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Étude d'un assemblage original de microvertébrés du Pléistocène moyen du nord-est de l'Algérie (Ben Kérat, Oued Zenati) et description de deux nouveaux muridés 阿尔及利亚东北部中更新世微脊椎动物原始组合的研究(Ben kerat, Oued Zenati)和两个新的murids的描述
IF 1.8 3区 地球科学 Q2 PALEONTOLOGY Pub Date : 2022-02-17 DOI: 10.5252/geodiversitas2022v44a8
Emmanuelle Stoetzel, M. Pickford
RÉSUMÉ La présente étude porte sur l'analyse d'un assemblage inédit de restes de microvertébrés récoltés en 1910 à Ben Kérat, dans le nord-est de l'Algérie. L'objectif principal de cette étude était d'évaluer le potentiel paléontologique du site, et de préciser son contexte géologique et chronologique. L'assemblage étudié obtenu à partir de plusieurs échantillons de brèche comprend 362 restes identifiés, appartenant à 44 individus et 11 taxons: six rongeurs appartenant aux genres Gerbillus Desmarest, 1804, Meriones Illiger, 1811, Paraethomys Petter, 1968, Mus Linnaeus, 1758, Praomys Thomas, 1915 et Ellobius Fischer, 1814, deux soricidés du genre Crocidura Wagler, 1832, un amphibien anoure, un lézard lacertidé et un serpent colubridé, indiquant un âge situé vers la fin du Pléistocène moyen. Bien que le spectre faunique global soit similaire à d'autres assemblages de sites maghrébins du Pléistocène moyen, les spécimens de Ben Kérat présentent des caractéristiques dentaires particulières, pouvant traduire soit une période d'amélioration climatique dans un contexte plus global d'aridification et de refroidissement croissant, soit une particularité régionale avec notamment des nouvelles formes de Paraethomys et de Mus ne s'insérant pas dans les lignées évolutives connues jusqu'ici en Afrique du Nord.
本研究分析了1910年在阿尔及利亚东北部本凯拉特采集的一组新的微型脊椎动物遗骸。本研究的主要目的是评估该遗址的古生物学潜力,并澄清其地质和时间背景。从多个缺口样本中获得的研究组合包括362个已鉴定的遗骸,属于44个个体和11个分类群:6个啮齿动物属,Gerbillus Desmarest,1804年,Meriones Illiger,1811年,Paraethomys Petter,1968年,Mus Linnaeus,1758年,Praomys Thomas,1915年和Ellobius Fischer,1814年,Crocidura Wagler属的两个杀山梨科,1832年,一个无尾两栖动物,一个Lacertid蜥蜴和一条Colubrid蛇,表示位于中更新世末期的年龄。尽管总体野生动物谱与中更新世马格里布遗址的其他组合相似,但Ben Kerat标本具有特殊的牙齿特征,这可能反映了在干旱化和降温增加的更广泛背景下的气候改善期,这是一个区域特征,特别是新形式的副精神病和MUS不属于北非迄今已知的进化谱系。
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引用次数: 0
First fossil of Varanus Merrem, 1820 (Squamata: Varanidae) from the Miocene Siwaliks of Pakistan 1820年巴基斯坦中新世Siwaliks地区Varanus Merrem的首个化石(鳞片目:Varanidae)
IF 1.8 3区 地球科学 Q2 PALEONTOLOGY Pub Date : 2022-02-14 DOI: 10.5252/geodiversitas2022v44a7
A. Villa, M. Delfino
ABSTRACT In the ‘50s of the XX century, a German expedition lead by Richard Dehm collected a large amount of fossil remains from northern Pakistan. Among these was an isolated trunk vertebra of a lizard, which is here referred to Varanus sp. The collecting site of this specimen is not precisely known, but it most likely comes from middle to early late Miocene Siwalik sediments of the Chinji Formation. This is the first published record of a fossil lizard from the Neogene of Pakistan and adds to the very patchy record of Varanus Merrem, 1820 in Asia. It further supports previous reconstructions of a warm climate for the middle to early late Miocene of the Siwaliks.
