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Isopod crustacean fossils from the Cenomanian stratotype: five new species in suborders Cymothoida, Asellota and Valvifera 盖诺曼尼亚层型等足甲壳类化石:Cymothoida亚目、Asellota亚目和Valvifera亚目五新种
IF 1 4区 地球科学 Q3 Earth and Planetary Sciences Pub Date : 2022-01-01 DOI: 10.1016/j.annpal.2022.102538
George D.F. Wilson , Nicolas Morel

We report on a significant fossil collection of five isopod crustaceans that belong to five families from three suborders. These fossils were discovered in Le Mans (western part of Paris Basin, France) during the second half of the nineteenth century by the French paleontologist Edouard Guéranger. The historical quarry has been studied a few years before by the French paleontologist Alcide d’Orbigny and used as part of his Cenomanian stratotype (Leach, 1814). The collection consists of two species in the Cymothooidea Dana, 1852 (Cirolanidae Dana, 1852; Lantoceramiidae fam. nov.), two Valvifera G. O. Sars, 1883 (Chaetiliidae Dana, 1853; Idoteidae Samouelle, 1819) and one Asellota Latreille, 1802 (Stenetriidae Hansen, 1905), which were previously unknown from Upper Cretaceous strata as old as the Cenomanian age (roughly 93–99 ma). Although phylogenetic dating based on extant taxa has assigned Permian to Triassic age of origin for the Asellota, the Stenetriidae Hansen, 1905 specimen is the first fossil record for this family. Another asellotan, Fornicaris calligarisi Wilson and Selden, 2016, is known from the Triassic (Norian) dating from approximately 210–215 ma. The valviferans were unknown from Mesozoic strata, previously being found in the Oligocene of Romania and the Fur Formation, Upper Paleocene/Lowermost Eocene of Denmark (Polz, 2007). This diverse assemblage of fossils with taxa assignable to extant families and one new family provides evidence for the presence of a substantially modern isopod fauna as early as the Cenomanian.

我们报告了五种等足类甲壳类动物的重要化石收集,它们属于三个亚目的五个科。这些化石是19世纪下半叶在勒芒(法国巴黎盆地西部)由法国古生物学家爱德华·古萨兰杰发现的。几年前,法国古生物学家Alcide d 'Orbigny对这个历史上的采石场进行了研究,并将其用作其Cenomanian层型的一部分(Leach, 1814)。该标本包括2种,属Cymothooidea Dana, 1852 (cirrolanidae Dana, 1852;Lantoceramiidae家人。11月),两个Valvifera g.o. Sars, 1883 (Chaetiliidae Dana, 1853;Idoteidae Samouelle, 1819年)和Asellota Latreille, 1802年(Stenetriidae Hansen, 1905年),这是以前未知的上白垩纪地层,可追溯到Cenomanian时代(大约93-99 ma)。尽管基于现存分类群的系统发育定年将Asellota的起源年龄划分为二叠纪至三叠纪,但1905年的Stenetriidae Hansen标本是该科的第一个化石记录。另一种asellotan, Fornicaris calligarisi Wilson和Selden, 2016年发现,来自三叠纪(诺里亚),距今约210-215 ma。valviferans在中生代地层中是未知的,以前在罗马尼亚渐新世和丹麦上古新世/始新世最底层的Fur组中发现过(Polz, 2007)。这一多样化的化石组合,其分类群可划分为现存的科和一个新的科,为早在塞诺曼尼亚时期就存在实质上现代的等足类动物群提供了证据。
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引用次数: 0
A new protopristocerine wasp (Hymenoptera: Bethylidae) from the mid-Cretaceous Burmese amber 文章标题中白垩世缅甸琥珀中一新种原夜蛾蜂属(膜翅目:夜蛾科)
IF 1 4区 地球科学 Q3 Earth and Planetary Sciences Pub Date : 2021-10-01 DOI: 10.1016/j.annpal.2021.102522
Corentin Jouault , Manuel Brazidec

Ramageoptera platycephala gen. et sp. nov., a new protopristocerine wasp, is described and figured from a female individual preserved in mid-Cretaceous amber from Tanai, northern Myanmar. Ramageoptera platycephala gen. et sp. nov. shows most of the known characters used to circumscribe the extinct subfamily Protopristocerinae, and highlights the putative status of stem-Pristocerinae that may occupy the Protopristocerinae. Ramageoptera platycephala gen. et sp. nov. is unique among the Protopristocerinae, at least, for possessing a flat and square shaped head; eyes glabrous and small; mandibles with four conspicuous sharp teeth; antenna with short scape; forewing with Rs + M vein absent; vein M straight; cell 2R1 distally opened; cell 1Cu closed, sub-equal to R cell; cell 2Cu open.

