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Chemical processing of fossil phyllocarid cuticle: A comparison of micro- and macrofossil remains 化石层状和角质层的化学加工:微观和宏观化石遗迹的比较
4区 地球科学 Q3 PALEONTOLOGY Pub Date : 2023-10-17 DOI: 10.1080/01916122.2023.2267642
Vojtěch Kovář, Matěj Šilinger, Oldřich Fatka, Rainer Brocke
ABSTRACTCuticular remains of the phyllocarid crustaceans Caryocaris? and Ceratiocaris? obtained via maceration of macrofossil remains in hydrofluoric acid are compared with dispersed fragments isolated using both the low-manipulation HF extraction and the standard HCl–HF–HCl method. The studied specimens were obtained from Ordovician and Silurian shales of the Šárka and Požáry formations (Prague Basin, Barrandian area, Czech Republic). This methodological approach allows a direct comparison of macrofossil remains with microfossils in residue. The differences in residues obtained through the two bulk-rock extraction methods are also discussed.KEYWORDS: phosphatic microfossilsorganic-walled microfossilsextraction methodsearly PaleozoicPrague BasinDisclaimerAs a service to authors and researchers we are providing this version of an accepted manuscript (AM). Copyediting, typesetting, and review of the resulting proofs will be undertaken on this manuscript before final publication of the Version of Record (VoR). During production and pre-press, errors may be discovered which could affect the content, and all legal disclaimers that apply to the journal relate to these versions also. AcknowledgmentsThe authors would like to thank Ondřej Zicha (Prague) for providing specimens of Caryocaris and Saltericaris for the in situ study. We would further like to thank Monika Uhlířová and Zuzana Strossová (both Charles University Prague) for aid with collecting samples at Kosov quarry, Lena Kraus (Senckenberg Gesellschaft für Naturforschung) for assistance during laboratory processing of bulk rock samples from the Šárka Formation and Jana Čepičková (Charles University Prague) for providing literature, Lucy Muir for language editing, and finally to Vera Korasidis, Ben Slater and an anonymous referee for providing valuable feedback that significantly improved the manuscript.Disclosure statementNo potential conflict of interest was reported by the authors.FundingVK acknowledges support by the project “Grant Schemes at CU “(reg. no. CZ.02.2.69/0.0/0.0/19_073/0016935). OF acknowledges support by Cooperatio GEOL of the Ministry of Education, Youth and Sports of the Czech Republic.Figure 1 (A) Distribution of lower Paleozoic rocks within the Prague Basin and its location within the Czech Republic. (B) Stratigraphical subdivision of the Ordovician and Silurian of the Prague Basin with highlighted Šárka and Požáry formations (sampled in this study).Display full sizeFigure 2 Remains of Caryocaris? on a bedding plane (Ba, Ca, Da) and from microfossil residues (A, Bb, Cb, Db) placed upon a reconstruction. Scale bars represent 100 µm for all specimens except Cb. In Cb, the scale bar represents 50 µm.Display full sizePlate 1. Cuticular fragments of Caryocaris? sp. extracted (a) directly from macrofossil (2,7), (b) via ‘low-manipulation HF extraction’ process (4–6, 8–11, 13–18), (c) via HCl-HF-HCl processing (3) and a coniform conodont (12) from the locality Praha - Na Salátce q
根茎类和甲壳类动物的表皮残骸?和Ceratiocaris吗?通过氢氟酸浸渍获得的大型化石遗骸与使用低操作HF萃取和标准HCl-HF-HCl方法分离的分散碎片进行了比较。研究样品取自捷克巴兰地地区布拉格盆地Šárka组和Požáry组奥陶系和志留系页岩。这种方法可以直接比较大化石和微化石的残留物。讨论了两种大块岩石提取方法所得到的残余量的差异。关键词:磷微化石有机壁微化石提取方法早期古生代布拉格盆地免责声明作为对作者和研究人员的服务,我们提供此版本的接受稿件(AM)。在最终出版版本记录(VoR)之前,将对该手稿进行编辑、排版和审查。在制作和印前,可能会发现可能影响内容的错误,所有适用于期刊的法律免责声明也与这些版本有关。作者要感谢Ondřej Zicha (Prague)为现场研究提供的Caryocaris和Saltericaris标本。我们还要感谢Monika Uhlířová和Zuzana strossov(布拉格查尔斯大学)在科索沃采石场收集样本的帮助,Lena Kraus (Senckenberg Gesellschaft f r Naturforschung)在实验室处理Šárka组的大量岩石样本时的帮助,Jana Čepičková(布拉格查尔斯大学)提供文献,Lucy Muir进行语言编辑,最后感谢Vera Korasidis。本·斯莱特(Ben Slater)和一位匿名审稿人,感谢他们提供了宝贵的反馈,极大地改进了手稿。披露声明作者未报告潜在的利益冲突。中大资助计划获“中大资助计划”(注册)资助。不。CZ.02.2.69/0.0/0.0/19_073/0016935)。论坛感谢捷克共和国教育、青年和体育部GEOL合作组织的支持。图1 (A)布拉格盆地下古生代岩石分布及其在捷克境内的位置。(B)布拉格盆地奥陶系和志留系地层划分,突出显示Šárka和Požáry组(本研究取样)。