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Luzon predators: Clues from a fossil with bite marks 吕宋岛的掠食者带有咬痕的化石提供的线索
IF 1 4区 地球科学 Q4 PALEONTOLOGY Pub Date : 2023-01-01 DOI: 10.1016/j.annpal.2023.102649
Meyrick U. Tablizo , Rebekka Volmer , Allan Gil S. Fernando , Juan C. Rofes
Luzon Island remained isolated throughout its emergent history with its paleofauna often considered depauperate. However, the terrestrial vertebrate fossil record of the island is still poorly known. A new fossil-bearing paleochannel deposit was discovered in La Union Province, northwestern Luzon, Philippines. This paleochannel is inferred to be of Pleistocene age and has yielded fossils tentatively classified as a bovid (Bubalus sp.), a cervid (Rusa sp.), a suid (Sus sp.), and a large crocodile (Crocodylidae). Interestingly, a proximal fragment of the left tibia of a cervid shows multiple bite marks. This study analyzes the bite marks and modifications on the specimen to infer its possible trace maker and explore its implications in understanding the ancient predatory guild of Luzon. Three sets of modifications were recognized on the specimen, namely the bite marks, the broken proximal end and the dry bone fractures. A crocodilian agent for the bite marks is supported by the bisections, their occurrence in the paleofaunal assemblage and the fluvial setting of the deposit. However, an unknown carnivoran agent interpretation cannot be fully disregarded given the ambiguity of bisected marks and the possibly gnawed proximal end of the specimen. Unfortunately, given the lack of in situ context and apparent overprinting, the extent of the peri- and post-mortem modification remains uncertain which hinders the identification of the predatory agent. Verification of the observations reported in this study would require the systematic collection and detailed taphonomic analysis of additional fossil specimens from the deposit.
卢松岛在历史上一直与世隔绝,其古生物群通常被认为是贫乏的。然而,人们对该岛的陆生脊椎动物化石仍然知之甚少。在菲律宾吕宋岛西北部的拉乌尼翁省发现了一个新的古河道化石矿床。这条古河道被认为可以追溯到更新世,出土的化石被暂时归类为牛科(sp.)、鹿科(sp.)、麂科(sp.)和大型鳄鱼科(Crocodylidae)。值得注意的是,左侧颈鹿胫骨的近端碎片上有多处咬痕。我们对这些咬痕的分析表明了它们的来源。这对吕宋岛古代食肉动物群的组成具有重要意义。在描述的标本上发现了三个系列的变化:咬痕、近端断裂和干骨上的骨折。观察到的痕迹可能与动物群中鳄鱼的痕迹一致,并且与沉积物的水流环境相吻合。然而,由于咬痕和近端被啃咬的痕迹模糊不清,不能完全排除另一种未知食肉动物的解释。遗憾的是,由于缺乏背景资料和明显的叠印,这些痕迹的改变程度仍不确定,从而阻碍了对食肉动物的最终鉴定。需要对遗址中的其他化石标本进行进一步观察、系统收集和详细的岩石学分析。
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引用次数: 0
Morphology of teleosaurid osteoderms from the Phu Kradung Formation of Thailand 泰国普卡东地层远东龙类骨器的形态学
IF 1 4区 地球科学 Q4 PALEONTOLOGY Pub Date : 2023-01-01 DOI: 10.1016/j.annpal.2023.102653
Supanut Bhuttarach , Uthumporn Deesri , Prapasiri Warapeang , Nonthiwat Taesuk , Komsorn Lauprasert
This is a morphological analysis of the osteoderm patterns of Indosinosuchus from the Late Jurassic Phu Noi excavation site in Kalasin Province, Thailand. Data on 203 specimens indicated that 197 osteoderms belongs to the teleosaurid genus Indosinosuchus. Out of 203 total, 109 specimens are dorsal osteoderm morphotypes (M1 to M5) with sub-square, sub-rectangular, or rhomboidal in shape, including keel and peg. Another 88 are ventral osteoderms (M6 and M7) with rectangular or sub-square osteoderms, obscure keel and peg. Large subcircular ellipsoid pits with an anterior facet bar are discovered in both dorsal and ventral osteoderms. Data on morphological features enabled osteoderm positions of Indosinosuchus to be described. The anterior cervical osteoderms are characterized by an obscure keel and a small sub-square shape (M1). In the posterior trunk region, the osteoderms increase in width and adopt a sub-rectangular shape (M2). These osteoderms decrease in size at the sacral level (M3) with a more pronounced keel. The osteoderms gradually decrease in size as they adopt a rhombus shape towards the caudal region (M4) until the caudal end (M5). The ventral osteoderms, with a lack of keel and peg, exhibit a rectangular shape in the central region (M6), while a square or sub-square shape is observed along its lateral margin (M7). Six specimens are could not be assigned to Indosinosuchus and were described as morphotype 8 (M8). This morphotype has distinct characteristics resembling the ventral osteoderms of Mesoeucrocodylia that are yet to be described. The discovery of Indosinosuchus and Mesoeucrocodylia osteoderms suggests a diverse range of crocodyliformes in the Phu Noi fossil excavation site with potential for future studies.
