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Acritarch Clusters from the Cambrian (Miaolingian) of the Příbram-Jince Basin, Czech Republic 捷克共和国Příbram-Jince盆地寒武纪(苗岭阶)的Acritach星团
IF 1.5 4区 地球科学 Q2 Earth and Planetary Sciences Pub Date : 2022-08-23 DOI: 10.1080/01916122.2022.2115574
Vojtěch Kovář, O. Fatka, J. Vodička
ABSTRACT Monogeneric clusters of five acritarch genera (Adara, Cymatiosphaera, Eliasum, Synsphaeridium and Timofeevia) were discovered in palynological residues obtained from fine-grained siliciclastic samples from the middle Cambrian (Miaolingian) Jince Formation of the Příbram–Jince Basin (Czech Republic). The clusters consist of two to more than 100 individual specimens and lack a common pattern of organisation. Acritarchs within clusters fall within a similar size range, regardless of generic affinity. Proposed mechanisms for the formation of these acritarch clusters are discussed: for the Jince Formation clusters, aggregation within algal blooms or primary colonial behaviour are the most plausible explanations. In addition, a summary of reports of acritarch clusters from lower Palaeozoic strata is included: clusters of a variety of acritarch genera have been documented from rocks of early Cambrian to Late Devonian age.
摘要/ ABSTRACT摘要:在捷克Příbram-Jince盆地中寒武世(庙岭期)Jince组的细粒硅屑孢粉残留中,发现了Adara、Cymatiosphaera、Eliasum、Synsphaeridium和Timofeevia 5个针状纲属的单属群。这些集群由2到100多个个体标本组成,缺乏共同的组织模式。集群内的Acritarchs落在相似的大小范围内,而不考虑亲缘关系。本文讨论了这些尖藻群形成的机制:对于Jince formation集群,藻华内的聚集或初级殖民地行为是最合理的解释。此外,本文还总结了下古生代岩石圈中各种各样的树状类群的报道:从早寒武世到晚泥盆世的岩石中记录了各种树状类群。
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引用次数: 1
Biostratigraphically Significant Palynofloras from the Paleocene–Eocene Boundary of the USA 美国古新世-始新世边界具有生物地层学意义的孢粉植物
IF 1.5 4区 地球科学 Q2 Earth and Planetary Sciences Pub Date : 2022-08-19 DOI: 10.1080/01916122.2022.2115159
Vera A. Korasidis, S. Wing, G. Harrington, T. Demchuk, Julia Gravendyck, P. Jardine, D. Willard
ABSTRACT Pollen and spores were recovered from the Paleocene Fort Union Formation and Paleocene–Eocene Willwood Formation of the Bighorn Basin (BHB), northwestern Wyoming, USA. In many local stratigraphic sections in the BHB, the base of the Eocene has been identified by the characteristic negative carbon isotope excursion (CIE) that marks the beginning of the Paleocene–Eocene Thermal Maximum (PETM). The palynotaxa from outcrop samples were examined using light microscopy (LM) and scanning electron microscopy (SEM). Seven new species are formally described (Tricolpites vegrandis, Rousea spatium, Striatricolporites astutus, Striatopollis calidarius, Friedrichipollis geminus, Retistephanocolporites modicrassus and Retistephanocolporites pergrandis). The temporal and geographic distributions of many of these palynotaxa suggest that hotter and more seasonally dry climates facilitated their northward range shifts during the PETM from the tropics or subtropics of the USA. For the temperate palynotaxa, the hotter and seasonally dry conditions resulted in local extirpation. A reevaluation of the palynostratigraphic schemes established for the Paleocene–Eocene boundary confirms that the first appearance of Platycarya platycaryoides denotes the Paleocene–Eocene boundary in the Rocky Mountains region. A new Striatopollis calidarius Subzone, associated with early Wasatchian (Wa) Wa-0 and Wa-R faunas, is also recognized for CIE body localities in the BHB.