图2石雕的遗迹?在层理平面(Ba, Ca, Da)和微化石残留物(a, Bb, Cb, Db)上进行重建。除Cb外,所有样品的比例尺均为100µm。在Cb中,比例尺代表50µm。显示全尺寸板核caris的表皮碎片?sp. (a)直接从巨化石(2,7)中提取,(b)通过“低操作HF提取”过程(4,6,8 - 11,13 - 18),(c)通过HCl-HF-HCl处理(3)和在当地Praha - Na Salátce采石场(中奥陶统Darriwilian Šárka组)中提取的锥形牙形石(12)。比例尺表示:50µm (1,3,4,6,8 - 10,12,14 - 16,18);200µm (2);100µm(5、7、11、17);20µm(13)。2、7号标本直接从大化石中提取,1、4 ~ 6、8 ~ 18号标本采用“低操作HF提取”方法提取,3号标本采用标准孢粉处理方法分离。幻灯片ŠS-JEN-29b, EF: W42(3)2。滑动ŠS-JEN-Caryocaris_a, EF: V51(1)幻灯片ŠS-JEN-30a, EF: E394。幻灯片ŠS-JEN-29b, EF: E36(2)5。滑动ŠS-JEN-29b, EF: P30(4)6。幻灯片ŠS-JEN-29b, EF: Q64(4)7。幻灯片ŠS-JEN-Caryocaris_a, EF: R458。幻灯片ŠS-JEN-29b, EF: E54(3)9。幻灯片ŠS-JEN-29c, EF: B6410。幻灯片ŠS-JEN-29b, EF: O59(3)11。幻灯片ŠS-JEN-29b, EF: C53(4)12。幻灯片ŠS-JEN-29b, EF: H46(3)13 -18。Stub saf_1显示全尺寸板2。角龙的遗迹?sp.(1 - 3,6,7),未知节肢动物(9)和牙形刺(4,5),来自Kosov采石场(Požáry组,Přídolí,晚志留世)。比例尺表示:400µm(1,2), 100µm(3), 50µm(4-7)。标本1-3直接从大化石中提取,4、5、8、9采用“低操作HF提取”工艺提取,6、7采用标准孢粉处理方法提取。滑块PS-K-FY-1, EF: W48(3)2。滑块PS-K-FY-1, EF: K44(1)3。滑块PS-K-FY-1, EF: W47(1)4。滑块PS-K-L-6c, EF: G29(4)5。滑块PS-K-L-6b, EF: G38(3)幻灯片PS-K-8c, EF: K43(3)幻灯片PS-K-8c, EF: G39(2)8。滑块PS-K-L-2, EF: F38(4)幻灯片PS-K-L-4c, EF: N37(2)显示全尺寸
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引用次数: 0
Outstanding botanical resources for Plebeia mansita (Apidae, Meliponini) in Northern Argentine Yungas: botanical characterization of its honey and main nesting substrates 阿根廷北部云加斯地区曼西塔小蜂的杰出植物资源:其蜂蜜和主要筑巢基质的植物学特征
4区 地球科学 Q3 PALEONTOLOGY Pub Date : 2023-10-05 DOI: 10.1080/01916122.2023.2266487
Fabio FernandoFlores, Liliana Concepción Lupo, Norma Inés Hilgert
AbstractPlebeia mansita is a native stingless bee endemic to the Yungas whose populations are distributed in sectors above 1000 m a.s.l. Its honeys and other products are used by local residents for various purposes, from food to medicinal uses. With the purpose of knowing the botanical resources that contribute nectar and nesting to this species of bee in the town of Baritú (22° 28’ 54.1” S; 64° 45’ 39.4” W; at 1546 m a.s.l.), it was analyzed 41 honey samples collected between September 2011 and September 2015 and recorded the main host trees. The samples it was examined with palynological techniques with acetolysis, observing the use of pollen types belonging to native plants, mainly of arboreal habit. Pollen richness per sample varied from 13 to 31 pollen types (mean = 22.7) in which where the family Asteraceae had the highest number of pollen types (n = 9) and nectar relevance, followed by the Myrtaceae family. Relevant pollen types correspond to Blepharocalyx salicifolius, Ilex argentina, Myrcianthes pseudomato and Allophylus edulis, followed by Sambucus peruviana, Parapiptadenia excelsa, Handroanthus lapacho, Myrsine coriacea and Zanthoxylum. Other recorded pollen types with importance index values less than 3.5% were Eupatorium, Dendrophorbium bomanii, Senecio, Pseudognaphalium and pollen of the families Asteraceae and Rhamnaceae. On the other hand, when evaluating nesting substrates it was recorded a frequency of use of trunks of Ilex argentina (55.9%), Parapiptadenia excelsa (10.3%), Myrcianthes pungens (7.4%), M. pseudomato (5.9%), Juglans australis (4.4%), Handroanthus lapacho (2.9%) and Blepharocalyx salicifolius, Myrsine coriacea, Ocotea porphyria, Viburnum seemenii, Myrcianthes mato, Roupala montana var. brasiliensis and Erithrina falcata with a value of 1.5%. each. The results obtained provide valuable information on the plants that provide nectar and nesting sites for this stingless bee. This information should be considered for the preparation of conservation or production plans for their hives and pollination activities.Keywords: Melissopalynologystingless beesmelliferous floranestingDisclaimerAs a service to authors and researchers we are providing this version of an accepted manuscript (AM). Copyediting, typesetting, and review of the resulting proofs will be undertaken on this manuscript before final publication of the Version of Record (VoR). During production and pre-press, errors may be discovered which could affect the content, and all legal disclaimers that apply to the journal relate to these versions also.