本研究对泰国加拉信府 Phu Noi 上侏罗纪遗址出土的骨龙进行了形态分析。对 203 个标本收集的数据表明,197 个骨皮属于远程龙科。在这203件标本中,有109件是背侧的骨皮形态(M1至M5),形状为近方形、近矩形或斜方形,有一个突起和一个小齿。另外88个是腹部骨皮形态(M6和M7),呈长方形或近方形,没有明显的突缘或齿状突起。背侧和腹侧的骨膜具有大的近圆形椭圆杯和一个条形的前关节面。形态特征数据可用于描述骨膜的位置。颈椎前部骨膜的特征是有一个不明显的突起,形状近方形(M1)。在躯干的后部,骨膜的宽度增加,呈亚矩形(M2)。这些骨膜在骶骨处缩小(M3),其骨突更加明显。骨膜的尺寸逐渐减小,向尾部(M4)呈菱形,直至尾端(M5)。腹侧的骨膜没有突缘和齿状突起,在中央区域呈长方形(M6),而侧面的骨膜呈正方形或近正方形(M7)。有 6 个标本无法归入 M8 形态型,被描述为 M8 形态型。该形态型具有与中目栉水母腹侧骨膜相似的明显特征,但尚未被描述。Mesoeucrocodylia和M8骨膜的发现表明在富诺化石遗址中还存在几种鳄形目,这可能是未来研究的主题。
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引用次数: 0
A new genus of praeaulacid wasp (Hymenoptera: Evanioidea) from the mid-Cretaceous Kachin amber (Myanmar) 白垩纪中期克钦琥珀中的一个新属(膜翅目:Evanioidea)(缅甸)
IF 1 4区 地球科学 Q4 PALEONTOLOGY Pub Date : 2023-01-01 DOI: 10.1016/j.annpal.2023.102599
Corentin Jouault , Simon Rosse-Guillevic

The diversity of praeaulacid wasps in Kachin amber is relatively poorly documented. Here, a new genus and species of praeaulacid wasp are described and illustrated from the mid-Cretaceous Kachin amber as Archeogastrinus kachinensis gen. et sp. nov., and placed in the subfamily Praeaulacinae. This new genus is distinguishable from the other Praeaulacidae because of its particular tibial spurs formula 1-2-2; its forewings with a long and thin pterostigma, a 1-M longer than 1-Rs, a 1-Rs longer than its distance to pterostigma, a crossvein 2r-rs aligned (or nearly) with 2rs-m and originating near the middle of pterostigma, a 2rs-m present, a cell 3rm long, a 1cu-a postfurcal to 1-M, a vein 2-M distinct; its hind wings cu-a reaching Cu distad M + Cu fork; and its metasoma with a first segment petiole-like (i.e., long and thin) and ventrally inflated in its distal half. The position of the genus Mesevania is briefly discussed, as well as the validity of the genus Paleosyncrasis, both described from the mid-Cretaceous Kachin amber.