20世纪50年代,理查德·德姆率领的一支德国探险队在巴基斯坦北部收集了大量化石遗骸。在这些化石中,有一个孤立的蜥蜴躯干椎骨,这里称之为Varanus sp.。该标本的采集地点尚不清楚,但很可能来自中新世中晚期的Chinji组Siwalik沉积物。这是首次公布的来自巴基斯坦新近纪的蜥蜴化石记录,并为1820年亚洲瓦拉纳斯·梅里姆(Varanus Merrem)的非常不完整的记录增添了新的内容。它进一步支持了先前关于中新世中期到晚中新世早期的温暖气候的重建。
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引用次数: 2
A re-appraisal of the middle-late Miocene fossil decapod crustaceans of the ‘Faluns’ (Anjou-Touraine, France) 中新世中晚期“法轮人”十足目甲壳类动物化石的重新评估(法国安茹-图林)
IF 1.8 3区 地球科学 Q2 PALEONTOLOGY Pub Date : 2022-02-10 DOI: 10.5252/geodiversitas2022v44a6
À. Ossó, Cyril Gagnaison, Olivier Gain
ABSTRACT The access to a new collection of decapod crustaceans collected from the middle-late Miocene ‘Faluns’, and examination of new specimens available, have allowed to expand the previous compilations by Couffon (1908) and Ossó & Gagnaison (2019), elevating to nineteen the species reported, representatives of seventeen genera and fourteen families. Squat lobsters and parthenopid crabs are reported for the first time in the Atlantic Miocene of France. The expected presence of Xantho moldavicus (Yanakevich, 1977) and Lobocarcinus sismondai (von Meyer, 1843), hitherto known only by chelae, is confirmed herein. The status of Haydnella pulchellus (A. Milne-Edwards, 1864) is discussed. Moreover, the ventral features of Necronectes michelini (A. Milne-Edwards, 1861) are described for the first time. This decapod assemblage presents clear affinity with coeval decapod faunas from the Mediterranean and Paratethys realm, as evidenced by the presence of identical species such as Liocarcinus kuehni (Bachmayer, 1953) or Pilumnus mediterraneus (Lőrenthey, 1897), which is by far, the most common decapod in the ‘Faluns’ outcrops. Several different chelae and carapace remains are left in open nomenclature.
从中新世中晚期“Faluns”收集的新收集的十足甲壳类动物,以及对现有新标本的检查,使Couffon(1908)和Ossó & Gagnaison(2019)的先前汇编得以扩展,将报告的物种增加到19种,代表17属和14科。法国大西洋中新世首次报道了深蹲龙虾和孤雌蟹。本研究证实了迄今为止仅通过螯合动物得知的黄索moldavicus (Yanakevich, 1977)和Lobocarcinus sismondai (von Meyer, 1843)的存在。讨论了海螺的地位(A. Milne-Edwards, 1864)。此外,还首次描述了Necronectes michelini (A. Milne-Edwards, 1861)的腹侧特征。这种十足动物组合与来自地中海和Paratethys领域的同时期十足动物有明显的亲缘关系,如Liocarcinus kuehni (Bachmayer, 1953)或Pilumnus mediterraneus (Lőrenthey, 1897)等相同物种的存在就证明了这一点,后者是迄今为止“Faluns”露头中最常见的十足动物。几种不同的螯合肢和甲壳残骸被保留在开放的命名法中。
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引用次数: 1
Inventory of Cenozoic radiolarian species (Class Polycystinea) – 1834-2020 新生代放射虫(多胱虫纲)物种清查- 1834-2020
IF 1.8 3区 地球科学 Q2 PALEONTOLOGY Pub Date : 2022-02-03 DOI: 10.5252/geodiversitas2022v44a5
L. O’Dogherty, Noritoshi Suzuki, J. Caulet, P. Dumitrică
ABSTRACT This paper presents, for the first time ever, a complete list of Cenozoic Polycystinea reported between 1834 and 2020. It records 6898 names of taxa originally described as new species or subspecies, assigned to the Class of Polycystinea, the most important group in the infraphylum Radiolaria. This list only attempts to provide an objective record of available and unavailable names, the latter including nomina oblita, nomina nuda, homonyms, invalid nomenclatorial acts, species wrongly described as Polycystinea and nomina dubia species with inexistent name-bearing specimens.