【摘要】曼西蜂(plebeia mansita)是云南特有的一种原生无刺蜜蜂,分布在海拔1000米以上的地区。其蜂蜜和其他产品被当地居民用于各种用途,从食品到药用。为了了解在Baritú镇(22°28 ' 54.1 ' S;64°45′39.4”w;于2011年9月至2015年9月期间采集了41份蜂蜜样本,并记录了主要寄主树。用孢粉学技术对样品进行了鉴定,观察到所用的花粉类型属于本地植物,主要是树栖植物。每个样品的花粉丰富度从13 ~ 31种花粉类型(平均为22.7种)不等,其中菊科的花粉类型最多(n = 9),与花蜜的相关性最高,其次是桃金桃科。相关花粉类型依次为Blepharocalyx salicifolius、Ilex argentina、Myrcianthes pseudomato和Allophylus edulis,其次为sambuus peruviana、Parapiptadenia excelsa、Handroanthus lapacho、myrsincoracea和Zanthoxylum。其他重要指数小于3.5%的花粉类型有泽兰、波马尼石斛(Dendrophorbium bomanii)、苍耳草(Senecio)、假格兰(Pseudognaphalium)以及菊科和鼠李科花粉。另方面,在筑巢基质评价中,阿根廷冬青(55.9%)、金丝桃(10.3%)、桃金娘(7.4%)、假瘤木(5.9%)、南胡桃木(4.4%)、山核桃(2.9%)、水杨花、桃金娘、紫花桃、巴西山青桃和鹰嘴藤(1.5%)的树干被利用频率最高。每一个。获得的结果为这种无刺蜜蜂提供花蜜和筑巢地点的植物提供了有价值的信息。这些信息应考虑到为其蜂房和授粉活动制定保护或生产计划。【关键词】蜜蜂科植物科免责声明作为对作者和研究人员的服务,我们提供此版本的已接受稿件(AM)。在最终出版版本记录(VoR)之前,将对该手稿进行编辑、排版和审查。在制作和印前,可能会发现可能影响内容的错误,所有适用于期刊的法律免责声明也与这些版本有关。
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引用次数: 0
New dinoflagellate cyst species from the Middle Eocene of the Volgograd-Volga region, southwest Russia 俄罗斯西南部伏尔加格勒-伏尔加河地区中始新世甲藻囊新种
4区 地球科学 Q3 PALEONTOLOGY Pub Date : 2023-10-05 DOI: 10.1080/01916122.2023.2266491
Olga N. Vasilyeva
AbstractThe Keresta Formation, located in the Volgograd-Volga region (the Volga-Caspian subregion of southwest Russia) has been dated as a Middle Eocene, Lutetian based on dinoflagellate cysts as the Enneadocysta arcuata Zone and nannofossils from the Lutetian Subzones CP13c-CP14a. Within this interval, four new species of dinoflagellate cysts were described: Spiniferella grigorovichiae sp. nov., Impagidinium tuberculatum sp. nov., Hystrichostrogylon crassitunicatum sp. nov. and Hystrichosphaeropsis tenerum sp. nov. The new species present a high correlation potential for Middle Eocene stratigraphy over the southern part of the Russian Platform. Also, the assemblage of organic-walled microphytoplankton from the Keresta Formation indicates warm water marine environments within the marginal part of the Paratethys Basin.Keywords: Dinoflagellate cystsMiddle EoceneLutetiansouthwest RussiaVolgograd-Volga regionDisclaimerAs a service to authors and researchers we are providing this version of an accepted manuscript (AM). Copyediting, typesetting, and review of the resulting proofs will be undertaken on this manuscript before final publication of the Version of Record (VoR). During production and pre-press, errors may be discovered which could affect the content, and all legal disclaimers that apply to the journal relate to these versions also.
摘要俄罗斯西南部伏尔加格勒-伏尔加地区(伏尔加-里海亚区)的Keresta组为中始新世,Lutetian为中始新世,基于鞭毛藻囊(Enneadocysta arcuata带)和Lutetian亚带CP13c-CP14a的纳米化石。在此期间,还发现了4个鞭毛藻囊新种:Spiniferella grigorovichiae sp. nov.、Impagidinium tuberculatum sp. nov.、Hystrichostrogylon crassitunicatum sp. nov.和Hystrichosphaeropsis tenerum sp. nov.,这些新种对俄罗斯地台南部中始新世地层具有较高的对比潜力。此外,Keresta组有机壁微型浮游植物的组合表明,Paratethys盆地边缘地区存在温暖的海洋环境。关键词:鞭毛藻囊肿中期初新世俄罗斯西南部伏尔加格勒-伏尔加河地区免责声明作为对作者和研究人员的服务,我们提供此版本的已接受稿件(AM)。在最终出版版本记录(VoR)之前,将对该手稿进行编辑、排版和审查。在制作和印前,可能会发现可能影响内容的错误,所有适用于期刊的法律免责声明也与这些版本有关。
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引用次数: 0
Pollen Morphology of Endemic Linum Species (Linoideae: Linaceae) from Mexico 墨西哥特有亚麻属植物(亚麻科)的花粉形态学
IF 1.5 4区 地球科学 Q3 PALEONTOLOGY Pub Date : 2023-10-02 DOI: 10.1080/01916122.2023.2222293
Juan González-Velasco, Iris G. Galván-Escobedo, Mireya Burgos-Hernández, G. Castillo‐Campos
ABSTRACT The morphological characteristics of pollen grains of Linum species in Mexico have not previously been evaluated in quantitative terms. This study reports statistical values of pollen characteristics – polar axis, equatorial axis, shape, polar area, exine thickness, sexine thickness, nexine thickness, and height and diameter of ornamental pollen elements – of 11 of the 13 endemic species accepted to date and for which material was available: L. cruciata, L. flagellare, L. lasiocarpum, L. longipes, L. mexicanum, L. modestum, L. orizabae, L. pringlei, L. rzedowskii, L. scabrellum, and L. tenellum. The detailed light and scanning electron microscopy study revealed that the pollen is prolate-spheroidal to subprolate, tricolpate and with semitectate exine sculptured with bacula, clavae, gemmae or spines whose height and diameter follow a homogeneous, intermediate, or heterogeneous pattern. The principal component analysis found that five characters, namely the height of the ornamental elements, sexine thickness, exine thickness, diameter of ornamental elements, and nexine thickness, explained the highest percentage of the variance between species. The cluster and discriminant analyses classified some of the species into three groups and differentiated three of them according to their morphology. The difficulty in distinguishing some species through a conventional approach was solved conveniently using palynological traits since the ornamentation pattern of pollen grains was unique and because grains of various species were statistically different, thus representing a tool for identifying Linum species.
