity of euungulate species to their fossil relatives. Detailed and standardized descriptive reference works of the basicranium for a range of living artiodactylans are not, however, readily available. Here I describe exemplar species from the four major extant terrestrial and semiaquatic artiodactylan clades (Hippopotamidae, Ruminantia, Suina, and Camelidae) and illustrate the anatomy of the ear region with the auditory bulla both in place and removed. Terrestrial artiodactyls exhibit varying degrees of expansion of the bony external acoustic meatus laterally relative to the mediolateral dimensions of the rounded, medial aspect of the auditory bulla, a characteristic that is least developed in Tragulidae. A relatively elongate external acoustic meatus has previously been described as entirely absent in living and fossil cetaceans and in some fossil species such as Diacodexis pakistanensis. Variation also exists in the proximity of the petrosal-bullar complex to midline basicranial bones. Isolation of these bones from other basicranial structures has been previously interpreted as functionally important for underwater hearing in Cetacea. Many artiodactylans have contact between the auditory bulla and the basioccipital but no contact between the deeper pars cochlearis of the petrosal bone and the basioccipital/basisphenoid. Exceptions are species of Hippopotamidae in which both the bulla and the petrosal are separated from midline bones. The functional interpretation of this separation has previously been linked to aquatic hearing, but this association may be more complex than originally thought. Other features observed in the basicrania of terrestrial artiodactylans described here are a general coalescence of basicranial foramina (i.e., the basicapsular fissure, carotid foramen, piriform fenestra, and sometimes the foramen ovale), the development of large and ornate styliform processes in species of Ruminantia, and widespread contact between the auditory bulla and the paracondylar process of the exoccipital.
摘要:近年来,耳部的比较数据在现存和已灭绝偶蹄动物的关系以及偶蹄动物在euungula中的地位的综合系统发育分析中发挥了重要作用。这些研究对于确定鲸目动物的系统发育位置以及了解多种有蹄类动物与其化石近亲的关系也具有重要意义。然而,关于一系列现存偶蹄纲动物的基本纲的详细和标准化的描述性参考著作并不是现成的。在这里,我描述了现存的四种主要陆生和半水生偶蹄动物分支(河马科、反刍动物科、鼩鼱科和骆驼科)的典型物种,并举例说明了耳区在听球存在和移除的情况下的解剖结构。陆生偶蹄动物表现出不同程度的骨外声道的横向扩张,相对于圆形的中外侧尺寸,听球内侧,这一特征在鹰科中最不发达。在现存鲸类和化石鲸类以及一些化石物种(如巴基斯坦双足鲸)中,相对较长的外声通道被描述为完全不存在的。岩石-公牛复合体与中线颅底骨之间也存在差异。这些骨骼与其他颅底结构的分离先前被认为对鲸目动物的水下听力具有重要的功能。许多偶蹄动物在听球和枕基之间有接触,但在岩骨较深的耳蜗部和枕基/枕状骨之间没有接触。例外的是河马科的物种,它们的大骨和岩骨都是从中线骨中分离出来的。这种分离的功能解释以前与水生听觉有关,但这种联系可能比最初想象的要复杂得多。本文所描述的陆生偶蹄动物基底突的其他特征包括颅底孔的普遍合并(即基底囊裂、颈动脉孔、梨状孔,有时还有卵圆孔),Ruminantia种类中大而华丽的花柱状突的发育,以及听球和枕外髁旁突之间的广泛接触。
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引用次数: 6
期刊
Bulletin of the American Museum of Natural History
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