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Brachiopodes toarciens de la coupe d’Aït Athmane, Haut Atlas centro-oriental (Maroc) 来自ait Athmane杯的toarcian腕足类,高阿特拉斯中东部(摩洛哥)
IF 1 4区 地球科学 Q3 Earth and Planetary Sciences Pub Date : 2022-10-01 DOI: 10.1016/j.annpal.2022.102572
Mohamed Benzaggagh , Mariusz A. Salamon , Hanane Khaffou , Touria Hssaida , Mohamed El Ouali , Badre Essafraoui

The Aït Athmane section is located in the eastern part of the central High Atlas, about 20 km north of Errachidia. Its Toarcian series consists of silty marl and silty marly limestone. Its lower part, of early (Levisoni Zone?) to middle Toarcian (Bifrons and Gradata? Zones) age, is rich in brachiopods (rhynchonellides and terebratulides). Its upper part, of late Toarcian age, is rich in terebratulids. Among eighteen reported brachiopod taxa, six are new for the central High Atlas. Their specific assemblages and stratigraphic distributions are similar to those of the Toarcian brachiopods reported from several basins of northern and southern margins of the Tethys and the Subboreal domain, particularly of western France.

Aït Athmane部分位于中部高阿特拉斯的东部,距离erachidia北部约20公里。它的陶里安系列由粉质泥灰岩和粉质泥灰岩组成。它的下半部分,早期(Levisoni带?)到中期(Bifrons和Gradata?)带)年龄大,有丰富的腕足类动物(颈足类动物和无脊椎动物)。它的上部,在晚托尔西亚时代,是丰富的无脊椎动物。在已报道的18个腕足动物分类群中,有6个是新发现的。它们的具体组合和地层分布与特提斯南北缘和亚北方地区(特别是法国西部)几个盆地中报道的托瓦西亚腕足类相似。
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引用次数: 1
Crassatella baudeti (Coquand, 1862) du Cénomanien de Batna (NE algérien) : paléobiogéographie, paléoécologie et biostratigraphie Crassatella baudeti (Coquand, 1862),来自Batna(阿尔及利亚东北部)的cenomanian:古生物地理学、古生态学和生物地层学
IF 1 4区 地球科学 Q3 Earth and Planetary Sciences Pub Date : 2022-07-01 DOI: 10.1016/j.annpal.2022.102569
Riadh Aouissi , Sihem Salmi-Laouar , Izzet Hoşgör

The lower Cenomanian strata from the Batna region (NE Algeria) constitute the richest and most diversified deposits in terms of macro-invertebrates, mainly dominated by bivalves both in terms of specimens and taxa. Among them, Crassatellidae are restricted to the lower Cenomanian stratigraphical levels. They are represented by the single species Crassatella baudeti. The species is easily distinguished by its flattened boat anchor shape and quite strong commarginal lamellae, and is very abundant. It is present in levels ranging from Sharpeiceras schlueteri Subzone to Cunningtoniceras inerme Zone. The species is, therefore, a good marker of the last levels of the lower Cenomanian and the beginning of the midlle Cenomanian. A precise palaeontological description and an environmental discussion were conducted. Thus, Cr. baudeti would be indicative of a shallow environment, with cold but calm waters, normal salinity and moderately soft substrate. Recorded exclusively from the Cenomanian of a few Tethys localities, the exclusive presence of Cr. baudeti reflects a specific thermal gradient and climatic zonation.

Batna地区(阿尔及利亚东北部)的下Cenomanian地层构成了大型无脊椎动物最丰富和最多样化的矿床,在标本和分类群方面主要以双壳类为主。其中,草苔科仅局限于塞诺曼阶地层的下层。它们的代表是单种博德蒂(Crassatella baudeti)。该物种很容易通过其扁平的船锚形状和相当强的近缘片层来区分,并且数量非常丰富。它存在于从Sharpeiceras schlueteri亚带到Cunningtoniceras inme带的各个层次。因此,该物种是下Cenomanian晚期和中Cenomanian早期的一个很好的标志。进行了精确的古生物学描述和环境讨论。因此,鲍德迪河可能是一个浅水环境的标志,具有寒冷但平静的水域,正常的盐度和中等柔软的基质。仅从特提斯的几个地方的塞诺曼尼亚记录,Cr. baudeti的独特存在反映了特定的热梯度和气候带。
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引用次数: 2
The heteromorph ammonite genus Ammonitoceras Dumas, 1876 (Ancyloceratidae) in the lower Aptian (Lower Cretaceous) of the Les Ferres Aptian Basin (southeastern France) 法国东南部Les Ferres Aptian盆地下阿普提亚(下白垩世)的异形菊石属Ammonitoceras Dumas, 1876(凤尾虫科)
IF 1 4区 地球科学 Q3 Earth and Planetary Sciences Pub Date : 2022-07-01 DOI: 10.1016/j.annpal.2022.102570
Stéphane Bersac , Didier Bert