摘要 以前从未对墨西哥亚麻属植物花粉粒的形态特征进行过定量评估。本研究报告了迄今为止已接受的 13 个地方物种中 11 个物种的花粉特征统计值,包括极轴、赤道轴、形状、极区、外轮廓厚度、性轴厚度、内轮廓厚度、观赏花粉元素的高度和直径:L. cruciata、L. flagellare、L. lasiocarpum、L. longipes、L. mexicanum、L. modestum、L. orizabae、L. pringlei、L. rzedowskii、L. scabrellum 和 L. tenellum。详细的光镜和扫描电子显微镜研究表明,花粉呈球状至近球状、三柱状,具有半ectate外皮,外皮上雕刻有bacula、clavae、gemae或刺,其高度和直径遵循同质、中间或异质模式。主成分分析发现,装饰要素的高度、sexine 厚度、exine 厚度、装饰要素的直径和 nexine 厚度这五个特征解释了物种间差异的最大百分比。聚类分析和判别分析将部分物种分为三组,并根据形态将其中三个物种区分开来。由于花粉粒的装饰图案是独一无二的,而且不同物种的花粉粒在统计学上存在差异,因此利用古植物学特征可以很方便地解决传统方法难以区分某些物种的问题,从而成为鉴别亚麻属植物物种的工具。
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引用次数: 0
Pollen morphology of Ipomoea L. species from Garhwal Himalaya, India and its taxonomic implication 印度Garhwal - himalayan地区Ipomoea L.种花粉形态及其分类意义
4区 地球科学 Q3 PALEONTOLOGY Pub Date : 2023-09-28 DOI: 10.1080/01916122.2023.2264357
Prabhawati Tiwari, Ruchita Rawat, Shivani Negi, Dinesh Singh Rawat
AbstractPollen morphology serves as a valuable tool for identification, phylogenetic analysis and evolutionary studies. The present study aims to investigate the pollen morphology of 12 species of the genus Ipomoea L. (a diverse and economically important genus within the family Convolvulaceae) from Garhwal Himalaya, India. Fresh flower buds were collected from the study area and pollen preparation was done using the acetolysis method. The morphology of the pollen grains was examined using light microscopy (LM) and scanning electron microscopy (SEM). Pollen grains of all the studied species were apolar, spheroidal, pantoporate, subtectate with fine reticulation and echinate but they differed from one another in their size, number of spines and pores, spine length and spine types. The size of grains ranged from 53.3 μm to 128.7 μm. Cluster analysis was performed based on pollen characters to show the relationship among the species and a key was prepared and discussed. The observed variation in pollen morphological traits gives important taxonomic information that is useful for distinguishing Ipomoea species and establishing taxonomic and phylogenetic relationships.Keywords: IpomoeaPantoporateSpineSpheroidalCluster analysisDisclaimerAs a service to authors and researchers we are providing this version of an accepted manuscript (AM). Copyediting, typesetting, and review of the resulting proofs will be undertaken on this manuscript before final publication of the Version of Record (VoR). During production and pre-press, errors may be discovered which could affect the content, and all legal disclaimers that apply to the journal relate to these versions also.
