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Additions to the phytoplankton of water bodies of the Krasnoyarsk Pillars National Park 克拉斯诺亚尔斯克支柱国家公园水体浮游植物的增加
Pub Date : 2023-07-03 DOI: 10.14258/pbssm.2023082
K. A. Eichvald, O. Bazhenova
The article presents new data on the species composition and taxonomic structure of phytoplankton of some water bodies of the Krasnoyarsk Pillars National Park according to studies during the open water period of 2022. 95 species and intraspecific taxa were identified, including the taxonomic type of the species, including: Cyanoprokaryota - 7, Dinophyta - 1, Chrysophyta - 5, Euglenophyta - 10, Bacillariophyta - 42, Chlorophyta - 27, Charophyta - 3. 41 new taxa with a rank below the genus have been identified for the territory, including Cyanoprokaryota - 4, Dinophyta - 1, Chrysophyta - 2, Euglenophyta - 8, Bacillariophyta - 10, Chlorophyta - 15, Charophyta - 1.
文章介绍了根据 2022 年开放水域期间的研究得出的关于克拉斯诺亚尔斯克支柱国家公园部分水体浮游植物物种组成和分类结构的新数据。确定了 95 个物种和种内类群,包括物种的分类类型,其中包括其中包括:蓝藻门(Cyanoprokaryota)--7 个、叶绿体门(Dinophyta)--1 个、菊形叶绿体门(Chrysophyta)--5 个、叶绿体门(Euglenophyta)--10 个、芽孢杆菌门(Bacillariophyta)--42 个、叶绿体门(Chlorophyta)--27 个、藻类门(Charophyta)--3 个。在境内新发现了 41 个低于属级的类群,其中包括蓝藻门(Cyanoprokaryota)- 4 个、鼎藻门(Dinophyta)- 1 个、菊形门(Chrysophyta)- 2 个、优生优育门(Euglenophyta)- 8 个、芽孢杆菌门(Bacillariophyta)- 10 个、叶绿体门(Chlorophyta)- 15 个、叶绿体门(Charophyta)- 1 个。
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引用次数: 0
Assessment of the population status of the relict species Lomelosia austroaltaica (Bobrov) Sojak in the flora of the Kazakstan Altai 评估哈萨克斯坦阿尔泰植物区系中孑遗物种 Lomelosia austroaltaica (Bobrov) Sojak 的种群状况
Pub Date : 2023-07-03 DOI: 10.14258/pbssm.2023022
A. N. Danilova, Yury A. Kotukhov, O. A. Anufrieva, A. Sumbembayev
The article discusses the population parameters of a rare relict species Lomelosia austroaltaica in the territory of the Kalba Upland in the Kazakhstan Altai. It has been established that the population is located in the Eastern Kalba on the Medvedka Mountain (Koktau Mountains) as part of the feather grass-lomelosia phytocenosis. The total area occupied by phytocenosis with the participation of Lomelosia austroaltaica, is 2.5 hectares. Vegetation cover is poorly formed, represented by 30 species of flowering plants. The structure and appearance of the phytocenosis are determined by the dominants Stipa capillata- cop2, S. pennata - cop1. In ecological terms, the species composition of the higher vascular plants of the surveyed phytocenosis is formed by xeropetrophytes - 20 species (66.7 %), xeromesophytes - 7 species (23.3 %), mesophytes - 3 species (10 %). The total projective cover is 44.4 %, the proportion of Lomelosia austroaltaica in the phytobiota is 22.2 %, i. e. according to its coenotic features, the species is an assectator. Individuals are scattered and unevenly distributed over the area, their height ranges from 28 to 48 cm (33.6 ± 7.4.6), forming loose sods 47.9 ± 5.2 cm in diameter from 46.3 ± 7.2 cm vegetative and 51.3 ± 10.5 cm generative shoots. A direct dependence of seed productivity on the number of formed heads on a generative shoot was revealed. Mass flowering of individuals was recorded in the second decade of June, the period of seed ripening is late August - early September. The population is normal, incomplete, the age spectrum is left-sided. The ratio of young and adult fractions (pl + j + im) : (v + g) = 11.8 : 7.8 indicates a stable state of the population at the present stage. Mainly seeds maintain the number of individuals. The species needs state protection.