摘要研究了美国怀俄明州西北部大角盆地(Bighorn Basin, BHB)古新世Fort Union组和古新世-始新世Willwood组的花粉和孢子。在BHB的许多局部地层剖面中,始新世的基底被特征负碳同位素偏移(CIE)所识别,标志着古新世-始新世极热期(PETM)的开始。利用光镜和扫描电镜对露头孢粉类群进行了研究。正式描述了7个新种(vegrandis Tricolpites、Rousea spatium、Striatricolporites astutus、Striatopollis calidarius、Friedrichipollis geminus、Retistephanocolporites modicrassus和Retistephanocolporites pergrandis)。这些孢粉植物的时间和地理分布表明,在PETM期间,炎热和季节性干燥的气候促进了它们从美国热带或亚热带向北转移。对于温带孢粉植物,炎热和季节性干燥的环境导致了局部灭绝。对古新世-始新世界线所建立的孢粉地层方案的重新评价证实,Platycarya platycaryoides的首次出现代表了落基山脉地区古新世-始新世界线。与早期Wasatchian (Wa) Wa-0和Wa- r动物群相关的一个新的Striatopollis calidarius亚带也被认为是BHB的CIE体位。
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引用次数: 2
Comments on Mertens et al. (2022): the taxonomic identity of Micracanthodinium setiferum (Lohmann) Deflandre (Dinophyceae incertae sedis) remains elusive, and its epitypification is not achieved 点评Mertens et al. (2022): Micracanthodinium setiferum (Lohmann) Deflandre (Dinophyceae incertae sedis)的分类身份仍然难以捉摸,未能实现其典型化
IF 1.5 4区 地球科学 Q2 Earth and Planetary Sciences Pub Date : 2022-07-26 DOI: 10.1080/01916122.2022.2106322
M. Gottschling, M. Elbrächter, U. Tillmann
Abstract Micracanthodinium setiferum is a very distinctive but poorly known dinophyte with characteristically delicate, unbranched, filiform and pointed setae. A recent study by Mertens et al. (2022, Palynology 46: 1934908) aims to clarify the taxonomy of the species and to introduce two new formae acknowledging the presence or absence of setae in environmentally collected cells. We assert that the epitype choice of illustrations by these authors is inappropriate, but also not achieved following the rules of the International Code of Nomenclature for algae, fungi, and plants (ICN). Additionally, the two new formae are not described in accordance with the rules of the ICN. A more suitable epitype may therefore be chosen, once corresponding physical material from the type locality off Sicily is at hand. The taxonomic identity of M. setiferum meanwhile remains elusive.
Micracanthodinium setiferum是一种非常独特但鲜为人知的恐龙植物,其特征是精致,不分枝,丝状和尖的刚毛。Mertens等人(2022,孢粉学46:1934908)最近的一项研究旨在澄清该物种的分类,并引入两种新的形式,承认在环境收集的细胞中存在或不存在刚毛。我们断言,这些作者的插图的表型选择是不适当的,但也没有实现以下规则的国际命名法藻类,真菌和植物(ICN)。此外,这两种新形式没有按照ICN的规则进行描述。因此,一旦从西西里岛附近的模式产地得到相应的物理材料,就可以选择更合适的表型。与此同时,M. setiferum的分类身份仍然难以捉摸。
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引用次数: 1
An Overview of the Palynology and Physicochemistry of Monofloral Honeys from the Dry and Humid Chaco of Argentina 阿根廷干湿查科地区单花蜂蜜的孢粉学和物理化学综述
IF 1.5 4区 地球科学 Q2 Earth and Planetary Sciences Pub Date : 2022-07-21 DOI: 10.1080/01916122.2022.2103748
Cristina Renee Salgado Laurenti, M. C. Tellería, J. M. Coronel, S. Sobrado
ABSTRACT This article is the first contribution to the characterization of monofloral honeys from the Dry and Humid Chaco that combines palynological and physicochemical parameters. The study focuses on the most represented honeys. Physicochemical parameters such as color, electrical conductivity, moisture, acidity and pH were analyzed for a total of 116 honeys classified as monofloral by pollen analysis. All the parameters considered were within the accepted range. Color and electrical conductivity were the most distinctive features, as stressed by multivariate (cluster analysis (CA) and principal component analysis (PCA)) and correlation analysis. The darkest honeys (i.e. black, dark amber and amber) were those from Tessaria integrifolia, Schinopsis balansae, S. lorentzii and Baccharis–Eupatorium-type pollen, and the lightest honeys (i.e. light amber and extra light amber) were those from Cynophalla retusa, Eugenia uniflora, Copernicia alba, Prosopis alba and Helianthus annuus. Mean values for electrical conductivity ranged from 0.4 to 0.8mS cm–1, with the highest values detected in the darkest honeys, which are mainly produced in the Dry Chaco. The intense color of these honeys also suggests the presence of antioxidants. Thus, the color, electrical conductivity and antioxidant properties are important factors for future research to consider, to obtain a reliable authentication of the botanical and geographical features of these honeys.