克钦琥珀中前叶胡蜂的多样性文献记载较少。本文描述了中白垩世克钦琥珀中一个新属和新种,称为Archeogastrinus kachinensis gen. et sp. nov.,并将其归入预胡蜂亚科。这个新属由于其特殊的胫刺公式1-2-2而与其他前肢酸科区分开来;其前翅具一细长的翼柱头,长1 ~ 1米,长1 ~ 2米,距翼柱头1 ~ 2米,与翼柱头近正中平行(或近)的横脉2 ~ 2米,有1 ~ 2米长,室长3米,后径1 ~ 1米,脉2 ~ 2米明显;其后翅Cu -a伸至Cu - distad M + Cu叉;其下半部的瘤具有第一节叶柄状(即长而薄),腹侧膨大。本文简要讨论了Mesevania属的位置,以及古syncrasis属的有效性,这两个属都是在中白垩世克钦琥珀中描述的。
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引用次数: 0
Middle Eocene Ostracods from southeast Fayoum area, Egypt: Systematics, paleobathymetry and paleobiogeography 埃及法尤姆东南部始新世中期介形虫:系统学、古测深学和古生物地理学
IF 1 4区 地球科学 Q4 PALEONTOLOGY Pub Date : 2023-01-01 DOI: 10.1016/j.annpal.2023.102598
Sobhi A. Helal , Sherif M. El Baz

This study deals with the Middle Eocene ostracods from southeast Fayoum area, Egypt. To carry out this investigation, three outcrops are examined, Gebel Elwe El Breig, Gebel Sath El Hadid and Gebel Munqar El-Shinnara. This study is based on the Midawara Formation that consists mainly of fossiliferous limestone, marly limestone and clay. The studied ostracod assemblage includes thirty-one species and subspecies, belonging to twenty-three genera and seventeen families. The ostracod group belonging to Cytheroidea Baird, 1850 represents the most abundant group that reaches 81.99% in Munqar El-Shinnara section, 81.88% in Sath El Hadid section, and 61.12% in Elwe El Breig section. The structure of the investigated assemblages (abundance of ostracods, number of taxa, Fisher's alpha, Shannon index, and equitability) indicates deposition in an environment belonging to the neritic zone. Moreover, this study tries to use the multivariate analyses to distinguish the paleobiogeographic provinces in Tethyan region during the Middle Eocene. These analyses are applied on a matrix consists of some nominated Middle Eocene species from 13 countries. The results point to two distinctive provinces, Southern Tethys Province (Algeria, Tunisia, Libya, Egypt and Jordan) and the Northern Tethys Province (England, Germany, Spain, France, Belgium, Ukraine, Hungary and Turkey). The reasonable similarities between these provinces may reflect a possible connection between both sides of the Tethys during the Middle Eocene age.

本文研究了埃及法尤姆东南部地区中始新世介形类。为了进行这项调查,考察了三个露头:Gebel Elwe El Breig、Gebel Sath El Hadid和Gebel Munqar El- shinnara。本研究以Midawara组为研究对象,该组主要由化石灰岩、泥灰岩和粘土组成。所研究的介形类包括17科23属31种和亚种。属Cytheroidea Baird, 1850的介形虫群数量最多,Munqar El- shinnara剖面为81.99%,Sath El Hadid剖面为81.88%,Elwe El Breig剖面为61.12%。所调查组合的结构(介形虫丰度、分类群数量、Fisher’s alpha、Shannon指数和公平性)表明沉积环境属于浅海带。此外,本研究尝试运用多元分析方法来区分特提斯地区中始新世的古生物地理省。这些分析应用于一个由来自13个国家的一些指定的中始新世物种组成的矩阵。结果指向两个不同的省份,南特提斯省(阿尔及利亚、突尼斯、利比亚、埃及和约旦)和北特提斯省(英国、德国、西班牙、法国、比利时、乌克兰、匈牙利和土耳其)。这些省份之间合理的相似性可能反映了始新世中期特提斯河两岸之间可能存在的联系。
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引用次数: 0
A new species of glypheid lobster, Glyphea pisuergae (Crustacea, Glypheoidea), from the Early Jurassic of Palencia, Basque-Cantabrian Basin, Spain 西班牙巴斯克-坎塔布里亚盆地帕伦西亚早侏罗世的一种新的雕纹龙虾,皮纹龙虾(甲壳纲,雕纹总科)
IF 1 4区 地球科学 Q4 PALEONTOLOGY Pub Date : 2023-01-01 DOI: 10.1016/j.annpal.2023.102596
Sylvain Charbonnier , Alessandro Garassino , Mikel A. López-Horgue

A new glypheid lobster (Crustacea, Glypheidae), Glyphea pisuergae sp. nov. is described from the Early Jurassic (late Pliensbachian–early Toarcian) of Salinas de Pisuerga, Palencia, Spain. This species represents the second record for the genus in the Early Jurassic.