文章讨论了哈萨克斯坦阿尔泰地区卡尔巴高地的稀有孑遗物种 Lomelosia austroaltaica 的种群参数。已确定该种群位于东卡尔巴的梅德韦卡山(科克套山)上,是羽毛草-Lomelosia植物群落的一部分。在奥斯特罗尔塔洛美洛斯(Lomelosia austroaltaica)的参与下,植物区系占据的总面积为 2.5 公顷。植被覆盖率很低,只有 30 种开花植物。植物区系的结构和外观是由占优势的穗状花序(Stipa capillata)--cop2、穗状花序(S. pennata)--cop1 决定的。从生态学角度来看,所调查的植物群落中高等维管束植物的物种组成为旱生植物--20 种(66.7%)、旱生植物--7 种(23.3%)、中生植物--3 种(10%)。总投影覆盖率为 44.4%,Lomelosia austroaltaica 在植物区系中所占比例为 22.2%,也就是说,根据其同源特征,该物种属于食虫植物。个体分散且不均匀地分布在整个区域,高度从 28 厘米到 48 厘米不等(33.6 ± 7.4.6),由 46.3 ± 7.2 厘米的无性芽和 51.3 ± 10.5 厘米的生成芽形成直径为 47.9 ± 5.2 厘米的松散草皮。结果表明,种子产量直接取决于生成芽上形成的头数。个体的大规模开花期在 6 月下旬,种子成熟期在 8 月底至 9 月初。种群正常,不完整,年龄谱呈左侧分布。幼虫和成虫的比例(pl + j + im):(v + g)= 11.8:7.8,表明现阶段种群处于稳定状态。主要由种子维持个体数量。该物种需要国家保护。
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引用次数: 0
Genetic variability of populations of Pulsatilla species (Ranunculaceae) in the south of Yenisei Siberia 叶尼塞西伯利亚南部白头翁属(Ranunculaceae)种群的遗传变异性
Pub Date : 2023-07-03 DOI: 10.14258/pbssm.2023028
V. V. Zhivotov, K. K. Ryabova, I. E. Yamskikh, N. V. Stepanov, M. G. Kutsev
Pulsatilla is a genus of ephemeroid grasses characterized by high morphological diversity. The study assessed the genetic polymorphism of 9 populations represented by the species P orientali-sibirica Stepanov, P turczaninovii Krylov and their hybrid P. herba-somnii Stepanov, and P. multifida G. Prizel. As a result of amplification of genomic DNA with 4 ISSR primers, 61 DNA amplicons were obtained. The overall level of polymorphism is 98.36 %. The number of amplified DNA fragments, depending on the primer, varies from 11 (17899A) to 17 (17899 B, HB12). The maximum level of genetic variability was noted for hybrid populations. The coefficient of genetic subdivision (Gst) of the studied populations is 0.4718, which indicates a high level of their differentiation. The dendrogram divides the studied populations into 3 groups, each of which is represented by a separate species, while the hybrid population is combined with P. orientali-sibirica. Phylogenetic analysis of the studied populations combines P orientali-sibirica with P multifida and P turczaninovii with P. herba-somnii. These data demonstrate the intermediate position of the hybrid population of P. herba-somnii. In the study of hybrids, ISSRS are the most informative, since they are evenly distributed across the genome and can show similarities with two parent organisms, whereas chloroplasts are inherited, as a rule, through the maternal line.