摘要本文首次结合孢粉学和理化参数对干湿查科单花蜂蜜进行了表征。这项研究的重点是最具代表性的蜂蜜。对116种单花蜂蜜的颜色、电导率、水分、酸度和pH等理化参数进行了分析。所有考虑的参数都在可接受的范围内。多变量聚类分析(CA)、主成分分析(PCA)和相关分析强调,颜色和电导率是最显著的特征。颜色最深的蜂蜜(即黑色、深琥珀色和琥珀色)为产自荆芥(Tessaria integrifolia)、五味子(Schinopsis balansae)、S. lorentzii和bacchari - eupatorium型花粉的蜂蜜,颜色最浅的蜂蜜(即浅琥珀色和超浅琥珀色)为产自Cynophalla retusa、Eugenia uniflora、Copernicia alba、Prosopis alba和Helianthus annuus的蜂蜜。电导率的平均值在0.4 ~ 0.8mS cm-1之间,在颜色最深的蜂蜜中检测到的值最高,这些蜂蜜主要产自干查科。这些蜂蜜的强烈颜色也表明抗氧化剂的存在。因此,颜色、电导率和抗氧化性能是未来研究需要考虑的重要因素,以获得这些蜂蜜的植物和地理特征的可靠认证。
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引用次数: 0
Revisiting R.H. Tschudy’s fern-spore spike concept 40 years later: connecting Tschudy’s ‘disaster taxon’ concept with paleopolyploidization in Stenochlaena J. Sm 40年后重新审视R.H. tschstudy的蕨类孢子穗概念:将tschstudy的“灾难分类群”概念与Stenochlaena的古多倍体化联系起来
IF 1.5 4区 地球科学 Q2 Earth and Planetary Sciences Pub Date : 2022-07-19 DOI: 10.1080/01916122.2022.2103196
K. Berry
Abstract The Cretaceous/Paleogene (K/Pg) boundary fern-spore spike concept was first introduced by R.H. Tschudy 40 years ago and established the precept that ferns are so-called ‘disaster taxa’ that flourish after natural disasters because of the high colonization potential of their wind-blown spores. Among the least understood topics at the crux of contemporary K/Pg boundary fern-spike studies is whether Stenochlaena-like or other stem blechnaceous ferns were among those that flourished after the K/Pg event, contributing to the Laevigatosporites-dominated phase of the fern-spore spike. For instance, dispersed Stenochlaena-like spores – e.g. Polypodiisporites usmensis (van der Hammen) Khan & Martin – first appear in the upper Eocene stratigraphic record of South America, whereas megafossils of Stenochlaena-like ferns are first known from the upper Paleocene strata of this same region. Beyond this traditional focus on taphonomic and taxonomic uncertainty regarding the identity of dispersed spore producers, however, there is further discordance between contemporary paleobotanical and molecular phylogenetic perspectives on the timing of diversification of stem lineages of blechnaceous ferns in relation to the K/Pg boundary. This investigation reconciles these two perspectives by constraining molecular clock divergence time estimates using contemporary fossil data. If this reconciliation is correct, then paleopolyploidization (whole genome duplication or WGD) associated with the origin of the genus Stenochlaena J. Sm. predated evolution of polocytic stomata, a heavily ornamented exospore, and a hemi-epiphytic to epiphytic habit observed in the crown lineage. Critically, this novel perspective elicits a merger of the K/Pg boundary fern spike and K/Pg boundary-WGD concepts, suggesting a link between polyploidy and the ‘disaster taxon’ concept.