西班牙Palencia的Salinas de Pisuerga早侏罗世(pliensbachia - Toarcian早期)描述了一种新的形虾(甲壳纲,形虾科),Glyphea pisuergae sp. nov.。该物种代表了早侏罗世该属的第二个记录。
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引用次数: 0
A new Miocene and Pleistocene continental locality from Nakhon Ratchasima in Northeastern Thailand and its importance for vertebrate biogeography 泰国东北部那空拉差斯玛的中新世和更新世大陆新地点及其对脊椎动物生物地理学的重要性
IF 1 4区 地球科学 Q4 PALEONTOLOGY Pub Date : 2023-01-01 DOI: 10.1016/j.annpal.2023.102659
Wilailuck Naksri , Yuichiro Nishioka , Jaroon Duangkrayom , Grégoire Métais , Naoto Handa , Pratueng Jintasakul , Jeremy E. Martin , Saitong Sila , Wongsakon Sukdi , Kriangkrai Suasamong , Haiyan Tong , Julien Claude
Two new fossil assemblages are described from Sin Charoen sandpit exploited in Phimai District, Nakhon Ratchasima Province, Northeastern Thailand. In this province, vertebrate fossils have been yielded from the Miocene to the Pleistocene deposits. The Miocene fauna has been documented in detail by a rich fossil record from several localities in Chaloem Phra Kiat District, while the Pleistocene fauna has been well studied in Khok Sung sandpit in Mueang Nakhon Ratchasima District. The fossil assemblage from the upper section of Sin Charoen sandpit is similar to the Middle Pleistocene fauna of Khok Sung, which is composed of extinct and extant species of mammals and reptiles. Moreover, the finding of Gavialis cf. bengawanicus and Duboisia santeng demonstrates a strong faunal relationship with Early-Middle Pleistocene faunas of Java (e.g., Trinil). The occurrence of large fluviatile turtles and gharials from Sin Charoen sandpit indicates that the past river system was more developed than today and that the Khorat Plateau was at a lower elevation at the time of deposition. The fossil assemblage from the lower section includes two rhinocerotids and an advanced form of the genus Stegolophodon and is similar to the Late Miocene fauna found in situ in the sandpits of Chaloem Phra Kiat District.
本文描述了泰国东北部呵叻府披迈县 Sin Charoen 沙坑出土的两个新化石群。在该省,从中新世到更新世的沉积物中都发现了脊椎动物化石。中新世动物群在 Chaloem Phra Kiat 县的几个地方都有丰富的化石记录,对其进行了详细记录,而更新世动物群则在 Mueang Nakhon Ratchasima 县的 Khok Sung 沙坑进行了深入研究。Sin Charoen 沙坑上段的化石群与 Khok Sung 的中更新世动物群相似,都由哺乳动物和爬行动物的已灭绝和现存物种组成。此外,与爪哇早-中更新世动物群(如特立尼达)的关系也很密切。Sin Charoen 沙坑中出现的大型流纹龟和长吻龙表明,过去的河流系统比现在发达,而且沉积时的呼叻高原海拔较低。下段的化石群包括两种犀牛类和一种高级的犀牛属动物,与在 Chaloem Phra Kiat 区沙坑中就地发现的晚中新世动物群相似。
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引用次数: 0
A new enantiornithine bird from Upper Cretaceous non-marine deposits at Villespassans (Hérault, southern France) 一种来自上白垩纪Villespassans(Hérault,法国南部)非海洋沉积物的新对映鸟类
IF 1 4区 地球科学 Q4 PALEONTOLOGY Pub Date : 2023-01-01 DOI: 10.1016/j.annpal.2022.102585
Eric Buffetaut , Delphine Angst , Haiyan Tong

A coracoid of an enantiornithine bird from Upper Cretaceous (probably late Campanian) fluvial sediments at Castigno (Villespassans, Hérault, southern France) is described. It differs from all hitherto reported enantiornithine coracoids and is referred a new genus and species, Castignovolucris sebei. This bone is large and robust, indicating a bird that was among the largest known enantiornithines, possibly the size of a Canada Goose (Branta canadensis). The new taxon is an addition to the short list of Late Cretaceous birds from France and confirms that enantiornithines were an important component of European avifaunas until late in the Cretaceous.