摘要本文首次列出了1834年至2020年间报道的新生代多胱氨酸属的完整目录。它记录了6898个最初被描述为新种或亚种的分类群的名称,这些分类群归属于多胱菌纲,这是放射虫纲中最重要的类群。此列表仅试图提供现有和不可用名称的客观记录,后者包括名义上的义务,名义上的努达,同音异义,无效的命名行为,被错误地描述为polycystiinea的物种和不存在同名标本的名义上的dubia物种。
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引用次数: 1
A 100-million-year-old ensiferan with unusual mouthparts and comments on the evolution of raptorial appendages within Polyneoptera 1亿年前有不寻常口器的敏感动物,并评论了多翅目中猛禽附属物的进化
IF 1.8 3区 地球科学 Q2 PALEONTOLOGY Pub Date : 2022-01-27 DOI: 10.5252/geodiversitas2022v44a4
J. Haug, M. Hörnig, C. Kiesmüller, P. Pazinato, V. Baranov, C. Haug
ABSTRACT Myanmar amber is known to provide fossils of the group Insecta with surprising morphologies. Here we present fossils of the new species Gryllobencain patrickmuelleri n. gen., n. sp. from Myanmar amber that possess an overall “orthopteroid” morphology, hence resemble crickets. Unlike in most crickets, thorax appendages 1 and 2 (“legs”) are large and prominent, even in comparison to thorax appendage 3. Furthermore, thorax appendages 1 and 2 are able to fold against themselves and are armed with prominent spine-like setae, indicating that these legs were originally used for grasping prey. Most prominent is a large spine on the tibia of both appendages distantly reminding of the tibial spur or claw of praying mantises. Comparable prey-catching apparatuses have evolved repeatedly in the group Polyneoptera: gladiators (Mantophasmatodea), three ingroups of bush-crickets (Saginae, Austrosaginae, Listroscelidinae) as well as one species of fossil mantises (Santanmantis axelrodi Grimaldi, 2003) appear to have used thorax appendages 1 and 2 for prey catching. The new fossils do not seem to be closely related to any of these groups. They differ especially by prominent cerci equipped with numerous, probably mechano-sensorial setae. In the other five groups, the cerci are significantly smaller, often indistinct. The fossils furthermore have prominent maxillae with sickle-shaped proximal parts and well-armed large palps, indicating that the maxillae played an important role in processing the prey. While the thorax appendages hence represent a case of clear convergent evolution, the uniqueness of the feeding apparatus leads us to recognise the fossils as a highly specialised new species.
缅甸琥珀为昆虫类化石提供了令人惊讶的形态。在这里,我们展示了来自缅甸琥珀的新物种Gryllobencain patrickmuelleri n. gen., n. sp.的化石,它们具有整体的“直足类”形态,因此与蟋蟀相似。与大多数蟋蟀不同,1号和2号胸附肢(“腿”)大而突出,甚至与3号胸附肢相比也是如此。此外,胸部附属物1和2能够折叠起来,并具有突出的棘状刚毛,表明这些腿最初是用来抓猎物的。最突出的是两个附属物胫骨上的大脊柱,使人想起螳螂的胫骨刺或爪子。在多翅目中,类似的捕食工具也在不断进化:角斗士(Mantophasmatodea),三组灌木蟋蟀(Saginae, austrrosaginae, Listroscelidinae)以及一种化石螳螂(santan螳螂axelrodi Grimaldi, 2003)似乎都使用了胸部附件1和2来捕捉猎物。这些新发现的化石似乎与这些群体中的任何一个都没有密切的关系。它们的不同之处在于突出的尾部装有大量的,可能是机械感觉的刚毛。在其他五组中,cerci明显较小,通常不明显。此外,这些化石还具有突出的上颌骨,近端呈镰刀状,前臂发达,表明上颌骨在处理猎物方面发挥了重要作用。虽然胸部附属物因此代表了一个明显趋同进化的例子,但进食装置的独特性使我们认识到这些化石是一个高度专业化的新物种。
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引用次数: 3
A Jacobson's beetle from Cretaceous Charentese amber (Coleoptera: Jacobsoniidae) 白垩纪夏伦氏琥珀中的一种雅各布森甲虫(鞘翅目:雅各布森甲虫科)
IF 1.8 3区 地球科学 Q2 PALEONTOLOGY Pub Date : 2022-01-20 DOI: 10.5252/geodiversitas2022v44a3
Erik Tihelka, D. Peris, Chen-yang Cai, V. Perrichot
ABSTRACT Jacobsoniidae is a species-poor family of minute polyphagan beetles distributed in tropical and subtropical regions. Both extant representatives of the family as well as fossils are rare. Here we describe a new fossil species, Derolathrus capdoliensis n. sp., from latest Albian-earliest Cenomanian Charentese amber from the Cadeuil deposit, Charente-Maritime department in south-western France. The new species is defined by several unusual characters, most notably the co-joined but not fully fused two apical antennomeres. Derolathrus capdoliensis n. sp. is approximately contemporaneous with Kachin amber (burmite), filling an important geographical gap in the Mesozoic distribution of the family. The widespread distribution of jacobsoniid beetles in the Cretaceous, encompassing the Tethyan and Austral realms, indicates a more widespread distribution of the family during this time than in the present day and suggests that the current biogeographical range of Jacobsoniidae may be a result of extinction in northern regions. The new species shows remarkable similarity to extant members of the genus and provides further evidence of prolonged morphological, and probably also ecological, stasis in Jacobsoniidae since at least the Cretaceous.