白垩纪/古近纪(K/Pg)边界蕨类-孢子穗概念是由R.H. Tschudy在40年前首次提出的,并确立了蕨类是所谓的“灾难分类群”,由于其风吹孢子的高定植潜力,在自然灾害后繁盛。在当代K/Pg边界蕨类穗研究的关键问题中,最不为人所知的主题是,在K/Pg事件后繁盛的蕨类植物中,是否有类似窄尾蕨或其他茎白垩类蕨类植物,它们对蕨类孢子穗的laevigatosporites主导阶段做出了贡献。例如,分散的类stenochlaena孢子——如Polypodiisporites usmensis (van der Hammen) Khan & Martin——首次出现在南美洲始新世上部地层记录中,而类stenochlaena蕨类植物的巨型化石则首次出现在同一地区的古新世上部地层中。然而,除了对分散孢子产生者身份的地理和分类不确定性的传统关注之外,当代古植物学和分子系统发育观点在与K/Pg边界相关的毛囊蕨类茎系多样化时间方面存在进一步的不一致。本研究通过使用当代化石数据来限制分子钟发散时间估计,从而调和了这两种观点。如果这种和解是正确的,那么古多倍体化(全基因组复制或WGD)与Stenochlaena jsm属的起源有关。在冠系中观察到多细胞气孔,大量装饰的外孢子和半附生到附生习性的早期进化。重要的是,这种新观点引发了K/Pg边界蕨类穗和K/Pg边界- wgd概念的合并,表明多倍体和“灾难分类单元”概念之间存在联系。
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引用次数: 0
Age Constraints of the Guttulapollenites hannonicus –Cladaitina veteadensis Biozone in Argentina Reveal the First Record of Early Triassic (Olenekian) Palynofloras in Western Gondwana 阿根廷汉氏Guttulapollenites hannonicus–Cladaitina veteadensis生物带的年龄限制揭示了冈瓦纳大陆西部早三叠纪(Olenekian)Palynofloras的首次记录
IF 1.5 4区 地球科学 Q2 Earth and Planetary Sciences Pub Date : 2022-07-05 DOI: 10.1080/01916122.2022.2097329
S. N. Césari, C. Limarino, Patricia L. Ciccioli, C. Wall
ABSTRACT The upper Palaeozoic basins of central-western Argentina include continuous fossiliferous successions spanning the Carboniferous-Permian interval. The palynostratigraphic biozones comprise Late Mississippian, Pennsylvanian and Cisuralian assemblages. Recently, new palynofloras of the La Veteada Formation were referred to the Lopingian (late Permian). However, they are characterized by spores and tetrads of Lundbladispora spp. and Densoisporites spp., pollen grains of Lunatisporites pellucidus, L. noviaulensis and Protohaploxypinus samoilovichii, and the alga Syndesmorion stellatum, that distinguish the post-Permian recovery floras worldwide. A new uranium-lead chemical abrasion-isotope dilution-thermal ionization mass spectrometry (U-Pb CA-ID-TIMS) age confirms the Olenekian age of this stratigraphic unit and allows the identification of the first Early Triassic palynofloras in this region of western Gondwana. Comparison and correlation with similar assemblages from the southern and northern hemispheres supports the Early Triassic turnover with an increase of lycopsid cavate spores associated with some diagnostic species of taeniate and non-taeniate bisaccate pollen.