摘要花粉形态是花粉鉴定、系统发育分析和进化研究的重要工具。本文研究了印度Garhwal喜玛拉雅地区12种旋花属植物(Ipomoea L.)的花粉形态。从研究区采集新鲜花蕾,用乙酰解法制备花粉。用光镜和扫描电镜观察了花粉粒的形态。所有种类的花粉粒均为极粒状、球形、泛孔状、细网状的隐盖状和棘刺状,但在花粉粒大小、棘孔数、棘长和棘型等方面存在差异。晶粒尺寸为53.3 ~ 128.7 μm。根据花粉性状进行聚类分析,揭示种间的亲缘关系,并编制和讨论了一个关键字。所观察到的花粉形态特征的变化提供了重要的分类学信息,有助于区分Ipomoea种,建立分类学和系统发育关系。关键词:ipomoeapantoporatespinespheroidal聚类分析免责声明作为对作者和研究人员的服务,我们提供此版本的已接受稿件(AM)。在最终出版版本记录(VoR)之前,将对该手稿进行编辑、排版和审查。在制作和印前,可能会发现可能影响内容的错误,所有适用于期刊的法律免责声明也与这些版本有关。
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引用次数: 0
Palynotaxonomy of species of Dendrophorbium (Asteraceae: Senecioneae) occurring in Brazil 产自巴西的石斛属植物的孢粉分类
4区 地球科学 Q3 PALEONTOLOGY Pub Date : 2023-09-28 DOI: 10.1080/01916122.2023.2262530
Thayná Rosa Batista Martins, Raquel Maria Batista Souza-Souza, Wellerson Picanço Leite, Roberto Lourenço Esteves, Cláudia Barbieri Ferreira Mendonça, Vania Gonçalves-Esteves
AbstractThis article presents a pollen study of 13 species of Dendrophorbium (Asteraceae: Senecioneae) occurring in Brazil. Dendrophorbium has species distributed in the South and Southeast regions of the Brazilian territory that were previously grouped in Senecio Myriocephalus sect. and later Dendrophorbium was elevated to the rank of genus, considering the section Myriocephalus as a synonym of Dendrophorbium. The aim of this study is to advance knowledge of the pollen characteristics of the genus and thus contribute to the taxonomy of the group. The botanical material was collected from specimens deposited in national herbaria. Floral buds were acetolysed for light microscopic analysis. Pollen sediments were mounted on slides with glycerinated gelatin and analyzed under a light microscope. Measurements were taken of the diameters and thickness of exine layers of pollen grains. Examination showed pollen grains to be monadic, isopolar, medium-sized, oblate-spheroidal to prolate-spheroidal, and 3-colporate. The endoaperture has median constriction in most species, except in D. pellucidinerve. The sexine is thicker than the nexine and echinate, with large perforations at the base of echinae. The exine is caveate. Based on the results, it can be concluded that the pollen grains of the 13 Dendrophorbium species are homogeneous and can be differentiated by aperture shape and dimensions and echinus characteristics.Keywords: CompositaemorphologyPalynologytaxonomyDisclaimerAs a service to authors and researchers we are providing this version of an accepted manuscript (AM). Copyediting, typesetting, and review of the resulting proofs will be undertaken on this manuscript before final publication of the Version of Record (VoR). During production and pre-press, errors may be discovered which could affect the content, and all legal disclaimers that apply to the journal relate to these versions also. Declaration of Competing InterestThe authors declare that they have no known competing financial interests or personal relationships that could have appeared to influence the work reported in this paper.The authors declare that have no conflict of interest.Data availabilityAll data supporting the findings of this study are available from the corresponding author.AcknowledgementsV. Esteves and C. B. F. Mendonça are grateful to the Brazilian National Council for Scientific and Technological Development (CNPq) for the research grant (nos. 304910/2019-8, 31618/2021-9). All authors thank the Rio de Janeiro State Research Foundation (FAPERJ) (grant nos. 26.0003/015254/2021 and 260003/015254/2021). We are grateful to the Brazilian Federal Agency for Support and Evaluation of Graduate Education (CAPES) and the Post-Graduate Support Program (PROAP) for awarding scholarships. We also thank the herbarium curators for providing access to their collections and the Scanning Microscopy Laboratory, National Museum, Federal University of Rio de Janeiro, Brazil, for conducting the
摘要本文报道了巴西13种石斛属植物(菊科:石斛科)的花粉研究。Dendrophorbium有分布在巴西南部和东南部地区的种,以前被归为Senecio Myriocephalus组,后来将Dendrophorbium提升到属的级别,将Myriocephalus组视为Dendrophorbium的同义词。本研究的目的是提高对该属花粉特征的认识,从而有助于该类群的分类。植物材料来源于国家植物标本馆保存的标本。花蕾被乙酰化用于光镜分析。将花粉沉积物装在甘油明胶载玻片上,在光学显微镜下进行分析。测定了花粉粒的外皮直径和厚度。花粉粒为单形、等极性、中等大小、扁球状至长球状和3合生。大多数种的内孔有中间缩窄,除了在D. pellucidinerve。阴部比内阴部和棘刺粗,在棘刺基部有大的穿孔。外壁呈空泡状。结果表明,13种石斛属植物的花粉粒具有较强的同质性,可通过花粉孔的形状、大小和棘突特征进行区分。关键词:合成词形词类分类免责声明作为对作者和研究人员的服务,我们提供此版本的已接受稿件(AM)。在最终出版版本记录(VoR)之前,将对该手稿进行编辑、排版和审查。在制作和印前,可能会发现可能影响内容的错误,所有适用于期刊的法律免责声明也与这些版本有关。竞争利益声明作者声明,他们没有已知的竞争经济利益或个人关系,可能会影响本文所报道的工作。作者声明没有利益冲突。数据可得性支持本研究结果的所有数据均可从相应作者处获得。Esteves和C. B. F. mendonapera感谢巴西国家科学技术发展委员会(CNPq)的研究资助(no . 304910/2019- 8,31618 /2021-9)。所有作者感谢里约热内卢州研究基金会(FAPERJ)(批准号26.0003/015254/2021和260003/015254/2021)。我们感谢巴西联邦研究生教育支持和评估机构(CAPES)和研究生支持计划(PROAP)颁发奖学金。我们也感谢植物标本馆馆长提供他们的藏品,感谢巴西里约热内卢联邦大学国家博物馆扫描显微镜实验室进行分析。
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引用次数: 0
Contribution to pollen morphology of Astragalus L. section Aegacantha Bunge (Galegeae, Fabaceae) and its systematic significance 黄芪(gaacantha Bunge)剖面花粉形态的贡献及其系统意义
4区 地球科学 Q3 PALEONTOLOGY Pub Date : 2023-09-28 DOI: 10.1080/01916122.2023.2264352
Amjad Khan, Mushtaq Ahmad, Amir Sultan, Zakir Ullah, Khawar Majeed, Tariq Ullah, Muhammad Zafar