描述了在法国南部hsamault的Castigno (Villespassans, hsamault)的上白垩纪(可能是坎帕尼亚晚期)河流沉积物中发现的一种反鸟目鸟类的喙。它不同于迄今报道的所有反鸟氨酸喙虫,被称为新属和新种,Castignovolucris sebei。这块骨头又大又结实,表明它是已知最大的对鸟目鸟类之一,可能和加拿大鹅一样大。这个新分类群是法国白垩纪晚期鸟类名单上的一个新成员,它证实了在白垩纪晚期之前,对鸟目是欧洲鸟类的一个重要组成部分。
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引用次数: 0
New material of turtles from the Upper Jurassic of Phu Noi, NE Thailand: Phylogenetic implications 泰国东北部 Phu Noi 上侏罗世的龟类新材料:系统发育的影响
IF 1 4区 地球科学 Q4 PALEONTOLOGY Pub Date : 2023-01-01 DOI: 10.1016/j.annpal.2023.102656
Haiyan Tong , Phornphen Chanthasit , Wilailuck Naksri , Suravech Suteethorn , Julien Claude
New shell material of Phunoichelys thirakhupti and Kalasinemys prasarttongosothi are reported from their type locality, Phu Noi site, Khorat Plateau, NE Thailand. These new specimens complete our knowledge of their anatomy. Intraspecific and ontogenetic variations are observed in P. thirakhupti; and reconstruction of the shell is provided for both species. Both Phunoichelys and Kalasinemys are aquatic turtles, but their shell morphology is distinct, suggesting different mode of life. A phylogenetic scenario is presented where the previously character based Xinjiangchelyidae appear to be paraphyletic, suggesting more phylogenetically diverse lineages were present in Eastern and South-east Asia during the Jurassic.
我们从泰国东北部呼叻高原的普诺伊(Phu Noi)海龟模式产地带回了新的海龟甲壳。这些新标本增加了我们对其解剖学的了解。我们观察到了这两个物种在种内和个体发育过程中的变异,并提供了它们的腕部重建图。根据之前推导的特征,提出了新疆蜓科的系统发育设想,即新疆蜓科属于旁系,这表明侏罗纪时期东亚和东南亚存在更多的系统发育世系。
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引用次数: 0
Transgressing the limits of palaeoenvironmental data for Southeast Asian Pleistocene faunal assemblages: A critical review to go further 超越东南亚更新世动物群古环境数据的限制:为更进一步而进行的严格审查
IF 1 4区 地球科学 Q4 PALEONTOLOGY Pub Date : 2023-01-01 DOI: 10.1016/j.annpal.2023.102657
Valéry Zeitoun , Chinnawut Winayalai , Prasit Auetrakulvit , Régis Debruyne , Jean-Baptiste Mallye , Arnaud Lenoble
For the Pleistocene of South-east Asia, the ‘savannah corridor’ hypothesis put forward by Lawrence Heaney in 1991 on the basis of a few pollen records rightly aroused a certain amount of enthusiasm and was the source of many attempts to confirm or refute a significant regression in forest cover, depending on the taxa considered, during the Last Glacial Maximum. The work carried out to verify this hypothesis used a number of tools, including palaeontology, the results of which seem to us to have been greatly embellished given the paucity of reliable data available. Indeed, it is clear that the hypothesis concerns a geographical area that is largely inaccessible today, located at a depth of 120 meters in the South China Sea, and that the faunal assemblages taken into account remain imperfectly defined to allow such a hypothesis to be tested. We provide a summary of the main work carried out to date and a critical assessment of the articles published over the last few decades. It emerges that the fossil series considered (essentially mammals) are neither well chronologically set nor well defined from a taphonomic point of view, notably because they are based on unsatisfactory references, which are themselves poorly dated or based on mixtures of different faunas. These shortcomings also call into question the renewed and attractive hypothesis of the flexibility of the diet and change of habitat (ecological niche) of certain taxa, based on isotopic analyses, as recently proposed. We advocate the implementation of systematic fine excavations that take into account the taphonomy of the sites, but also show that progress has been made to go further.