在缅甸北部塔奈发现的中白垩世琥珀中,发现了一种新的原始原始黄蜂——Ramageoptera platycephala gen. et sp. nov.。Ramageoptera platycephala gen. et sp. 11 .显示了大部分已知的特征,用于划定已灭绝的原猿亚科,并强调了可能占据原猿亚科的干猿亚科的假定地位。至少在原始鸟科中,有一个扁平的方形头是独一无二的;眼睛无毛和小;下颌骨有四颗明显的尖牙;具有短柱面的天线;前翼无Rs + M脉;脉M直;2R1细胞远端打开;单元1Cu闭合,次等于R单元;2Cu单元打开。
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引用次数: 2
Upper Pliocene bivalve shell concentrations from the Lower Chelif basin (NW Algeria): Systematics, sedimentologic and taphonomic framework 下切里夫盆地(阿尔及利亚西北部)上新世双壳类贝类的富集:系统学、沉积学和地貌学框架
IF 1 4区 地球科学 Q3 Earth and Planetary Sciences Pub Date : 2021-10-01 DOI: 10.1016/j.annpal.2021.102509
Madani Benyoucef , Mohamed Bendella , Mauro Brunetti , Bruno Ferré , Tomas Koci , Imad Bouchemla , Rafika Slami , Asma-Fethia Ghenim

Sedimentologic and palaeontological investigation of the Upper Pliocene Slama Formation in the Lower Chelif Basin (NW Algeria) led us to collect important bivalve assemblages for taxonomic and taphonomic purposes. A rather comprehensive inventory list of Upper Pliocene bivalves from northwestern Algeria is now available and consists of 30 species, 17 of which are extinct ones. Four principal taphonomic attributes were analysed: bioerosion, encrustation, fragmentation, and abrasion. Physical and biogenic sedimentary structures are used for palaeoenvironmental interpretations. The taphonomic, sedimentologic and ichnological characteristics of most of the deposits suggests they originated from discontinuous processes of winnowing and bypassing of sediments, probably due to the action of storms in shallow waters, mainly in the shoreface depositional environment. The bivalve assemblage is dominated by disarticulated valves and displays significant taphonomic alteration in the shells. Sclerobionts traces in shells particulary affect the oyster shells. Bioerosion traces are predominately those of clionid sponges (Entobia isp.), polychaetes (Maeandropolydora isp. and Caulostrepsis isp.), bivalves (Gastrochaenolites isp.), and of predatory gastropods (Oichnus isp.). Among the sclerobionts, the identified encrusters were juvenile oyster recruits, barnacles, polychaetes (serpulid tubeworms), bryozoans (Microporella sp. and Acanthodesia sp.), and vermetid gastropods (Petaloconchus intortus).