ABSTRACTAstragalus L. sect. Aegacantha Bunge has 57 described species mostly confined in Southwest Asia and Central Asia. Here, we studied the pollen morphology of 17 species from this section present in Pakistan. Pollen morphology of sect. Aegacantha species is poorly investigated despite its systematic importance. The main aim of this study is to determine and document the pollen characters that can help in the taxonomic identification of species in this difficult section. Pollen slides were prepared following acetolysis and examined by both light and scanning electron microscopy. Pollen size varies, with the polar axis ranging from 32.91-43.10 μm and the equatorial axis from 22.91-28.47 μm. The shape of pollen is sub-prolate (1.15-1.33) to prolate (1.34-2.00), small to medium in size, radially symmetrical, isopolar, trizonocolporate to tricolporate and monads. Sculpturing variations (i.e. reticulate, microreticulate, perforate, microperforate, regulate, microregulate and granulate) were observed around the apertural, meridional and polar regions. Ordination (PCA) analysis revealed that the quantitative characters of pollen morphology are the most reliable characters for delimitation of section Aegacantha species. Cluster (UPGMA) analysis shows a partial relationship between the Aegacantha taxa clustered based on pollen characters and supports the general morphology. Further, this study shows that the pollen traits provide a baseline for phylogenetic optimization in the investigated Aegacantha section.KEYWORDS: TaxonomypalynologyphylogenySEM-analysisPCAUPGMAPakistanDisclaimerAs a service to authors and researchers we are providing this version of an accepted manuscript (AM). Copyediting, typesetting, and review of the resulting proofs will be undertaken on this manuscript before final publication of the Version of Record (VoR). During production and pre-press, errors may be discovered which could affect the content, and all legal disclaimers that apply to the journal relate to these versions also.
摘要/ abstract摘要:加棘属(Aegacantha Bunge)有57种,主要分布在西南亚和中亚。本文对分布于巴基斯坦的17种植物的花粉形态进行了研究。尽管其在系统上具有重要意义,但其花粉形态的研究却很少。本研究的主要目的是确定和记录花粉特征,以帮助在这一困难的剖面中进行物种分类鉴定。乙酰解后制备花粉载玻片,用光镜和扫描电镜观察。花粉的极轴在32.91 ~ 43.10 μm之间,赤道轴在22.91 ~ 28.47 μm之间。花粉的形状为近长形(1.15-1.33)至长形(1.34-2.00),大小小至中等,径向对称,等极性,三聚体至三聚体和单胞体。在孔洞区、子午区和极区周围观察到雕刻变化(即网状、微网状、穿孔、微穿孔、调节、微调节和颗粒状)。排序(PCA)分析结果表明,花粉形态的数量特征是最可靠的物种划分特征。聚类分析(UPGMA)表明,依花粉特征聚类的Aegacantha类群具有一定的相关性,支持其一般形态。此外,本研究表明,花粉性状为所调查的Aegacantha区段的系统发育优化提供了基线。作为对作者和研究人员的服务,我们提供此版本的已接受稿件(AM)。在最终出版版本记录(VoR)之前,将对该手稿进行编辑、排版和审查。在制作和印前,可能会发现可能影响内容的错误,所有适用于期刊的法律免责声明也与这些版本有关。
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引用次数: 0
Palynotaxonomy of the subtribe Anisopappinae (Athroismeae, Asteraceae) and the genus Duhaldea (Inuleae, Asteraceae) based on their exine ultrastructure 基于外叶超微结构的山葵亚族(蕨目,菊科)和杜鹃属(茵叶目,菊科)孢粉分类
4区 地球科学 Q3 PALEONTOLOGY Pub Date : 2023-09-27 DOI: 10.1080/01916122.2023.2261997
António PereiraCoutinho, Santiago Ortiz, Daniela Cordeiro, Miguel Serrano, Mónica Zuzarte
AbstractThe exine ultrastructure of eighteen species of Anisopappus (Anisopappinae, Athroismeae, Asteraceae) and three species of Duhaldea (Inuleae, Asteraceae) was studied using Transmission Electron Microscopy (TEM). Two ultrastructural patterns were identified: Senecioid and Helianthoid. The Senecioid pattern is characteristic of the species included in the basal clades of the phylogenetic tree of Anisopappus, as published in earlier studies. The Helianthoid pattern is restricted to the taxa placed in the derived clades, including the core group of species of Anisopappus where A. chinensis, the type species, is included. The observed ultrastructural differences seem to suggest that a future splitting of Anisopappus into several different genera should not be disregarded and that the Helianthoid pattern represents, within this genus, the derived condition. The ultrastructure of the exine supports the inclusion, suggested by previous authors, of the species Duhaldea stuhlmannii, but not that of Philyrophyllum schinzii, in Anisopappus. In contrast, the three studied species of Duhaldea (D. cappa, D. nervosa, D. revoluta) present an exine with a Senecioid pattern, which is characteristic of the tribe Inuleae, in which they are included. The ectoaperture, mesoaperture, and endoaperture of all studied Anisopappinae and of Duhaldea, respectively intersect the tectum plus columellae, the foot layer and the outer part of the endexine, and the inner part of the endexine. This feature seems to characterize, in a consistent way, the “Heliantheae alliance”, and is also present in all the studied Asteroideae (excl. Anthemideae), Dicomoideae, and Carduoideae.Keywords: AnisopappusAnisopappinaeDuhaldeaexinemesoaperturespalynotaxonomyultrastructureDisclaimerAs a service to authors and researchers we are providing this version of an accepted manuscript (AM). Copyediting, typesetting, and review of the resulting proofs will be undertaken on this manuscript before final publication of the Version of Record (VoR). During production and pre-press, errors may be discovered which could affect the content, and all legal disclaimers that apply to the journal relate to these versions also.