Ammonitoceras Dumas, 1876 is a genus of heteromorph ammonites characterized by the presence of a peculiar ontogenetic stage in its inner whorls: the Ammonitoceras stage. But in spite of its wide paleogeographic and biostratigraphic extension throughout the Aptian (Lower Cretaceous), this genus remains poorly known. In the present work we study specimens of Ammonitoceras from the lower Aptian Deshayesites multicostatus (Deshayesites deshayesi Zone) to Dufrenoyia furcata (Dufrenoyia furcata Zone) subzones of the Les Ferres Aptian Basin (southeastern France). The results are as follows: (1) representatives of Ammonitoceras from this area are regarded dimorphic with criocone macroconchs and ancylocone microconchs, (2) their ontogenetic sequence is described, (3) their intraspecific variability is significant and concerns the adult size and the duration of the ontogenetic stages, especially the Ammonitoceras stage, (4) two species are recognized: the earlier Ammonitoceras ucetiae Dumas, 1876, characterized by a brief Ammonitoceras stage on average, and the latter Ammonitoceras lahuseni (Sinzow, 1906), characterized by a longer Ammonitoceras stage on average. The sample of the Deshayesites grandis Subzone (Deshayesites deshayesi Zone) is composed of specimens too fragmentary to be identified at species level.

Ammonitoceras Dumas, 1876是一个异型菊石属,其特征是在其内轮中存在一个特殊的个体发育阶段:Ammonitoceras期。但是,尽管它在整个Aptian(下白垩纪)具有广泛的古地理和生物地层扩展,但该属仍然鲜为人知。本文研究了法国东南部Les Ferres Aptian盆地下Aptian Deshayesites多共岩(Deshayesites deshayesi Zone)至Dufrenoyia furcata (Dufrenoyia furcata Zone)亚带的菊石类标本。结果表明:(1)该地区有代表性的螺壳类具有环孔大螺和环孔小螺的二态性;(2)描述了螺壳类的个体发生序列;(3)种内变异性显著,与成虫体型和个体发生阶段(尤其是环孔螺阶段)的持续时间有关;早期的Ammonitoceras ucetiae Dumas, 1876年,其特点是平均较短的Ammonitoceras阶段,而后期的Ammonitoceras lahuseni (Sinzow, 1906年),其特点是平均较长的Ammonitoceras阶段。Deshayesites deshayesi亚带(Deshayesites deshayesi带)的标本过于破碎,无法在物种水平上进行鉴定。
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引用次数: 0
A new anaxyelid wood wasp (Hymenoptera: ‘Symphyta’) from the mid-Cretaceous Burmese amber 文章标题中白垩纪缅甸琥珀中一种新的无翅目木蜂(膜翅目:合虫亚目)
IF 1 4区 地球科学 Q3 Earth and Planetary Sciences Pub Date : 2022-07-01 DOI: 10.1016/j.annpal.2022.102568
Corentin Jouault , Gi-Soo Nam , Alexandr P. Rasnitsyn

A new genus and species of anaxyelid wasp is described and figured from the mid-Cretaceous Burmese amber as Curvitexis kopylovi gen. et sp. nov. The placement of this new genus within the Anaxyelidae is corroborated by its wing venation and the configuration of its mesosoma. This new genus differs from all other Syntexinae from the mid-Cretaceous Burmese amber biota, inter alia, owing to its forewing with the vein 1Rs curved and longer than 1M, the crossvein 1r-rs absent, the vein 2Rs + M present; the hind wing with the abscissa 2M + Cu present, the vein m-cu absent, and the cell r closed. The recently described Paraxiphydria resinata Gao et al., 2022 is transferred to Anaxyelidae: Syntexinae, and the subfamily Paraxiphydriinae Gao et al., 2022 is synonymized under Syntexinae.