白头翁属(Pulsatilla)是一种昙花草属植物,具有形态多样性的特点。本研究评估了以 P orientali-sibirica Stepanov、P turczaninovii Krylov 及其杂交种 P. herba-somnii Stepanov 和 P. multifida G. Prizel 为代表的 9 个种群的遗传多态性。用 4 个 ISSR 引物扩增基因组 DNA 后,得到 61 个 DNA 扩增子。总体多态性水平为 98.36%。根据引物的不同,扩增 DNA 片段的数量从 11 个(17899A)到 17 个(17899B,HB12)不等。杂交种群的遗传变异水平最高。研究种群的遗传细分系数(Gst)为 0.4718,这表明它们的分化程度很高。树枝图将所研究的种群分为 3 组,每组代表一个单独的物种,而杂交种群则与 P. orientali-sibirica 结合在一起。所研究种群的系统发生分析将 P orientali-sibirica 与 P multifida 和 P turczaninovii 与 P. herba-somnii 结合在一起。这些数据证明了 P. herba-somnii 杂交种群的中间位置。在杂交种的研究中,ISSRS 的信息量最大,因为它们在基因组中分布均匀,能显示出与两个亲本生物的相似性,而叶绿体通常是通过母系遗传的。
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引用次数: 0
Genetic mechanisms of progressive evolution and speciation in plants: Gene multiplication as a way of an adequate response of the genome to the action of environmental factors 植物逐步进化和物种分化的遗传机制:基因繁殖是基因组对环境因素作用做出适当反应的一种方式
Pub Date : 2023-07-03 DOI: 10.14258/pbssm.2023061
Родионов А. В Rodionov
One of the mechanisms related to the progressive evolution of plants probably is gene duplications. Paralogous and orthologous gene copies can appear in the genome either as a result of whole genome duplication (WGD), or after segmental or tandem duplications. The proportion of duplicated genes among the protein-coding genes in plant genomes is high (64 % on average), but it is several times less than would be expected if all genes duplicated during WGD were preserved - all plant genomes in that or otherwise lost some of the duplicated genes and transposons. Due to the need to respect the gene dose, for duplicated genes resulting from WGD, the rule applies: genes whose products work as part of multiprotein complexes are preserved, genes following the rule “one gene - one enzyme” prefer a monogenic existence. On the contrary, if duplicated copies of genes appeared in the genome as a result of tandem or segmental duplication, then in the first place, for the same reason, “extra” copies of the genes of the components of heteromeric complexes are lost. The role of tandem duplications in plant evolution, apparently, consists primarily in the fact that they correct the results of the distribution of alleles that remains in the genome of a polyploid or diploid as a result of processes of secondary diploidization and fractionation of neopolyploid genomes, stochastic changes in the set of alleles in comparison with stabilized (passed a long selection).
与植物逐步进化有关的机制之一可能是基因复制。基因组中的旁系和同源基因副本可能是全基因组复制(WGD)的结果,也可能是节段或串联复制的结果。在植物基因组的蛋白质编码基因中,重复基因所占的比例很高(平均为 64%),但如果在 WGD 过程中重复的所有基因都被保留下来,那么重复基因所占的比例就会比预期的低几倍--在这种情况下,所有植物基因组都会丢失一些重复基因和转座子。由于需要尊重基因剂量,对于 WGD 产生的重复基因,适用的规则是:其产物作为多蛋白复合物的一部分起作用的基因被保留下来,遵循 "一个基因-一种酶 "规则的基因倾向于单源存在。相反,如果基因的复制拷贝是由于串联复制或片段复制而出现在基因组中,那么出于同样的原因,首先会丢失异源复合体成分基因的 "额外 "拷贝。显然,串联复制在植物进化中的作用主要在于,它们纠正了多倍体或二倍体基因组中因二次二倍体化和新多倍体基因组分化过程而残留的等位基因的分布结果,以及等位基因集与稳定的(经过长期选择的)等位基因集相比发生的随机变化。
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引用次数: 0
Distribution of Rhaponticum serratuloides in Kazakhstan 蚕沙皂苷在哈萨克斯坦的分布情况
Pub Date : 2023-07-03 DOI: 10.14258/pbssm.2023041
S. Mamyrova
The distribution of Rhaponticum serratuloides on the territory of Kazakhstan has been studied. Rh serratuloides is a typical steppe species with a Pontic-Kazakh-Altai type of habitat. In Kazakhstan, it grows in the steppe zone with a rare occurrence in the northern deserts (Balkhash-Adakol floral district), where it occurs in saline and saline meadows, banks and bottoms of drying salt lakes and rivers. As a result of field research and viewing materials of herbariums AA, LE, MW, KUZ, KG, NUR, the species was found in 13 floristic districts and subdistricts of Kazakhstan. The species was found for the first time in the Tarbagatai floristic district.