摘要利用透射电子显微镜(TEM)研究了18种异角蕨科(异角蕨科,刺尾蕨科)和3种杜鹃科(刺尾蕨科,菊科)的外壁超微结构。发现两种超微结构模式:Senecioid和Helianthoid。在早期的研究中发表的研究结果表明,Senecioid模式是异角猿系统发育树的基础分支中包含的物种的特征。Helianthoid模式仅限于被置于衍生枝中的分类群,包括以模式种中国古猿(A. chinensis)为代表的异距鸟(Anisopappus)核心类群。观察到的超微结构差异似乎表明,不应忽视未来异角猿分裂为几个不同属的可能性,而在这个属中,Helianthoid模式代表了衍生的条件。外壁的超微结构支持了先前作者提出的物种Duhaldea stuhlmannii的包含,而不是Philyrophyllum schinzii的包含。相比之下,所研究的Duhaldea (D. cappa, D. nervosa, D. revoluta)的三个种呈现出具有Senecioid模式的外壁,这是Inuleae部落的特征,它们被包括在其中。所研究的异松科和杜哈尔亚科的外孔、中孔和内孔分别相交于顶盖加小柱、足部与叶尖外侧和叶尖内侧。这一特征似乎是“Heliantheae联盟”的一贯特征,也存在于所有研究的Asteroideae(不包括Anthemideae)、Dicomoideae和Carduoideae中。关键词:异角蝶,异角蝶,duhaldeexineme, esoaperture, respalyno, taxonomy,超微结构免责声明作为对作者和研究人员的服务,我们提供这个版本的接受稿件(AM)。在最终出版版本记录(VoR)之前,将对该手稿进行编辑、排版和审查。在制作和印前,可能会发现可能影响内容的错误,所有适用于期刊的法律免责声明也与这些版本有关。
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引用次数: 0
Pollen morphology of Matthiola and the related genus Dvorakia (Brassicaceae) in Iran 标题伊朗芸苔科Matthiola及其相关属Dvorakia的花粉形态
4区 地球科学 Q3 PALEONTOLOGY Pub Date : 2023-09-23 DOI: 10.1080/01916122.2023.2260457
Somayeh Karami, Massoud Ranjbar
ABSTRACTOur study focused on the pollen morphology of 13 Matthiola species and the closely related genus Dvorakia. We used light and scanning electron microscopes to examine the importance of pollen characteristics in classifying these genera. Pollen grains were acetolysed, measured, photographed, and described based on qualitative and quantitative characteristics. Here, we present statistical analyses and multivariate statistics for quantitative data. Pollen in both genera is tricolpate and varies in shape from subprolate to prolate. We distinguish two pollen types based on lumen width. Matthiola pollen has both reticulate and macroreticulate ornamentation, while Dvorakia alyssifolia pollen is reticulate. The thickness of the exine and the width of the mesocolpium are important traits in addition to exine ornamentation. Iranian Dvorakia and Matthiola species' pollen size can be divided into four types, one of which is Type I (Dvorakia alyssifolia-type). Among the examined species, M. tomentosa has the smallest pollen grains, while D. alyssifolia has the biggest grains. Pollen morphology confirms the transfer of D. alyssifolia from Matthiola to Dvorakia. Additionally, we propose a species key for the genera.KEYWORDS: palynologyscanning electron microscopic studytaxonomytricolpateDisclaimerAs a service to authors and researchers we are providing this version of an accepted manuscript (AM). Copyediting, typesetting, and review of the resulting proofs will be undertaken on this manuscript before final publication of the Version of Record (VoR). During production and pre-press, errors may be discovered which could affect the content, and all legal disclaimers that apply to the journal relate to these versions also.