在中白垩纪缅甸琥珀中发现了一种新属,命名为Curvitexis kopylovi gen. et sp. 11 .。该属的翅脉和介膜结构证实了该属在Anaxyelidae中的位置。该新属与中白垩世缅甸琥珀生物群中所有其它琥珀科的不同之处,主要在于其脉1Rs弯曲且长于1M,无交叉脉1r-rs,脉2Rs + M存在;后翅横坐标2M + Cu存在,静脉m-cu缺失,细胞r闭合。最近描述的paraxphydria resinata Gao et al., 2022被转移到Anaxyelidae: Syntexinae,而paraxphydriinae Gao et al., 2022在Syntexinae下同义。
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引用次数: 1
First description of the most complete Metriorhynchus aff. superciliosus (Thalattosuchia) specimen from the Callovian of the Vaches-Noires cliffs (Normandy, France) and limitations in the classification of Metriorhynchidae 法国诺曼底Vaches-Noires悬崖Callovian最完整Metriorhynchus affer . superciliosus (Thalattosuchia)标本的首次描述及Metriorhynchidae分类的局限性
IF 1 4区 地球科学 Q3 Earth and Planetary Sciences Pub Date : 2022-07-01 DOI: 10.1016/j.annpal.2022.102539
Jonas Le Mort , Jeremy E. Martin , Laurent Picot , Stéphane Hua

The metriorhynchids from the Jurassic of Normandy have been known since the 19th century. However, the remains studied until now have rarely been found in anatomical connection. The acquisition of specimen MPV 2010.3.610 that was found in the Callovian of Vaches-Noires cliffs by the Paleospace Museum in 2010, allowed the first description of the most complete Metriorhynchidae from this locality so far. The analysis of this Metriorhynchidae raises questions about the taxonomic value of several criteria currently used to distinguish between the different Callovian and Oxfordian species. The study of the cranial and postcranial elements shows that this specimen presents diagnostic criteria of different Metriorhynchidae such as the shape of the prefrontals, the mandible, the palato-maxillary suture, the atlas/axis, the cervical vertebrae as well as the extension of the jugal bone and the morphology of the teeth. A discussion highlights the limitations of the current classification of Metriorhynchidae.

自19世纪以来,人们就已经知道了诺曼底侏罗纪的米氏婴。然而,迄今为止研究的遗骸很少有解剖学上的联系。2010年,古空间博物馆在Vaches-Noires悬崖Callovian发现了标本MPV 2010.3.610,首次对该地区最完整的Metriorhynchidae进行了描述。对这一Metriorhynchidae的分析提出了关于目前用于区分不同Callovian和Oxfordian物种的几种标准的分类价值的问题。颅后结构的研究表明,该标本在额骨形状、下颌骨、腭颌缝合、寰椎/轴、颈椎、颧骨的延伸和牙齿的形态等方面具有不同的诊断标准。一篇讨论强调了目前的甲鼠科分类的局限性。
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引用次数: 1
Hypowaagenia? bodenbenderi (Behrendsen): A rare aspidoceratid ammonite from the Tithonian of the Neuquén Basin, Argentina Hypowaagenia吗?bodenbenderi (Behrendsen):一种罕见的蜘蛛壳菊石,产自阿根廷neuqu<s:1>盆地的梯通世
IF 1 4区 地球科学 Q3 Earth and Planetary Sciences Pub Date : 2022-07-01 DOI: 10.1016/j.annpal.2022.102571
Horacio Parent , Günter Schweigert , Armin Scherzinger , Alberto C. Garrido

Among the ammonite collection from the Neuquén and Mendoza provinces made by G. Bodenbender, stands Aspidoceras bodenbenderi Behrendsen. This rare aspidoceratid is assigned to the lower Alternans Zone because of the associated Parodontoceras calistoides (Behrendsen), and can be most likely attributed to the recently established genus Hypowaagenia Schweigert & Schlampp.

在G. Bodenbender收集的neuqusamen和Mendoza省的菊石中,有一种是Aspidoceras bodenbenderi Behrendsen。由于与之相关的pardontoceras calistoides (Behrendsen),这种罕见的蛛形虫被分配到下隔生带,并且最有可能归因于最近建立的Hypowaagenia Schweigert属;Schlampp。
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引用次数: 0
Crinoids from the Ouarsenis Massif (Algeria) fill the Lower Cretaceous (Berriasian and Valanginian) gap of northern Africa 来自阿尔及利亚Ouarsenis地块的海百合填补了北非下白垩世(Berriasian和Valanginian)的缺口
IF 1 4区 地球科学 Q3 Earth and Planetary Sciences Pub Date : 2022-04-01 DOI: 10.1016/j.annpal.2022.102555
Madani Benyoucef , Bruno Ferré , Bartosz J. Płachno , Imad Bouchemla , Mariusz A. Salamon

