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The rostral neurovascular system of Tyrannosaurus rex 霸王龙的嘴侧神经血管系统
IF 2 4区 地球科学 Q1 Earth and Planetary Sciences Pub Date : 2022-01-01 DOI: 10.26879/1178
Florian Bouabdellah, Emily J Lessner, J. Benoit
The study of the rostral neurovascular system using CT scanning has shed new light on phylogenetic and palaeobiological reconstructions of many extinct tetrapods. This research shows a detailed description of the rostral neurovascular canals of Tyrannosaurus rex including the nasal, maxillary (dorsal alveolar), and mandibular (ventral alveolar) canals. Extensive comparisons with published descriptions show that the pattern of these canals in Tyrannosaurus is not unusual for a non-avian theropod. As in the non-avian theropod Neovenator, the maxillary canal shows several anastomoses of its branches. Differences from the plesiomorphic sauropsid condition are concentrated within the canal for the maxillary neurovasculature, which is primitively horizontal, tubular, and connected to a single row of supralabial foramina, whereas in Tyrannosaurus the main trunk of the canal is oriented more obliquely and dorsally displaced to give room to the deep tooth alveolae. As a result, the lateral branches that provide innervation and blood supply to the skin are dorsoventrally elongated compared to non-theropod taxa, and multiple rows of supralabial foramina are present. An overview of the literature suggests that the evolution of the trigeminal canals among sauropsids only weakly supports previous hypotheses of crocodile-like facial sensitivity in non-avian theropods (except, maybe, in semiaquatic taxa). More systematic studies of the rostral neurovascular canals in non-avian theropods may help answer the question of whether lips were present or not. Florian Bouabdellah. Universities of Poitiers, 15 Rue de l'Hôtel Dieu, TSA 71117, 86000 Poitiers, France. florian.bouabdellah@hotmail.com Emily Lessner. University of Missouri School of Medicine, Columbia, MO, 65211, USA. ejlessner@mail.missouri.edu Julien Benoit. Evolutionary Studies Institute and School of Geosciences, University of the Witwatersrand, Johannesburg 2050, South Africa, julien.benoit@wits.ac.za
利用CT扫描对吻侧神经血管系统的研究为许多已灭绝的四足动物的系统发育和古生物学重建提供了新的思路。本研究详细描述了霸王龙的鼻侧神经血管管,包括鼻、上颌(牙槽背侧)和下颌(牙槽腹侧)的神经血管管。与已发表的描述进行的广泛比较表明,暴龙身上这些沟渠的模式对于非鸟类兽脚亚目动物来说并不罕见。与非鸟类兽脚亚目Neovenator一样,上颌管显示出其分支的几个吻合处。与多形龙的不同之处在于上颌神经血管的管道内,它是原始的水平管状,与单排颚上孔相连,而暴龙的管道主干定向更斜,并向背侧移位,以便为深牙槽提供空间。因此,与非兽脚亚目动物相比,为皮肤提供神经支配和血液供应的侧支是背侧延长的,并且存在多行颚上孔。文献综述表明,蜥脚类动物三叉神经管的进化只能微弱地支持先前的假设,即非鸟类兽脚亚目动物具有类似鳄鱼的面部敏感性(可能半水生分类群除外)。对非鸟类兽脚亚目动物的吻侧神经血管管进行更系统的研究可能有助于回答嘴唇是否存在的问题。Florian Bouabdellah。普瓦捷大学,15 Rue de l'Hôtel Dieu, TSA 71117,法国普瓦捷86000。florian.bouabdellah@hotmail.com Emily Lessner。美国密苏里大学医学院,哥伦比亚,密苏里州,65211ejlessner@mail.missouri.edu朱利安·伯努瓦。威特沃特斯兰德大学进化研究所和地球科学学院,约翰内斯堡2050,南非,julien.benoit@wits.ac.za
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引用次数: 7
Amateur collectors are critical to the study of fossil vertebrates: A case study from two Neogene localities in Northern California (Santa Margarita and Purisima formations) 业余收藏家对脊椎动物化石的研究至关重要:来自北加州两个新近纪地区(圣玛格丽塔和普里西马地层)的案例研究