雅各布斯甲虫科是分布于热带和亚热带地区的一个物种贫乏的微小多角甲虫科。该家族现存的代表和化石都很罕见。在这里,我们描述了一个新的化石物种,Derolathrus capdoliensis n.sp.,来自法国西南部夏朗德海事部Cadeuil矿床的最新阿尔比-最早的Cenomanian夏朗德琥珀。新物种由几个不寻常的特征来定义,最显著的是两个顶端触角单体共同连接但未完全融合。capdoliensis n.sp.的Derolathrus与克钦琥珀(缅甸)大致同时代,填补了该科中生代分布的一个重要地理空白。雅各布斯甲虫在白垩纪的广泛分布,包括特提斯和澳大利亚地区,表明该科在这一时期的分布比现在更为广泛,并表明雅各布斯甲虫科目前的生物地理范围可能是北部地区灭绝的结果。该新物种与该属现存成员表现出显著的相似性,并提供了进一步的证据,证明至少自白垩纪以来,雅各布斯科的形态和生态都长期停滞。
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引用次数: 2
Equids from the late Middle Pleistocene to Early Holocene of the Apulia Peninsula (southern Italy): reassessment of their taxonomy and biochronology 意大利南部普利亚半岛中更新世晚期至全新世早期的马科动物:分类和生物年代学的重新评估
IF 1.8 3区 地球科学 Q2 PALEONTOLOGY Pub Date : 2022-01-14 DOI: 10.5252/geodiversitas2022v44a2
B. Mecozzi, Flavia Strani
ABSTRACT The Apulian Peninsula represents a key-region for the study of climatic changes and paleoenvironmental dynamics during the Quaternary. Both large and small-sized horses are well documented in this region during the Pleistocene and are frequently found associated. The caballoid horses from Middle to Late Pleistocene of Europe show a large intraspecific ecomorphological variation, emphasizing a strong link between skeletal adaptations and specific aspects of the palaeoenvironment. This large variability led to an extended debate concerning the taxonomy of equids and their evolutionary history. In the Italian Peninsula, fossils from the Middle to the Late Pleistocene have been historically referred to several species (or even subspecies), emphasizing the uncertainty of the taxonomic attribution. Here, a large craniodental sample of Equidae fossils from late Middle Pleistocene to the Early Holocene localities of the Apulia Peninsula are described for the first time. The comparison of the protocone index allowed us to determine the first occurrences of Equus mosbachensis von Reichenau, 1903 in the Apulian Peninsula, from a few localities chronologically referred to late Middle Pleistocene. Most of the material from the late Middle to the end of the Late Pleistocene is instead attributed to Equus ferus Boddaert, 1785. The results of this work thus add novel information on the biochronology of Italian fossil equids and their evolutionary history through the Middle and Late Pleistocene.