The Ouarsenis Massif belongs to the Algerian Tell domain, which is considered as the eastern part of the Maghrebian Tethys former margin. The Berriasian-Valanginian Oued Fodda Formation outcropping in the Kef Aïn El Hadjela section, at the foot of the great peak of the Ouarsenis Massif, is composed of marls and laminated and bioturbated limestone alternations. The marl facies reveals diverse microfauna, including crinoids. These latter are represented by: Isocrinida [isocrinids Balanocrinus cf. gillieroni (de Loriol), Percevalicrinus aldingeri Klikushin, Isocrinus? lissajouxi (de Loriol)], Cyrtocrinida [cyrtocrinids (Phyllocrinus sp., Hemibrachiocrinus sp.)], and Roveacrinida gen. indet. sp. indet. [roveacrinids]. All these, with exception of roveacrinids and phyllocrinids, are noted for the first time from Algeria and African continent (southern margin of Tethys in the Maghreb). Knowledge on Cretaceous crinoids formerly described from Algeria is presented. It is also shown that crinoid assemblage and associated invertebrates are typical for relatively shallow, distal depositional setting situated below storm wave base.

Ouarsenis地块属于阿尔及利亚的Tell域,被认为是马格里布特提斯前边缘的东部。Berriasian-Valanginian Oued Fodda组露头位于Kef Aïn El Hadjela剖面,位于Ouarsenis地块的大峰脚下,由泥灰岩和层状和生物扰动的石灰岩交替组成。泥灰岩相显示多种微动物群,包括海百合。后者的代表是:Isocrinida [isocrinids Balanocrinus cf. gillieroni (de Loriol)], Percevalicrinus aldingeri Klikushin, Isocrinus?lissajouxi (de Loriol)], Cyrtocrinida [cyrtocrinids (Phyllocrinus sp., Hemibrachiocrinus sp.)], Roveacrinida genindet。sp. indet。[roveacrinids]。除roveacrinids和phyllocrinids外,所有这些都是首次在阿尔及利亚和非洲大陆(马格里布特提斯南缘)发现的。介绍了以前在阿尔及利亚描述的白垩纪海百合的知识。海百合组合及其伴生的无脊椎动物是位于风暴波基底下方较浅的远端沉积环境的典型特征。
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引用次数: 4
Les palynomorphes du Bajocien supérieur : un premier PALYNO-DATA pour les Monts de Rhar Roubane (Algérie nord occidentale) 上bajocian的palynomorphes: Rhar Roubane山脉(阿尔及利亚西北部)的第一个palyno数据
IF 1 4区 地球科学 Q3 Earth and Planetary Sciences Pub Date : 2022-04-01 DOI: 10.1016/j.annpal.2022.102542
Assia Khellas , Abbas Marok , Louisa Samar

Apart from the few works carried out in the North Tethyan basins, the palynological study of Mesozoic outcrops is still rare. Thus, the present study devoted to the Zahra Marls Formation (upper Bajocian) of the Rhar Roubane Mountains (northwestern Algeria), allowed us for the first time to inventory 44 different taxa. The palynological assemblages studied are strongly dominated by trilete spores (17 genera and 26 species), the most abundant of which are Concavissimisporites variverrucatus, Cyathidites australis and Cyathidites minor. Pollen grains are relatively frequent (10 genera and 16 species), while acritarchs and algae are represented by only two taxa (Micrhystridium sp. and Botryococcus sp.). In addition, the sporopollinic spectra dominated by the terrestrial elements are represented mainly by the spores of Pteridophyta and Bryophyta and gymnosperm pollen.

除了在北特提斯盆地进行的少量工作外,对中生代露头的孢粉学研究仍然很少。因此,本研究致力于Rhar Roubane山脉(阿尔及利亚西北部)的Zahra Marls组(上Bajocian),使我们首次对44个不同的分类群进行了清查。所研究的孢粉组合以三孢子(17属26种)为主,其中最多的是concavisimisporites variverrucatus、Cyathidites australis和Cyathidites minor。花粉粒相对较多(10属16种),而真菌和藻类只有两个分类群(Micrhystridium sp.和Botryococcus sp.)。此外,以陆生元素为主的孢粉谱主要以蕨类植物和苔藓植物的孢子和裸子植物花粉为代表。
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引用次数: 0
Quantitative analysis of horizontal bioturbation from Brioverian (Ediacaran - Fortunian) deposits of Brittany (Armorican Massif, NW of France) 法国西北部布列塔尼(armoramerican地块)布里奥纪(埃迪卡拉纪-福图尼纪)沉积水平生物扰动定量分析
IF 1 4区 地球科学 Q3 Earth and Planetary Sciences Pub Date : 2022-04-01 DOI: 10.1016/j.annpal.2022.102543
Baptiste Coutret , Didier Néraudeau