IF 2 4区 地球科学 Q1 Earth and Planetary Sciences Pub Date : 2022-01-01 DOI: 10.26879/1199
R. Boessenecker
Vertebrate paleontology was born from the efforts of amateur and commercial fossil collectors in the nineteenth century. Amateur fossil collecting is a popular hobby in the USA, though owing to different ownership laws, American vertebrate paleontologists have less positive attitudes toward amateur collectors than in Europe where amateur and professional collectors work synergistically. Collections-based and literature surveys were conducted to evaluate the scientific contribution of amateurs to vertebrate paleontology near Santa Cruz, California. The first was a survey of museum collections identifying collector status (amateur or professional) of fossils from two formations (Santa Margarita Sandstone, Purisima Formation). The second was a com-prehensive literature survey for these two stratigraphic units, documenting whether fossils were collected by amateurs or professionals. The third was a literature survey of all published (2009-2021) Cenozoic marine vertebrate records for the Pacific coast of North America (Alaska to Baja California, Mexico). The first survey indicates that amateurs have contributed the most (75.9%) to Santa Margarita Formation specimens and over a third (38.1%) of Purisima Formation specimens. These contributions are of high quality as they are included in 40% of all publications through time in the study area, and amateur-collected fossils are reported in half (49%) of all publications from the Pacific coast over the past decade. These findings indicate that amateur collectors are not only capable of collecting scientifically significant specimens, but appear to be integral to the study of Cenozoic marine vertebrates on the Pacific coast. Dismissive attitudes towards amateur collectors are clearly unwarranted. Advice for cultivating strong professional-amateur relations is provided.
脊椎动物古生物学是在19世纪业余和商业化石收藏家的努力下诞生的。业余化石收集在美国是一种流行的爱好,尽管由于不同的所有权法律,美国脊椎动物古生物学家对业余收藏家的态度不如业余和专业收藏家协同工作的欧洲那么积极。通过收集资料和文献调查来评估业余爱好者对加州圣克鲁斯附近脊椎动物古生物学的科学贡献。首先是对博物馆藏品进行调查,确定两个地层(圣玛格丽塔砂岩和普里西马地层)化石的收藏家身份(业余或专业)。第二项是对这两个地层单位进行全面的文献调查,记录化石是由业余爱好者还是专业人士收集的。第三项是对北美太平洋沿岸(阿拉斯加至墨西哥下加利福尼亚州)所有已发表的新生代海洋脊椎动物记录(2009-2021年)的文献调查。第一次调查表明,业余爱好者对Santa Margarita地层标本的贡献最大(75.9%),对Purisima地层标本的贡献超过三分之一(38.1%)。这些贡献是高质量的,因为它们包含在研究区域所有出版物的40%中,并且在过去十年中,来自太平洋沿岸的所有出版物中有一半(49%)报告了业余收集的化石。这些发现表明,业余收藏家不仅能够收集具有科学意义的标本,而且似乎对太平洋沿岸新生代海洋脊椎动物的研究不可或缺。对业余收藏家的轻蔑态度显然是没有根据的。为培养专业与业余的牢固关系提供了建议。
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引用次数: 0
Phylogeny of iguanodontian dinosaurs and the evolution of quadrupedality 禽龙类恐龙的系统发育与四足动物的进化
IF 2 4区 地球科学 Q1 Earth and Planetary Sciences Pub Date : 2022-01-01 DOI: 10.26879/702
Karen Poole