阿普利亚半岛是研究第四纪气候变化和古环境动力学的关键区域。在更新世期间,该地区的大型和小型马都有很好的记录,并且经常被发现有关联。欧洲中更新世至晚更新世的caballoid马显示出较大的种内生态形态变异,强调了骨骼适应与古环境特定方面之间的紧密联系。这种巨大的变异导致了关于马科动物分类及其进化史的长期争论。在意大利半岛,从中新世到晚更新世的化石在历史上被称为几个物种(甚至亚种),强调了分类归属的不确定性。本文首次描述了阿普利亚半岛中更新世晚期至全新世早期的大型马科头骨化石样本。原始锥体指数的比较使我们能够确定1903年在阿普里亚半岛最早出现的Equus mosbachensis von Reichenau,从几个年代上被称为中更新世晚期的地方。大多数中晚期到晚更新世末期的材料被认为是Equus ferus Boddaert, 1785。因此,这项工作的结果为意大利化石马科动物的生物年代学及其在中新世和晚更新世的进化史增加了新的信息。
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引用次数: 2
The evolution of hermit crabs (Crustacea, Decapoda, Anomura, Paguroidea) on the basis of carapace morphology: a state-of-the-art-report 寄居蟹(甲壳纲、十足目、无尾目、异足目)在甲壳形态基础上的进化:最新报告
IF 1.8 3区 地球科学 Q2 PALEONTOLOGY Pub Date : 2022-01-06 DOI: 10.5252/geodiversitas2022v44a1
René H. B. Fraaije, B. V. van Bakel, J. Jagt, S. Charbonnier, G. Schweigert, G. Garcia, Xavier Valentin
ABSTRACT In our ongoing studies of both extinct and extant hermit crabs (Paguroidea Latreille, 1802), we have observed and recorded a number of morphological changes that have taken place between Early Jurassic times (c. 185 million years ago) and the present day. Species compositions of paguroid assemblages from marine Upper Jurassic and Lower Cretaceous rocks in Europe are here compared with those of modern marine settings. Basal paguroids with cylindrical carapaces predominated in reefal environments during the Late Jurassic, but were gradually replaced by hermit crabs with non-cylindrical carapaces. The most basal paguroids reveal a branchial groove, but branchial condensation towards a single (i.e., cervical) groove is here shown to have occurred early in their evolutionary history. In several extant, genera remnants of a branchial groove, in combination with several other basal features, can be seen. For this reason, these are here considered to be the most primitive members amongst extant paguroids. In this study, we resurrect the family Probeebeidae and erect a new family, Paguropsidae n. fam, to accommodate extant basal hermit crabs such as Paguropsis Henderson, 1888, Eopaguropsis Fraaije, Krzemiński, Van Bakel, Krzemińska & Jagt, 2012 and Paguropsina Lemaitre, Rahayu & Komai, 2018. Described are also one pair of Early Jurassic (Pliensbachian) and two pairs of Middle Jurassic (Callovian) isochelous paguroid chelae, all collected recently. A new specimen of Schobertella reveals some important morphological traits of the shield that had not been observed previously. The earliest known example to date of clearly heterochelous chelae in the fossil record of hermit crabs originates from upper Kimmeridgian/Tithonian strata in the Boulonnais (northwestern France). For the first time, a phylogenetic scheme of marine Paguroidea, as based on morphological data of carapaces, is presented.
在我们对已灭绝和现存寄居蟹(Paguroidea Latreille, 1802)的持续研究中,我们观察并记录了早侏罗世(约1.85亿年前)到现在发生的许多形态变化。本文将欧洲海相上侏罗统和下白垩统岩石中的类石组合的物种组成与现代海相环境中的类石组合进行了比较。在晚侏罗世的珊瑚礁环境中,具有圆柱形甲壳的基缘蜈蚣类占优势,但逐渐被具有非圆柱形甲壳的寄居蟹所取代。最基部的类鱼显示鳃裂沟,但此处显示向单一(即颈)沟的鳃裂凝结在其进化史的早期发生。在几个现存的属中,可以看到鳃沟的残余物,与其他几个基底特征结合在一起。因此,它们被认为是现存的类人猿中最原始的成员。在这项研究中,我们复活了Probeebeidae家族,并建立了一个新的家族,Paguropsidae n. fam,以容纳现有的基寄居蟹,如Paguropsis Henderson, 1888, Eopaguropsis Fraaije, Krzemiński, Van Bakel, Krzemińska & Jagt, 2012和Paguropsina Lemaitre, Rahayu & Komai, 2018。还发现了一对早侏罗世(Pliensbachian)和两对中侏罗世(Callovian)等掌类螯动物,均为近期发现。一个新的肖伯特拉标本揭示了一些以前没有观察到的盾的重要形态特征。在寄居蟹的化石记录中,迄今已知最早的明显异螯螯的例子来自法国西北部布龙尼斯的上基默里吉纪/提托尼统地层。本文首次根据甲壳的形态资料,提出了一种海洋巨蛛总科的系统发育方案。
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引用次数: 5
Jean-Claude Rage, a life dedicated to paleoherpetology Jean-Claude Rage,致力于古爬虫学的一生
IF 1.8 3区 地球科学 Q2 PALEONTOLOGY Pub Date : 2021-12-20 DOI: 10.5252/geodiversitas2021v43a27
J. Steyer, M. Augé, G. Métais
ABSTRACT Jean-Claude Rage (1943-2018) was a renowned vertebrates paleontologist who dedicated his career to fossil amphibians and reptiles, which earned him the highest reputation. He was CNRS Emeritus Director of Research in Paris and worked in the laboratory of paleontology of the Muséum national d'Histoire naturelle (MNHN) together with the authors of this article (JSS and MA were also his Master and PhD students). Jean-Claude has established himself as an outstanding herpetologist and taxonomist with numerous publications (more then 250 articles, see below) throughout his career. The aim of this article is not to describe in details his very rich career (see Steyer & Buffetaut 2012 for details) but to show how important his research was, and is still, and to go deeper in his personality to show that his intellectual legacy should be considered as a model today.