Ediacaran-Cambrian bioturbation on bedding planes provides physical and chemical data about the environmental conditions and biological activities of early metazoans. We propose a quantitative method to estimate horizontal disruption of the substrate using bedding-plane trace fossils from Brioverian deposits. This methodology provides the first quantitative analysis of the trace fossil assemblage from the Armorican Massif (Brittany, NW of France). The Ediacaran-Cambrian transition within the Brioverian series is characterized by the abundance of simple and horizontal trace fossils such as Helminthopsis, Helminthoidichnites, and Palaeophycus as well as rare Gordia and Spirodesmos. Recent U-Pb dating has been carried out on detrital zircon grains from the upper Brioverian series suggesting a late Ediacaran to Fortunian age of the fossiliferous deposits (ca. 550 to 530 Ma). With Arc-Gis software, we use a semi-quantitative approach to estimate the relative 2D bioturbation rate recorded on bedding planes. The dataset is used to discuss the feeding strategies and how the seafloor ecospace was colonized by the early bilaterian metazoans. This approach combines new values such as: the length of a trace fossil, the surface of a trace, the cumulative length, and the cumulative surfaces from all of the ichnofossils recorded on the same slate surface, to finally suggest that the bioturbation rate of the microbial grazers impacted the use of the seafloor ecospaces and feeding strategies through the biomats.

埃迪卡拉-寒武系层理平面生物扰动为早期后生动物的环境条件和生物活动提供了物理化学资料。我们提出了一种定量的方法来估计底物的水平破坏,利用从布里弗纪沉积物的层理面痕迹化石。该方法首次对来自法国西北部布列塔尼的美洲地块的微量化石组合进行了定量分析。埃迪卡拉-寒武纪在布里弗系的过渡时期,具有丰富的简单和水平的痕迹化石,如Helminthopsis、Helminthoidichnites和Palaeophycus,以及罕见的Gordia和Spirodesmos。最近对上布里奥世系列的碎屑锆石颗粒进行了U-Pb定年,表明化石矿床的年龄为埃迪卡拉世晚期至福图尼世(约550 ~ 530 Ma)。利用Arc-Gis软件,我们采用半定量方法估计了记录在层理平面上的相对二维生物扰动率。该数据集用于讨论早期双边后生动物的摄食策略以及海底生态空间是如何被殖民的。这种方法结合了新的价值,如:痕迹化石的长度、痕迹的表面、累积长度和在同一板岩表面上记录的所有鱼化石的累积表面,最终表明微生物食草动物的生物扰动率通过生物生物影响了海底生态空间的使用和摄食策略。
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引用次数: 1
Ammonites du Toarcien du Haut Atlas central (Maroc)
IF 1 4区 地球科学 Q3 Earth and Planetary Sciences Pub Date : 2022-04-01 DOI: 10.1016/j.annpal.2022.102540
Mohamed Benzaggagh , Hanane Khaffou , Mariusz A. Salamon , Touria Hssaida , Mohamed El Ouali , Badre Essafraoui

In the central High Atlas, the Toarcian Stage is represented by a marl and marly limestone series of variable thickness. Two sections have been studied: Amellago (500 m) and Aït Athmane (100 m). Ammonites from the two sections allowed to characterize the Polymorphum, Levisoni, Bifrons, Gradata, and Speciosum zones. The Polymorphum Zone provided, among others: Dactylioceras (Eodactylites) mirabile, D. (E.) pseudocommune, Dactylioceras (Orthodactylites) aff. crosbeyi and Neolioceratoides aff. hoffmanni; the Levisoni Zone provided a rich ammonite fauna composed of, Calliphylloceras nilssoni, Dactylioceras (Orthodactylites) cf. semiannulatum, Eleganticeras exaratum, Harpoceras falciferum, H. pseudoserpentinum, H. serpentinum, H. subplanatum, Hildaites cf. forte, H. levisoni, H. cf. serpentiniformis, H. cf. subserpentinus, H. striatus, H. wrighti, Lytoceras siemensi, Lytoceras sp., Maconiceras soloniacense, Phylloceras sp., Polyplectus pluricostatum et Polyplectus sp.; the Bifrons Zone provided Eleganticeras sp., Harpoceras subplanatum?, Hildoceras bifrons, H. lusitanicum, H. semipolitum, H. sublevisoni, Hildoceras sp., Porpoceras gr. vortex verticosum; the Gradata Zone was characterized by Pseudocrassiceras bayani, P. frantzi, Pseudocrassiceras sp., Pseudocrassiceras sp. indet.; and the Speciosum Zone provided an Hammatoceras aff. insigne. These species have been described and illustrated for the first time for the central High Atlas. Most of them are common to several basins of the north Tethyan margin and the Subboreal Domain of NW Europe.

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引用次数: 3
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Annales de Paleontologie
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