通过对下切里夫盆地(NW阿尔及利亚)上上新世火山岩组的沉积学和古生物学研究,我们收集了重要的双壳类组合,用于分类和分类目的。阿尔及利亚西北部有一份相当全面的上新世双壳类清单,包括30种,其中17种已经灭绝。分析了四种主要的埋藏属性:生物侵蚀、结壳、破碎和磨损。物理和生物成因的沉积构造被用于古环境解释。大部分沉积物的地貌学、沉积学和工艺学特征表明,它们可能是由于浅水(主要是滨面沉积环境)的风暴作用而形成的不连续的筛分和绕流过程。双壳类组合以分离的壳类为主,在壳中表现出明显的语音学变化。牡蛎壳中的固化剂对牡蛎壳的影响尤甚。生物侵蚀痕迹主要是clionid sponge (Entobia isp.), polychaetes (Maeandropolydora isp.)。和Caulostrepsis isp.),双壳类(Gastrochaenolites isp.)和掠食性腹足类(Oichnus isp.)。在硬壳动物中,鉴定出的壳类有幼生牡蛎、藤壶、多毛纲(绢状管虫)、苔藓虫(微孔虫和棘虫)和蚓状腹足类(Petaloconchus intortus)。
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引用次数: 4
Les radiolaires messiniens du Dahra (Bassin du Bas Chélif, Algérie) : systématique et intérêt biostratigraphique 达赫拉的墨西尼亚放射虫(阿尔及利亚下切利夫盆地):系统和生物地层学的兴趣
IF 1 4区 地球科学 Q3 Earth and Planetary Sciences Pub Date : 2021-10-01 DOI: 10.1016/j.annpal.2021.102520
Ali Mokhtar Samet , Abbas Marok , Matías Reolid , Shin-Ichi Kamikuri

The micropaleontological study of the Upper Miocene diatomites of the Dahra massif at Ouillis and Sidi Lakhdar sections (Lower Chelif Basin, Algeria), allowed for the first time in Algeria, the identification of 36 species and 22 genera of radiolarians dominated by families Actinommidae, Spongodiscidae and Theoperidae. The qualitative and quantitative analyses of the assemblages show significant specific and generic diversification in the first members. Towards the top of the sections, the progressive decrease in diversity and finally, the disappearance of siliceous microfauna are linked to a reoxygenation of the environment. Respect to the biostratigraphy, the radiolarian assemblage identified confirms a Messinian age (Biozone Stichocorys peregrina).

通过对阿尔及利亚下Chelif盆地Dahra块体Ouillis和Sidi Lakhdar剖面上中新世硅藻的微古生物学研究,在阿尔及利亚首次鉴定出放射虫属36种22属,以放线虫科、海绵虫科和针叶虫科为主。对这些组合的定性和定量分析表明,第一段具有明显的特异和一般多样性。在剖面的顶部,多样性逐渐减少,最后,硅质微动物群的消失与环境的再氧化有关。在生物地层学方面,所鉴定的放射虫组合确认了一个迈西尼亚时代(Biozone Stichocorys peregrina)。
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引用次数: 0
Les Pseudholaster (Echinoidea, Holasteroida) du Crétacé de France 法国白垩纪的伪holaster(棘总科,Holasteroida)
IF 1 4区 地球科学 Q3 Earth and Planetary Sciences Pub Date : 2021-10-01 DOI: 10.1016/j.annpal.2021.102519
Jean-Christophe Dudicourt

In the order of Holasteroida, the fossil record highlights a contradiction between the genus Pseudholaster that appears in the Aptian, whose plastron is prostostern close to the Jurassic ancestors and the genus Holaster, which appears in the Valanginian, whose meridostern plastron appears more derived. This inconsistency can be explained by the ignorance of the plastronal architecture on the part of the early authors. A review of the species of Pseudholaster from the Cretaceous period of France was therefore carried out. The objective was to statistically determine the discriminating morphological characters, and to study the modifications of the architecture of the interambulacrum 5 of the French species belonging to this genus, as well as to the species included in the genus Holaster incorrectly by earlier authors. This review of the species of the genus Pseudholaster begins with a study of the ontogeny of the species Holaster intermedius Münster in Goldfuss, 1826–1833, first representative of the genus Pseudholaster, which appears in the Hauterivian in the Parisian and Rhodano-vocontian basins. The modifications during growth concern the overall shape, but also the plastron architecture: the number of plastron plates increases while the number of plates located between the peristome and the periproct remains fixed. The plastron of this species is protosternal and not meridosternal as Lambert pointed out. The labrum is cupuliform in contact with the second sternal 5a2 by a narrow digitation. However, this arrangement differs from that observed on a protosternal breastplate. This apomorphism of the plastron plate pattern, called “labrotaxienne”, is found in all the Pseudholaster studied, and the study of the architecture of the interambulacrum 5 also reveals a gradual decrease in the number of preanal plates between the oldest (Hauterivian) and the younger (Cenomanian-Lower Turonian) species studied. Most of the French species have been revised, with some synonyms. A new species, P. neraudeaui, is the last known Pseudholaster dated from the upper Cenomanian and lower Turonian of southwestern France. Our study illustrates the evolution of the genus Pseudholaster between the Hauterivian and the early Turonian in France. The interest of the study is to show that the appearance of the genus Pseudholaster is older than that of the genus Holaster. Pseuholaster intermedius, of Hauterivian age, possesses a derived protostern plastron called here “labrotaxien” and not meridostern as defined historically by Lambert, and to reveal that the number of preanals decreases over geological time. This data is essential for future phylogenetic studies. On a palaeobiogeographical level, the study reveals the expansion of the genus Pseudholaster during early Cretaceous in western Europe, with diversification during t