关于东南亚的更新世,劳伦斯-海尼(Lawrence Haeney)于 1991 年根据一些花粉记录提出的 "热带草原走廊 "假说引起了人们一定程度的热情,许多人试图以此来证实或反驳在末次冰川最盛时期森林覆盖率的显著下降(取决于所考虑的分类群)。为验证这一假说而开展的工作使用了多种工具,特别是古生物学工具,但在我们看来,由于可用的可靠数据极少,这些工具的结果似乎被大大美化了。事实上,很明显,该假说涉及的地理区域今天基本上无法进入,位于南中国海 120 米深处,而且所考虑的动物群落仍不完善,无法对该假说进行检验。我们对迄今为止开展的主要工作进行了总结,并对过去几十年发表的文章进行了批判性评估。所考虑的化石系列(主要是哺乳动物)似乎既没有很好的年代学设定,也没有从岩石学的角度进行很好的界定,这主要是因为它们所依据的参考文献并不令人满意,这些参考文献本身就年代不清或依据的是不同动物群的混合物。这些缺陷也使最近提出的基于同位素分析的某些类群饮食灵活性和栖息地(生态位)变化的新的诱人假说受到质疑。我们主张进行系统的精细发掘,既要考虑到遗址的岩石学,又要表明已经取得了重大进展。
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引用次数: 0
Microbial influence in the Messinian sedimentation: Example of Cap Bon (NE Tunisia) 微生物对墨西尼亚沉积的影响:以突尼斯东北部的Cap Bon为例
IF 1 4区 地球科学 Q3 Earth and Planetary Sciences Pub Date : 2023-01-01 DOI: 10.1016/j.annpal.2023.102600
Jean-Paul Saint Martin , Sihem Hlel , Juliette Debrie , Nadia Ben Moktar , Simona Saint Martin , Beya Mannai-Taiech

Messinian sedimentation in Tunisia is characterized by the absence of large carbonate platforms known in many Mediterranean marginal basins. As a result, correlations are not easy to establish in order to integrate it into a general pattern of development at the regional level. However, the discovery of microbial constructions of metric scale makes it possible to precise uncertain points. Thus in the sector of Cap Bon in the northeast of Tunisia, stromatolitic constructions can form, over distances of several hundred meters, domes containing columnar structures at different scales as well as carbonate beds showing sedimentary ripples also of microbialite nature. The highlighting of an assemblage of microbialites, associated with oolitic deposits, allows linking these formations to the Terminal Carbonate Complex which finds its place at a part of the so-called Messinian Salinity Crisis.

突尼斯的迈西尼亚沉积的特点是在许多地中海边缘盆地中缺乏已知的大型碳酸盐台地。因此,不容易建立相互关系,以便将其纳入区域一级的一般发展格局。然而,公制尺度微生物结构的发现使得精确测量不确定点成为可能。因此,在突尼斯东北部的Cap Bon地区,叠层石结构可以在几百米的距离上形成包含不同规模柱状结构的圆顶,以及具有微生物岩性质的沉积波纹的碳酸盐层。与鲕状沉积物相关的微生物岩组合的突出显示,可以将这些地层与终端碳酸盐复合体联系起来,该复合体位于所谓的墨西尼亚盐危机的一部分。
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引用次数: 0
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Annales de Paleontologie
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