研究了哈萨克斯坦境内蚕豆草(Rhaponticum serratuloides)的分布情况。Rh serratuloides 是典型的草原物种,栖息地为庞迪-哈萨克-阿尔泰类型。在哈萨克斯坦,它生长在草原地带,很少出现在北部沙漠(巴尔喀什-阿达科尔花区),在那里,它出现在盐碱和盐碱草地、干涸的盐湖和河流的岸边和底部。通过实地考察和查阅标本馆 AA、LE、MW、KUZ、KG、NUR 的资料,在哈萨克斯坦的 13 个花卉区和分区发现了该物种。该物种首次出现在塔尔巴加泰花卉区。
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引用次数: 0
Materials to the creation of the “Klyukvennoe Lake” Natural Monument of regional importance (Altai Krai) 创建 "克柳克文诺湖 "地区重要自然纪念碑的材料(阿尔泰边疆区)
Pub Date : 2023-07-03 DOI: 10.14258/pbssm.2023026
N. Elesova
Materials on the flora and vegetation of the planned Nature Monument of regional importance “Klyukvennoe Lake” are given. Flora comprises 269 species of higher vascular plants, including 2 rare species: Dryopteris cristata, Sphagnum teres and 2 species belonging to the Appendix 1 of the Altai Krai Red Data Book (Plants and fungi taxa reguire a special attention to their.
提供了有关规划中的具有地区重要性的自然纪念碑 "克柳克文诺湖 "的植物区系和植被的资料。植物区系包括 269 种高等维管植物,其中有 2 种稀有物种:其中包括 2 个稀有物种:Dryopteris cristata、Sphagnum teres 和 2 个属于《阿尔泰边疆区红色数据手册》附录 1 的物种(植物和真菌类群需要特别关注。
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引用次数: 0
Features of the morphology of seeds of species of the genus Suaeda flora of the Aral-Balkhash region 咸海-巴尔喀什地区苏埃达属植物种子的形态特征
Pub Date : 2023-07-03 DOI: 10.14258/pbssm.2023075
S. Ussen, P. Vesselova, G. Kudabayeva
Genus Suaeda Forssk. ex J. F. Gmel. is one of the largest in the family Chenopodiaceae Vent. (Amaranthaceae Juss.). Its representatives are distributed throughout Central Asia and are found, as a rule, in coastal halophyte communities. Out of 40 Central Asian species, there are 18 species in the flora of Kazakhstan, 15 (83 %) of which are found in the Aral-Balkhash region. Suaeda species are distributed in 5 sections (Schanginia C. A. Mey., Lachnostigma Iljin, Conosperma Iljin, Physophora Iljin, Heterosperma Iljin). The aim of the research was to analyze the features of the morphology of seeds of species of the genus Suaeda of the flora of the Aral-Balkhash region, which includes 5 floristic districts (14 - Aral, 15 -Kyzylorda, 16 - Betpakdala, 17 - Muyunkum (Moyinkum) and 18 - Balkhash-Alakul). Of the 15 species found within the Aral-Balkhash region, seeds of 11 species were taken for comparative analysis of morphological features (Suaeda linifolia Pall., S. paradoxa Bunge, S. altissima (L.) Pall., S. dendroides (С. A. Mey.) Moq., S. arcuata Bunge, S. acuminata (С. А. Меу.) Moq., S. corniculata (С. А. Меу.) Bunge, S. heterophylla (Kar. et Kir.) Bunge, S. salsa (L.) Pall., S. prostrata Pall., S. crassifolia Pall.). A detailed study reveals sectional differences. In this respect, the seeds of Suaeda acuminata differ most clearly from the species of other sections. from the Conosperma section. The species of the Schanginia section have a generally similar granular structure of the seed surface, and the representatives of the Heterosperma section have a point-grid surface of seeds with a flattened edge.