在霍拉斯特目中,化石记录突出了出现在阿普tian的伪霍拉斯特属和出现在Valanginian的霍拉斯特属之间的矛盾,阿普tian的假霍拉斯特属的板状体是接近侏罗纪祖先的前直立的,而瓦兰吉尼亚的霍拉斯特属的子午线板状体似乎更接近侏罗纪祖先。这种不一致可以用早期作者对平面架构的无知来解释。因此,对法国白垩纪时期的假枪龙种进行了综述。目的是统计确定鉴别形态学特征,并研究属于该属的法国种的间肢5的结构修改,以及早期作者错误地包括在Holaster属中的种。本文从1826-1833年Goldfuss的Holaster intermedius m nster种的个体发育研究开始,这是假Holaster属的第一个代表,出现在巴黎和罗达诺-沃孔洲盆地的Hauterivian。生长过程中的修改涉及整体形状,但也涉及板层结构:板层板的数量增加,而位于围壁和围壁之间的板层数量保持不变。如Lambert指出的,这个物种的板是前胸骨的,而不是子骨的。唇形呈壶状,通过狭窄的指状与第二胸骨5a2接触。然而,这种排列不同于在原胸骨胸甲上观察到的排列。这种被称为“labrotaxienne”的板状板块模式的非同形性在所有被研究的伪holaster中都有发现,对interambulacrum 5结构的研究也揭示了在最古老(Hauterivian)和最年轻(Cenomanian-Lower Turonian)物种之间的前肛门板块数量逐渐减少。大多数法国种已被修订,有一些同义词。一个新种,P. neraudeaui,是已知的最后一种来自法国西南部上塞诺曼尼亚纪和下Turonian纪的假枪龙。我们的研究说明了法国Hauterivian和早期Turonian之间的假holaster属的进化。这项研究的兴趣在于证明假枪属的出现比枪属的出现更古老。Hauterivian时代的pseudoholaster intermedius拥有一个衍生的原始板,这里称为“labrotaxien”,而不是Lambert历史上定义的meridosien,并且揭示了前动物的数量随着地质时间的推移而减少。这些数据对未来的系统发育研究至关重要。在古地理水平上,研究揭示了假牙龙属在早白垩世在西欧的扩张,在阿尔比安时期的多样化,在巴黎盆地晚塞诺曼尼亚时期的消失,而在阿基塔盆地仍然存在,其对环流环境的偏好。
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引用次数: 0
Middle Permian ostracod fauna from the Khao Khad Formation (Indochina Terrane), Central Thailand
IF 1 4区 地球科学 Q3 Earth and Planetary Sciences Pub Date : 2021-10-01 DOI: 10.1016/j.annpal.2021.102521
Anisong Chitnarin , Chatchalerm Ketwetsuriya

Permian ostracods are reported for the first time from the Wordian (Middle Permian) Khao Khad Formation of Saraburi Group (Lopburi Province, Central Thailand). The ostracod fauna consists of 18 species belonging to 11 genera including Bairdia, Bohlenatia, Liuzhinia, Silenites, Acratia, Bairdiacypris, Basslerella, Aurigerites, Microcheilinella, Paraparchites and Shemonaella. The studied Khao Khad Limestone, which are rich in fusulinids, gastropods, ostracods, bivalves and brachiopods, was deposited in a shallow carbonate platform on the western margin of the Indochina Terrane. The ostracod assemblage is typically Palaeo-Tethyan and similar to faunas from the other Lower to Middle Permian limestones in central Thailand.