摘要阿根廷中西部的上古生代盆地包括横跨石炭-二叠纪的连续化石序列。坡缕岩生物带包括晚密西西比纪、宾夕法尼亚纪和西乌拉尔纪组合。近年来,拉Veteada组新的孢粉植物群被认为是洛平阶(晚二叠世)。然而,它们的特征是Lundbladispora spp.和Densoisporites spp.的孢子和四分体,Lunatisporites pellucidus、L.novaulensis和Protoploxypinus samoilovichii的花粉粒,以及藻类Syndesmorion stellatum,它们区分了世界各地的后二叠纪恢复区系。一个新的铀-铅化学磨蚀同位素稀释热电离质谱(U-Pb CA-ID-TIMS)年龄证实了该地层单元的奥列涅纪年龄,并允许识别冈瓦纳大陆西部该地区的第一个早三叠世孢粉植物群。与来自南半球和北半球的类似组合的比较和相关性支持了早三叠纪的更替,与一些带绦虫和非带绦虫双囊花粉的诊断物种相关的石松囊孢子增加。
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引用次数: 2
A palynological investigation of some taxa of the genus Ranunculus L. (Ranunculaceae) in Turkey and its taxonomic value 土耳其毛茛属部分分类群的孢粉学研究及其分类价值
IF 1.5 4区 地球科学 Q2 Earth and Planetary Sciences Pub Date : 2022-06-27 DOI: 10.1080/01916122.2022.2095053
M. Kurşat, Neriman Özdağ, B. Başer, FAHRETTİN ÖZBEY, I. Emre
Abstract Ranunculus L. (Ranunculaceae) is morphologically very diverse and palynological characters support the systematic studies of the genus. This study was aimed to perform palynological investigations of 30 different taxa of Ranunculus by using light microscope (LM) and scanning electron microscopy (SEM) methods. Polar-equatorial axis measurements, exine-intine layer thicknesses and colpi-pori numbers of pollen grains were examined using the Wodehouse method. Surface ornamentations were determined by electron microscope. The dendrogram of palynological characters analysed on the LM and SEM and observations made by multi-variate statistical package software partly support the monophyletic genera. According to the palynological dendrogram, Ranunculus taxa are divided into three types and R. arvensis clustered only in type 2. The results from present study suggest that the palynological characters can be useful for clarifying the taxonomic relationship in the genus.
摘要毛茛属植物形态多样,其孢粉学特征为系统研究提供了依据。本研究采用光学显微镜(LM)和扫描电子显微镜(SEM)对毛茛属30个不同类群进行了孢粉学研究。用Wodehouse方法测定了花粉粒的极赤道轴测量值、外壁-内层厚度和colpi pori数。表面纹饰用电子显微镜测定。LM和SEM上分析的孢粉学特征树状图以及多变量统计软件包的观察结果部分支持单系属。根据孢粉学树状图,毛茛属植物可分为三类,A.arvensis仅聚在第2类。本研究结果表明,该属的孢粉学特征有助于阐明该属的分类学关系。
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引用次数: 3
Palynomorphological and taxonomical investigations of some Apiaceae taxa from Bilecik, Turkey 土耳其Bilecik一些蜂科分类群的形态学和分类学研究
IF 1.5 4区 地球科学 Q2 Earth and Planetary Sciences Pub Date : 2022-06-24 DOI: 10.1080/01916122.2022.2094014
Aslı Özkök, Okan Sezer, O. Koyuncu, İ. Potoğlu Erkara
Abstract The objective of the present study is to investigate pollen morphology and taxonomy of some Apiaceae species in Bilecik, Turkey. The study area is restricted to Vezirhan in the south (Bilecik), Pamukova and Geyve in the north (Sakarya), İznik in the west (Bursa), Gölpazarı in the south-east (Bilecik) and Taraklı in the east (Sakarya). Apiaceae species were collected from the study area by visiting there in all four seasons. In this study, the pollen morphology of 22 taxa belonging to different genera of the Apiaceae family was investigated for the first time in detail using light microscope and scanning electron microscopy (SEM). The pollen is radially symmetrical, isopolar and tricolporate in all examined species. Pollen grains are perprolate, prolate and subprolate with the polar axis ranging from 17.53 µm (Non-acetolysed: N) and 18.28 µm (Acetolysed: A) to 39.71 µm (N) and 39.72 µm (A) and the equatorial axis from 11.77 µm (N) and 12.2 µm (A) to 23.77 µm (N) and 23.54 µm (A). The polar shapes of the studied taxa are triangular. Exine ornamentations of four pollen types have been observed by SEM. Two main groups were revealed in the dendrogram which was constructed by using Average Linkage from the examined data. It was determined that some pollen characteristics of investigated Apiaceae taxa are supporters of the classification of taxa with other taxonomical characteristics. Also, three components of investigated palynological characteristics were identified with principal component analysis. Polar axis (P), equatorial axis (E), the ratio of polar axis to equatorial axis (P/E) (pollen shape) and ornamentation in the polar and equatorial views are determined as the most important palynological characteristics for discrimination of the Apiaceae taxa.