摘要让-克洛德·拉格(Jean-Claude Rage,1943-2018)是一位著名的脊椎动物古生物学家,他的职业生涯致力于两栖爬行动物化石研究,这为他赢得了最高的声誉。他是CNRS在巴黎的名誉研究主任,与本文作者一起在国家自然历史博物馆(MNHN)的古生物学实验室工作(JSS和MA也是他的硕士和博士生)。让-克劳德在其职业生涯中发表了大量出版物(超过250篇文章,见下文),成为一位杰出的爬虫学家和分类学家。这篇文章的目的不是详细描述他非常丰富的职业生涯(详见Steyer&Buffetaut 2012),而是展示他的研究过去和现在是多么重要,并深入了解他的个性,以表明他的智力遗产应该被视为当今的典范。
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引用次数: 0
Bryozoan faunas at the Tortonian-Messinian transition. A palaeoenvironmental case study from Crete Island, eastern Mediterranean 托尔顿-墨西尼亚过渡时期的苔藓虫动物群。地中海东部克里特岛的古环境案例研究
IF 1.8 3区 地球科学 Q2 PALEONTOLOGY Pub Date : 2021-12-16 DOI: 10.5252/geodiversitas2021v43a26
P. Moissette, A. Antonarakou, G. Kontakiotis, J. Cornée, V. Karakitsios
ABSTRACT Four sedimentary sections were logged and sampled from upper Tortonian-lower Messinian outcrops on the island of Crete (Greece). The collected material yielded about 60 bryozoan species belonging to nine different colonial morphotypes. A few species are stenobathic, indicating either shallow- or deep-water environments, but most of them are eurybathic (with bathymetric ranges extending in some instances from the shelf down to several hundreds of metres). Bryozoan communities point to sea-level variations modulated by local tectonics. Deep circalittoral environments were recognized at the base of three sections (whereas the fourth section starts with infralittoral/shallow circalittoral environments passing to deep circalittoral. Shallow bathyal habitats follow upwards, succeeded in turn by assemblages indicative of deep circalittoral and subsequently shallow circalittoral/infralittoral depths. Inferred bathymetric fluctuations are diachronous throughout the island. Shallow-water species found associated, sometimes abundantly, with rarer deep-water faunas in a few levels, are interpreted as transported by currents. Several dysoxic episodes have been also detected: likely the result of water column stratification and enhanced productivity. These processes were probably boosted by a combination of changes in oceanic circulation, climate, global sea-level, and the local/regional tectonics (in Crete and/or the marine gateways between the Mediterranean and the Atlantic).
摘要从克里特岛(希腊)上托托阶-下墨西阶露头中记录并采样了四个沉积剖面。收集到的材料产生了大约60种苔藓虫,属于9种不同的群落形态类型。少数物种是狭带生物,表明它们要么是浅水环境,要么是深水环境,但它们中的大多数都是裸泳生物(在某些情况下,水深范围从大陆架延伸到数百米)。苔藓虫群落指出海平面变化受当地构造的调节。在三个部分的基础上识别出深环形环境(而第四节从近海/浅水环流环境开始,过渡到深水环流。浅水生境向上延伸,依次是指示深水环流和随后的浅水环流/近海深度的组合。推断的水深波动在整个岛屿上是跨时的,有时数量丰富,在几个层面上有罕见的深水动物群,被解释为由洋流输送。还发现了一些中毒事件:可能是水柱分层和生产力提高的结果。这些过程可能是由海洋环流、气候、全球海平面和当地/区域构造(克里特岛和/或地中海和大西洋之间的海洋门户)的变化共同推动的。
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引用次数: 0
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Geodiversitas
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