在泰国中部lophburi省Saraburi群的Wordian(中二叠统)Khao Khad组中首次报道了二叠纪介形类。介形类区系包括Bairdia、Bohlenatia、Liuzhinia、Silenites、Acratia、Bairdiacypris、Basslerella、Aurigerites、Microcheilinella、Paraparchites和Shemonaella等11属18种。介形类组合是典型的古特提斯,与泰国中部其他下二叠世至中二叠世石灰岩的动物群相似。
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引用次数: 0
Addendum à l’article « Les faunes coralliennes de l’Oligocène de Malte : biodiversité et paléoenvironnement. ». Annales de Paléontologie 107 (2021) 102508 “马耳他渐新世珊瑚群:生物多样性和古环境”文章的附录。»。古生物学年鉴107 (2021)102508
IF 1 4区 地球科学 Q3 Earth and Planetary Sciences Pub Date : 2021-10-01 DOI: 10.1016/j.annpal.2021.102523
Jean-Paul Saint Martin , Christian Chaix , Bruno Cahuzac , Pierre Moissette , Jean-Pierre André
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引用次数: 0
Les faunes coralliennes de l’Oligocène de Malte : biodiversité et paléoenvironnement 马耳他渐新世珊瑚群:生物多样性与古环境
IF 1 4区 地球科学 Q3 Earth and Planetary Sciences Pub Date : 2021-07-01 DOI: 10.1016/j.annpal.2021.102508
Jean-Paul Saint Martin , Christian Chaix , Bruno Cahuzac , Pierre Moissette , Jean-Pierre André

The study of the coral biodiversity of the Upper Oligocene sedimentary series (Lower Coralline Limestone, Chattian) of Malta permitted the identification of 25 scleractinian genera represented by 41 species. Three new species are proposed: Miophora naxxarensis n. sp., Nerthastraea maltensis n. sp. and Gyrosmilia maltensis n. sp. Observations made in various Oligocene sites in Malta, especially around Naxxar and Tal Bajjada, allow to establish a suite of coral associations which may vary spatially and vertically. The identified scleractinian associations can be represented by different types of coral assemblages and bioconstructions: isolated colonies, coral beds of varying density but of large spatial extent, more cohesive coral banks (coral carpets) or patch reefs forming morphologies with gentle lateral slopes. The colonial morphologies are closely linked to these different types of bioconstructions. These coral constructions have been established and developed in shallow marine areas. The coral biodiversity of the Oligocene of Malta fits well in the evolution of the Cenozoic reef phenomenon whose development reached its peak in the Oligocene with great coral richness in the Chattian in the Mediterranean area.

通过对马耳他上渐新世沉积系列(Chattian下珊瑚系灰岩)珊瑚生物多样性的研究,鉴定出了41种、25个硬核系属。提出了3个新种:Miophora naxxarensis n. sp、Nerthastraea maltensis n. sp和Gyrosmilia maltensis n. sp。在马耳他的多个渐新世遗址,特别是在Naxxar和Tal Bajjada附近的观测,允许建立一套可能在空间和垂直上变化的珊瑚组合。已确定的核结系组合可以由不同类型的珊瑚组合和生物结构来代表:孤立的群落,密度不同但空间范围大的珊瑚床,更有凝聚力的珊瑚滩(珊瑚地毯)或形成平缓侧坡形态的斑块珊瑚礁。殖民地形态与这些不同类型的生物结构密切相关。这些珊瑚建筑是在浅海地区建立和发展起来的。马耳他渐新世珊瑚的生物多样性与新生代珊瑚礁现象的演化非常吻合,渐新世是珊瑚礁发展的高峰,地中海地区的Chattian珊瑚非常丰富。
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引用次数: 3
Paléoécologie des foraminifères benthiques de nouvelles coupes de basse latitude du passage Crétacé-Paléogène : Coupes de l’Oued Es Smara et de l’Oued Abiod (région de Téjerouine, NW Tunisie) 白垩纪-古生代通道新低纬度剖面的底栖有孔虫古生态学:埃斯马拉河和阿比奥德河剖面(tejerouine地区,突尼斯西北部)
IF 1 4区 地球科学 Q3 Earth and Planetary Sciences Pub Date : 2021-07-01 DOI: 10.1016/j.annpal.2021.102491
Njoud Gallala , Moez Ben Fadhel