Suaeda Forssk.其代表植物分布在整个中亚地区,通常生长在沿海卤叶植物群落中。在 40 种中亚物种中,哈萨克斯坦植物区系中有 18 种,其中 15 种(83%)分布在咸海-巴尔喀什地区。Suaeda 种类分布在 5 个部分(Schanginia C. A. Mey.、Lachnostigma Iljin、Conosperma Iljin、Physophora Iljin、Heterosperma Iljin)。研究的目的是分析咸海-巴尔喀什地区植物区系中 Suaeda 属物种种子的形态特征,该地区包括 5 个植物区系(14 - 咸海、15 - 基兹洛达、16 - 贝特帕克达拉、17 - 穆因库姆(莫因库姆)和 18 - 巴尔喀什-阿拉库尔)。在咸海-巴尔喀什地区发现的 15 个物种中,有 11 个物种的种子被用于形态特征比较分析(Suaeda linifolia Pall.、S. paradoxa Bunge、S. altissima (L.) Pall、S. dendroides (С. A. Mey.) Moq.、S. arcuata Bunge、S. acuminata (С. А. Меу.) Moq.、S. corniculata (С. А. Меу.) Bunge、S. heterophylla (Kar. et Kir.) Bunge、S. salsa (L.) Pall.、S. prostrata Pall.、S. crassifolia Pall.)。详细的研究显示了节间差异。在这方面,Suaeda acuminata 的种子与其他科的物种差异最为明显。Schanginia 部分的物种种子表面的颗粒结构基本相似,而 Heterosperma 部分的代表物种种子表面呈点状网格状,边缘扁平。
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引用次数: 0
Algae flora of the Lower Irtysh and its tributary, the Tobol River 额尔齐斯河下游及其支流托博尔河的藻类植物群
Pub Date : 2023-07-03 DOI: 10.14258/pbssm.2023066
L. A. Semenova, M. S. Bondar
The results of algoflora studying the Lower Irtysh and its tributary, the Tobol River, are presented. A total of 432 taxa of species and subspecies ranks were identified from 124 of 58 families and 8 departments. For the first time, a systematic list of algae is given. The greatest species and intraspecific richness were characterized by green, diatoms, blue-green (cyanoprokaryotes) and euglenic algae (93 % of the total list). Planktonic species, cosmopolitan species and beta-mesosaprobes predominated. In the phytoplankton of the mouth of the Irtysh, 368 algae taxa with a rank below the genus were registered, in the modern period in the lower reaches - 247, in the Tobol River - 192. The algae of the Lower Irtysh plankton were distinguished by a large variety of green algae and an abundance of diatoms, in the Tobol River -cyanoprokaryotes (August). The estuary section of the Irtysh River was distinguished by the highest level of quantitative development of phytoplankton, diatomaceous plankton. The modern period is characterized by the development of small-cell green algae, diatom-chlorococcal plankton. A high level of development of summer algocenosis caused by the vegetation of cyanoprokaryotes and green algae was observed in the Tobol River. The appearance of cryptomonads in the 2000s indicates organic pollution of the water masses of the surveyed rivers. The obtained materials have a certain scientific and practical value and can be used in biomonitoring.
本文介绍了研究额尔齐斯河下游及其支流托博尔河的生物群落的结果。共鉴定出 432 个种级和亚种级分类群,分别来自 58 个科和 8 个系中的 124 个科。首次给出了藻类的系统列表。物种和种内丰富度最高的是绿藻、硅藻、蓝绿藻(蓝藻)和优藻(占总清单的 93%)。浮游物种、世界性物种和β-介囊藻占多数。在额尔齐斯河河口的浮游植物中,记录了 368 个属以下的藻类类群,在现代时期,在下游记录了 247 个类群,在托博尔河记录了 192 个类群。额尔齐斯河下游浮游生物中的藻类以种类繁多的绿藻和丰富的硅藻而著称,而在托博尔河则以蓝藻为主(八月)。额尔齐斯河河口段的特点是浮游植物、硅藻浮游生物的数量发展水平最高。现代时期的特点是小细胞绿藻、硅藻-氯球藻浮游生物的发展。在托博尔河(Tobol River)观察到,由蓝藻和绿藻植被引起的夏季藻类增生的发展水平很高。2000 年代出现的隐单胞菌表明调查河流的水体受到了有机污染。所获材料具有一定的科学价值和实用价值,可用于生物监测。
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引用次数: 0
Diversity in the morphology of clones of witches’ broom in Siberian stone pine 西伯利亚石松中女巫扫帚克隆形态的多样性
Pub Date : 2023-07-03 DOI: 10.14258/pbssm.2023052
O. I. Polyakova, E. A. Zhuk
Mutational witches’ broom (WB) is a fragment of a tree crown with abnormally dense branching and slow shoot growth compared with those of a normal crown. All WBs have thicker and denser crown compared to the normal crown part of their parental tree, but the crown density differs significantly among WBs. There are both relatively loose WBs, whose crown density differ from normal crown part not very strong, and WBs with a very dense crown. The aim of this study was to conduct a comparative analysis of morphological traits in Pinus sibirica WB clones with different crown densities and to determine which traits contribute most to their differences. The object of the study was clones from 11 mutational WB with different crown density. The density of WB was considered low if it exceeded the density of a normal crown by less than 2 times, medium - by 2-3 times, high - by more than 3 times. Morphological characteristics of crown and shoots were measured in 12-year-old clones. It was found that the higher the WB clone crown density the smaller and less elongated its crown shape, as well as less crown continuity. The basis of this phenotype formation was primarily the shorter length of annual shoots, the reduced level of apical dominance compared to other clone groups, and advanced branching. An additional factor that increased crown density was the significantly shorter internodes compared to the other clone groups. The needle length did not contribute to the differences in crown density of the clones.