摘要本研究对13种Matthiola属及其近缘属Dvorakia的花粉形态进行了研究。利用光镜和扫描电镜研究了花粉特征对这些属的分类的重要性。对花粉粒进行乙酰化、测量、拍照,并根据定性和定量特征进行描述。在这里,我们对定量数据进行统计分析和多元统计。两属的花粉均为三柄,形状从近长形到长形不等。我们根据管腔宽度来区分两种花粉类型。黄松花粉既有网状纹饰,也有大网状纹饰;黄松花粉为网状纹饰。外壁的厚度和中果皮的宽度是除外壁纹饰外的重要特征。伊朗Dvorakia和Matthiola的花粉大小可分为四种类型,其中一种为I型(Dvorakia alyssifolia-type)。在所调查的物种中,毛毛蓟的花粉粒最小,而白花蓟的花粉粒最大。花粉形态证实了D. alyssifolia从Matthiola转移到Dvorakia。此外,我们提出了一个属的种键。关键词:孢粉;扫描电镜研究;分类;免责声明作为对作者和研究人员的服务,我们提供此版本的已接受稿件(AM)。在最终出版版本记录(VoR)之前,将对该手稿进行编辑、排版和审查。在制作和印前,可能会发现可能影响内容的错误,所有适用于期刊的法律免责声明也与这些版本有关。
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引用次数: 0
‘The man who vacuum cleaned the Atlantic’ – the aerosol collector and Gunnar Erdtman’s attempts to measure pollen rain “用真空吸尘器清扫大西洋的人”——气溶胶收集器和加纳·厄德特曼测量花粉雨的尝试
4区 地球科学 Q3 PALEONTOLOGY Pub Date : 2023-09-19 DOI: 10.1080/01916122.2023.2260437
Kevin J. Edwards, Pia Östensson
ABSTRACTIn 1937, the Swedish palynologist Gunnar Erdtman (1897–1973) mounted two adapted vacuum cleaners atop an ocean-going liner and set out to obtain air samples as he crossed the Atlantic Ocean. The devices were able to capture samples of airborne pollen which were related to air volume and distance from land. The results of this investigation are still cited in the scientific literature, but a study of some overlooked publications demonstrates that the ocean study had land-based antecedents. Furthermore, archival investigation reveals the background to such studies, including the technical plans for Erdtman’s ‘aerosol collector’, records of raw data from the voyage, draft portions of the key publication, photographs and the precise location of the land-based equipment. A storage loft in the Swedish Museum of Natural History in Stockholm has been found to house the surviving aerosol collector.KEYWORDS: Gunnar Erdtmanarchiveslong distance pollen rainNorth Atlantic Oceanvacuum cleaneraerosol collectorhistory of scienceDisclaimerAs a service to authors and researchers we are providing this version of an accepted manuscript (AM). Copyediting, typesetting, and review of the resulting proofs will be undertaken on this manuscript before final publication of the Version of Record (VoR). During production and pre-press, errors may be discovered which could affect the content, and all legal disclaimers that apply to the journal relate to these versions also. Additional informationNotes on contributorsKevin J. Edwards KEVIN J. EDWARDS holds degrees from the Universities of St Andrews (MA, DSc) and Aberdeen (PhD). He is Emeritus Professor in physical geography and adjunct chair in archaeology in the University of Aberdeen, adjunct professor in the Graduate Center, City University of New York, and, at the University of Cambridge, Fellow Commoner of Clare Hall, Senior Fellow of the McDonald Institute for Archaeological Research and Emeritus Associate of the Scott Polar Research Institute. He has held academic posts and visiting research positions in a number of universities including those of Queen’s Belfast, Birmingham, Sheffield, Minnesota, Copenhagen and Oxford. A palynologist with a strong involvement in multidisciplinary Quaternary science, his interests include human-environment interactions during Lateglacial and Holocene times in Britain and Ireland, Norse impacts upon the landscapes of the North Atlantic region, methodology, and the history of science.Pia Östensson PIA ÖSTENSSON is curator of the Palynological Laboratory in the Department of Environmental Research and Monitoring of the Naturhistoriska riksmuseet (Swedish Museum of Natural History) in Stockholm. A graduate in biology (plant systematics) from Stockholm University, she administers the pollen monitoring network for central Sweden and has extensive experience of aerobiology and forecasting based on airborne pollen sampling, plant phenology, and long distance palynomorph transport. Her
1937年,瑞典孢粉学家Gunnar Erdtman(1897-1973)在一艘远洋客轮上安装了两台经过改装的真空吸尘器,开始在横渡大西洋时采集空气样本。该装置能够捕获空气中花粉的样本,这些样本与空气量和与陆地的距离有关。这项调查的结果仍然在科学文献中被引用,但对一些被忽视的出版物的研究表明,海洋研究有陆地研究的先例。此外,档案调查揭示了这些研究的背景,包括厄德特曼“气溶胶收集器”的技术计划、航行原始数据的记录、关键出版物的草稿部分、照片和陆基设备的精确位置。人们在斯德哥尔摩的瑞典自然历史博物馆找到了一个储藏阁楼,用来存放现存的气溶胶收集器。关键词:Gunnar erdtmanarchives远距离花粉雨北大西洋海洋真空吸尘器气溶胶收集器科学史免责声明作为对作者和研究人员的服务,我们提供此版本的接受稿件(AM)。在最终出版版本记录(VoR)之前,将对该手稿进行编辑、排版和审查。在制作和印前,可能会发现可能影响内容的错误,所有适用于期刊的法律免责声明也与这些版本有关。KEVIN J. Edwards拥有圣安德鲁斯大学(MA, DSc)和阿伯丁大学(博士)学位。他是阿伯丁大学自然地理学名誉教授和考古学兼职主席,纽约城市大学研究生中心兼职教授,剑桥大学克莱尔·霍尔研究员,麦克唐纳考古研究所高级研究员和斯科特极地研究所名誉副研究员。他曾在贝尔法斯特女王大学、伯明翰大学、谢菲尔德大学、明尼苏达大学、哥本哈根大学和牛津大学等多所大学担任学术职位和访问研究职位。作为一名孢粉学家,他对多学科的第四纪科学有着浓厚的兴趣,他的兴趣包括英国和爱尔兰冰川和全新世时期人类与环境的相互作用,北欧人对北大西洋地区景观的影响,方法论和科学史。Pia Östensson Pia ÖSTENSSON是斯德哥尔摩自然历史博物馆环境研究和监测部门孢粉实验室的馆长。她毕业于斯德哥尔摩大学生物学(植物系统学)专业,负责管理瑞典中部的花粉监测网络,并在空气生物学和基于空气花粉采样、植物物候学和长距离花粉形态运输的预测方面拥有丰富的经验。她之前的职位包括在斯德哥尔摩大学教授植物学,科学顾问,以及在瑞典国立自然历史博物馆植物观察系林奈植物标本馆担任高级助理。她是植物学家,也是《激情系统:linn och drömmen》一书的插画家。
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Palynology
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