Due to an impact of a bolide at the K/Pg boundary, the planktonic foraminifera have suffered sever mass extinction. However, no small Benthic Foraminifera species have documented mass extinction at the K/Pg boundary. Nevertheless, many species showed disturbance. The Maastrichtian assemblages may be different from those of the lower Paleogene by their species content, diversity and frequencies. At Oued Es Smara and Oued Abiod sections, the small benthic foraminifera indicate lower bathyal environment, and manifest significant faunal turnover. Until the uppermost Maastrichtian, their assemblages are highly diversified, with 77 species and 76 species respectively at Oued Es Smara and Oued Abiod sections. These are dominated by endobenthic morphotypes. At the K/Pg boundary, although 33 species (42,85%) (Oued Es Smara section) and 27 species (35,52%) (Oued Abiod section) of them seem to disappear, but only few species have really extinct such as Arenobulimina obesa. Nevertheless, the majority of species persist elsewhere at the Danian (e.g., Pseudoglandulina manifesta, Cibicioides proprius, Clavulinoides amorpha, Coryphostoma plaitum, Pullenia coryelli). At the lower Danian, the survivor Maastrichtian species are of 58% (Oued Es Smara) and 65% (Oued Abiod). Throughout the Parasubbotina pseudobulloides subzone, 4 others species were progressively disappeared. They are oligotrophic and low oxygen tolerant. About the Masstrichtian species, at the two studied sections (e.g. Gaudryina inflata and Tritaxia midwayensis) they seem to be more trophic exigent. Consequently, the benthic Foraminifera did not suffer massive extinction at the K/Pg boundary, but their assemblages underwent a significant faunal turnover which reflects important environmental changes. These changes are compatible with the catastrophic scenario induced by the large asteroid impact.

由于K/Pg边界的一次撞击,浮游有孔虫遭受了严重的大灭绝。然而,在K/Pg边界没有记录到小型底栖有孔虫物种的大规模灭绝。然而,许多物种表现出干扰。马斯特里赫特组合在物种含量、多样性和频率上可能与下古近纪不同。在Oued Es Smara和Oued Abiod剖面中,小底栖有孔虫显示了较低的深海环境,并表现出明显的动物更替。到马斯特里赫特最上层,它们的组合高度多样化,在Oued Es Smara和Oued Abiod区段分别有77种和76种。它们以底栖生物形态为主。在K/Pg界线上,虽然有33种(42.85%)(Oued Es Smara剖面)和27种(35.52%)(Oued Abiod剖面)似乎消失了,但真正灭绝的只有少数几种,如Arenobulimina obesa。然而,大多数物种在大年系的其他地方仍然存在(例如,Pseudoglandulina manifesta, Cibicioides proprius, Clavulinoides amorpha, Coryphostoma plaitum, Pullenia coryelli)。在大年河下游,幸存的马斯特里赫特物种为58% (Oued Es Smara)和65% (Oued Abiod)。在伪球拟虎目亚区,其他4种逐渐消失。它们是低营养和低氧耐受性的。在两个研究区(例如Gaudryina inflata和Tritaxia midwayensis), Masstrichtian的物种似乎更营养迫切。因此,在K/Pg界线处,底栖有孔虫并没有大规模灭绝,但它们的组合经历了显著的动物群更替,反映了重要的环境变化。这些变化与大型小行星撞击引起的灾难性情景是相容的。
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引用次数: 0
Corrigendum to “Biostratigraphy and Paleobiogeographic implications of the Cenomanian – Early Turonian Ostracods of Egypt” [Ann. Paleontol. 106 (2020) 102408] “埃及塞诺曼尼亚-早期土尔onian介形类的生物地层学和古生物地理学意义”的勘误表[Ann.]。古生物学通报,2006 (5):349 - 349 [j]
IF 1 4区 地球科学 Q3 Earth and Planetary Sciences Pub Date : 2021-07-01 DOI: 10.1016/j.annpal.2021.102505
Mohamed M. Khalil
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Annales de Paleontologie
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