变异女巫扫帚(WB)是树冠的一个片段,与正常树冠相比,它的分枝异常密集,嫩枝生长缓慢。与亲本树的正常树冠部分相比,所有 WB 的树冠都更粗更密,但不同 WB 的树冠密度差异很大。既有树冠密度与正常树冠部分相差不大的相对疏松的 WB,也有树冠非常密集的 WB。本研究的目的是对具有不同树冠密度的西伯利亚红松 WB 克隆的形态特征进行比较分析,并确定哪些特征对它们之间的差异贡献最大。研究对象是来自 11 个不同树冠密度的变异西伯利亚红松克隆。如果WB的密度超过正常树冠密度的2倍以下,则被视为低密度;超过正常树冠密度的2-3倍,则被视为中密度;超过正常树冠密度的3倍,则被视为高密度。对 12 年克隆的树冠和嫩枝的形态特征进行了测量。结果发现,WB 克隆的树冠密度越高,其树冠形状就越小、越不修长,树冠的连续性也越差。这种表型形成的基础主要是一年生枝条长度较短、顶端优势程度比其他克隆组低,以及分枝能力较强。增加树冠密度的另一个因素是节间明显短于其他克隆组。针叶长度并没有造成克隆树冠密度的差异。
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引用次数: 0
Distribution of fungi of the genera Nectria and Neonectria in the southeast of Kazakhstan 哈萨克斯坦东南部 Nectria 和 Neonectria 属真菌的分布情况
Pub Date : 2023-07-03 DOI: 10.14258/pbssm.2023004
A. M. Assylbek, Y. Rakhimova, L. Kyzmetova, G. Sypabekkyzy
The article presents data on the distribution of microscopic fungi of the genera Nectria and Neonectria in the territory of the south-east of Kazakhstan, developing as saprotrophs on fallen or dead branches of the host, or as wound parasites. The material for the research was the authors’ own collections on the territory of the Ile Alatau, Dzhungarian Alatau, Kungey Alatau, the Ketmen, Terskey and Altyn-Emel ridges, as well as herbarium specimens stored in the mycological herbarium of the Institute of Botany and Phytointroduction. 5 species of Nectria and Neonectria were identified, developing on 25 species of feeding plants. Most of the samples are represented by the conidial stage. The most common species is Nectria cinnabarina. Among the most affected species are representatives of the genus Ribes L., as well as the genera Acer L., Betula L., Malus Mill. and Rhamnus L. The maximum number of Nectria and Neonectria species was recorded at an altitude of 1200-1600 m above sea level, which corresponds to the belt of steppes, small-leaved and dark coniferous forests. In the steppe belt, representatives of Nectria and Neonectria are found in floodplain forests and shrubs.
文章介绍了哈萨克斯坦东南部地区Nectria属和Neonectria属显微真菌的分布数据,这些真菌在寄主倒下或枯死的树枝上以嗜渍菌的形式生长,或以伤口寄生菌的形式生长。研究材料是作者自己在 Ile Alatau、Dzhungarian Alatau、Kungey Alatau、Ketmen、Terskey 和 Altyn-Emel 山脊上采集的,以及植物学和植物诱导研究所真菌标本室保存的标本。已鉴定出 5 种 Nectria 和 Neonectria,它们生长在 25 种食料植物上。大部分样本以分生孢子阶段为代表。最常见的种类是 Nectria cinnabarina。在海拔 1200-1600 米的地区,Nectria 和 Neonectria 的数量最多,这与草原、小叶林和暗色针叶林带相对应。在草原带,Nectria 和 Neonectria 的代表出现在冲积平原森林和灌木丛中。
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引用次数: 0
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