摘要本研究的目的是研究土耳其比勒西克一些Apiaceae物种的花粉形态和分类学。研究区域仅限于南部的Vezirhan(Bilecik)、北部的Pamukova和Geyve(Sakarya)、西部的伊兹尼克(Bursa)、东南部的Gölpazarı(Bilezik)和东部的Taraklı(Sakaria)。Apiaceae物种是通过在所有四个季节的访问从研究区域收集的。本研究首次利用光学显微镜和扫描电子显微镜对Apiaceae科不同属的22个分类群的花粉形态进行了详细的研究。在所有被检查的物种中,花粉呈放射状对称、等极性和三角状。花粉粒为全长形、长形和亚板状,极轴在17.53之间 µm(未裂解:N)和18.28 µm(丙酮溶解:A)至39.71 µm(N)和39.72 µm(A)和赤道轴从11.77 µm(N)和12.2 µm(A)至23.77 µm(N)和23.54 µm(A)。研究分类群的极性形状是三角形的。用扫描电镜观察了四种花粉类型的外部纹饰,用平均连锁法构建的树状图显示了两个主要类群。已确定所调查的Apiaceae分类群的一些花粉特征是与其他分类群特征相结合的分类群分类的支持。同时,利用主成分分析法对所研究的孢粉学特征中的三个成分进行了鉴定。极轴(P)、赤道轴(E)、极轴与赤道轴之比(P/E)(花粉形状)和极赤道观中的纹饰被确定为区分Apiaceae分类群的最重要的孢粉学特征。
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引用次数: 0
Prestigious early Roman gardens across the Empire: the significance of gardens and horticultural trends evidenced by pollen 帝国早期著名的罗马花园:花粉证明了花园和园艺趋势的重要性
IF 1.5 4区 地球科学 Q2 Earth and Planetary Sciences Pub Date : 2022-06-15 DOI: 10.1080/01916122.2022.2089928
D. Langgut
Abstract The present study has two main goals. The first is to reconstruct the botanical components that grew in the impressive garden of Villa Arianna (Stabiae). The garden, which was extensively destroyed and covered by tephra ash in 79 CE, is considered the largest peristyle garden in the Vesuvian region. Its plants were revealed based on a unique palynological-archaeological method involving the extraction of pollen from plaster attached to structures that faced the garden. The second aim is to compare this prestigious garden with other early elite Roman gardens, located in the eastern part of the Empire, to trace the importation of plants, horticultural trends, etc. For this purpose, gardens of Herod the Great, the client king of Judaea, which the author recently studied palynologically (in Caesarea, Herodium and Jericho), were compared with the new pollen results of Villa Arianna. The comparison between the gardens’ botanical components and their different landscapes led to the following conclusions: (1) Plants were imported from both ends of the Empire as elite products (rather than cash crops). Hazelnut (Corylus) and cedar (Cedrus) were introduced from west to east, while the date palm (Phoenix dactylifera) was introduced along an east-west axis. (2) The gardening trend of tree dwarfism was observed both at Villa Arianna and Jericho. (3) The gardens flourished in challenging habitats. At Villa Arianna and Caesarea, efforts were devoted to sustaining splendid gardens in the relatively harsh, saline Mediterranean Sea environment; at Herodium and Jericho, special efforts were required for the success of Mediterranean plants in semi-arid climate. (4) Herod’s mausoleum garden in Herodium, whose dark evergreen trees on the whitish slopes of the artificial tumulus could be seen from the Temple in Jerusalem, may have been inspired by the architectural arrangement of the Pantheon and the Mausoleum of Augustus, the patron of Herod.
