Zou Y. Zhao P. Wu N. et al. 2025 “rarestR: An R Package Using Rarefaction Metrics to Estimate α- and β-Diversity for Incomplete Samples.” Diversity and Distributions 31: e13954. https://doi.org/10.1111/ddi.13954.
In both equations, there should be no ‘—’ in some parts. That is, the intended hypergeometric probability terms should be expressed using binomial coefficients (combinations), rather than as division.
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