摘要本研究有两个主要目标。首先是重建在Villa Arianna(Stabiae)令人印象深刻的花园中生长的植物成分。该花园在公元79年被火山灰广泛摧毁并覆盖,被认为是维苏威地区最大的潜望式花园。它的植物是根据一种独特的孢粉考古方法发现的,该方法涉及从附着在花园结构上的灰泥中提取花粉。第二个目的是将这个著名的花园与位于帝国东部的其他早期罗马精英花园进行比较,以追踪植物的进口、园艺趋势等。为此,作者最近研究了犹太客户国王希律大帝的花园(在凯撒利亚、希律王和杰里科),与Villa Arianna的新花粉结果进行了比较。花园的植物成分及其不同景观之间的比较得出了以下结论:(1)植物是作为精英产品(而不是经济作物)从帝国两端进口的。榛子(Corylus)和雪松(Cedrus)是从西向东引入的,而椰枣(Phoenix dactylifera)是沿着东西轴线引入的。(2) 在Villa Arianna和Jericho都观察到了树木矮化的园艺趋势。(3) 花园在富有挑战性的栖息地蓬勃发展。在Villa Arianna和Caesarea,他们致力于在相对恶劣、盐碱的地中海环境中维持辉煌的花园;在希罗迪姆和杰里科,地中海植物在半干旱气候中的成功需要特别的努力。(4) 希律王在希律殿的陵墓花园,从耶路撒冷的圣殿可以看到人工陵墓白色斜坡上的深色常青树,其灵感可能来自万神殿和希律王的赞助人奥古斯都陵墓的建筑布置。
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引用次数: 4
Pollen Morphology of Lactuca L. (s. lat.) (Cichorieae: Asteraceae) from Hindukush, Western Himalayan and Karakorum Ranges and Its Taxonomic Significance 兴都库什山脉、西喜马拉雅山脉和喀喇昆仑山脉的乳苣苔(l.s lat.)(菊科:菊科)花粉形态及其分类意义
IF 1.5 4区 地球科学 Q2 Earth and Planetary Sciences Pub Date : 2022-06-15 DOI: 10.1080/01916122.2022.2089927
R. Abid, M. Qaiser
ABSTRACT Pollen morphology of 15 species of the genus Lactuca L. (s. lat.) of the tribe Cichorieae from the Hindukush, Western Himalayan and Karakorum Ranges was studied using light microscopy and scanning electron microscopy. The pollen morphology of six speciesm viz. L. erostrata Bano & Qaiser, L. glaucifolia Boiss., L. persica Boiss., L. crambifolia (Bunge) Boiss., L. chitralensis (Tuisl.) Ghafoor, Qaiser & Roohi Bano and L. picridiformis Boiss., was never studied before. Pollen were found to be isopolar, spheroidal to oblate-spheroidal, elliptic to rarely circular in equatorial view, hexagonal in polar view, trizonocolporate and echinolophate with 15 lacunae. Morphometric analysis of the pollen characters was conducted using agglomerative cluster analysis with Euclidean distance and Ward's method to compute the dissimilarity matrix, and a dendogram was prepared. Two pollen types, based primarily on the diameter of the polar and equatorial axes of the pollen grain, were recognized. Keys to the pollen types, subtypes and species are provided. The present palynological findings are to a great extent in agreement with the results of recent phylogenetic studies based on molecular evidence. Moreover, both the pollen types are well correlated with the general morphology of the investigated species of the genus Lactuca L. (s. lat) and provide additional micromorphological characters for the delimitation of the species.
摘要利用光学显微镜和扫描电子显微镜对印度库什山脉、喜马拉雅山脉西部和喀喇昆仑山脉菊科莴苣属15种植物的花粉形态进行了研究。对六个种的花粉形态进行了研究,它们分别是L.erostrata Bano&Qaiser,L.glaucifolia Boiss。,L.persica Boiss。,L.crambifolia(Bunge)Boiss。,L.chitralensis(Tuisl.)Ghafoor、Qaiser和Roohi Bano以及L.picridiformis Boiss。,以前从未学习过。花粉为等极性、球形至扁球形、赤道视为椭圆形至很少圆形、极视为六边形、三棱状和棘状,有15个陷窝。利用欧氏距离凝聚聚类分析和Ward方法计算相异矩阵,对花粉性状进行形态计量分析,并制备了树状图。主要根据花粉粒的极轴和赤道轴的直径,识别出两种花粉类型。提供了花粉类型、亚型和种类的关键。目前的孢粉学发现在很大程度上与最近基于分子证据的系统发育研究结果一致。此外,这两种花粉类型都与所研究的乳酸菌属(Lactuca L.(s.lat))物种的一般形态密切相关,并为该物种的划界提供了额外的微观形态特征。
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引用次数: 1
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Palynology
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