Iguanodontians are a large and biogeographically widespread group of dinosaurs, known from every modern continent, with a temporal range from the Late Jurassic through the Late Cretaceous. While the nested hadrosauroids have been studied extensively, the phylogeny of non-hadrosauroid iguanodontians remains less clear. This study presents a character matrix with 323 characters, and both parsimony and time-calibrated Bayesian analyses. While these result in different topologies, they both recover a Thescelosauridae outside of Iguanodontia. Within Iguanodontia, they both recover Muttaburrasaurus and Tenontosaurus as sister taxa to Rhabdodontidae, with a larger group of Gondwanan Rhabdodontoidea (nomen cladi novum) in the Bayesian analysis. A small Dryosauridae forms the sister group to Ankylopollexia, which has Uteodon and Camptosaurus as the most basally branching taxa. Within Styracosterna two distinct clades are recovered: Iguanodontidae, and a group of taxa with robust forelimbs. The holotype of Mantellisaurus is sister to “Dollodon”, supporting the hypothesis that these taxa are synonymous. The “hatchet-shaped” sternal thought to be a synapomorphy of Styracosterna occurs in two taxa recovered outside that group: Macrogryphosaurus and the unnamed taxon from the Kirkwood Formation of South Africa. Characters associated with quadrupedality are mapped on the phylogeny, indicating a transition from bipedality to quadrupedality occurred in a stepwise manner at the base of Ankylopollexia. Based on synapomorphies of the groups, the major innovations in Ankylopollexia were postcranial, while those of hadrosauroids were centered on dentition and the dentaries. There is a clear faunal succession from the Late Jurassic to Early Cretaceous non-hadrosauroid ankylopollexians to the Late Cretaceous hadrosauroids. Karen Poole. Department of Basic Sciences, New York Institute of Technology College of Osteopathic Medicine at Arkansas State University, Wilson Hall, 2405 Aggie Rd., Jonesboro Arkansas 72401, USA. kpoole@nyit.edu
禽龙是一种生物地理上分布广泛的大型恐龙,在现代各大洲都有发现,其时间范围从晚侏罗世到晚白垩纪。虽然巢类鸭嘴龙已经被广泛研究,但非鸭嘴龙禽龙的系统发育尚不清楚。本文提出了一个包含323个字符的字符矩阵,并进行了简约性和时间校准贝叶斯分析。虽然这些结果导致不同的拓扑结构,但它们都恢复了禽龙目之外的这些龙目。在禽龙目中,他们都恢复了muttaburrasurus和Tenontosaurus作为Rhabdodontidae的姐妹分类群,并在贝叶斯分析中发现了一个更大的Gondwanan Rhabdodontoidea (nomen cladi novum)群。一个小的旱龙科形成了甲龙的姐妹类群,甲龙的最基本分支类群是虎齿龙和喜树龙。在Styracosterna中恢复了两个不同的分支:禽龙科和一组具有强壮前肢的分类群。Mantellisaurus的完整型是“Dollodon”的姐妹,支持了这些分类群是同义词的假设。被认为是Styracosterna近亲的“斧头状”胸骨出现在该类群之外的两个分类群中:巨形龙和来自南非柯克伍德组的未命名分类群。与四足相关的特征在系统发育上进行了映射,表明从两足到四足的过渡是在强直阅读的基础上逐步发生的。从类群的突触形态来看,甲龙的主要进化发生在颅后,而鸭嘴龙的进化主要集中在齿列和牙列。从晚侏罗世到早白垩世非鸭嘴龙类甲龙类到晚白垩世鸭嘴龙类有明显的区系演替。凯伦普尔。阿肯色州立大学纽约理工学院骨科医学院基础科学系,Wilson Hall, Jonesboro Aggie Rd. 2405, Arkansas 72401, USAkpoole@nyit.edu
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引用次数: 5
Feeding strategies of circum‑Mediterranean hipparionins during the late Miocene: Exploring dietary preferences related to size through dental microwear analysis 中新世晚期环地中海海马的摄食策略:通过牙齿微磨损分析探索与尺寸相关的饮食偏好
IF 2 4区 地球科学 Q1 Earth and Planetary Sciences Pub Date : 2022-01-01 DOI: 10.26879/990
Guillem Orlandi-Oliveras, M. Köhler, J. Clavel, R. Scott, Serdar Mayda, T. Kaya, G. Merceron
The adaptive radiation of hipparionins after their Old World dispersal was linked with a trend towards smaller body sizes. The appearance of the small-sized forms has usually been associated to open environments and grazing diets. A recent approach, moreover, highlights the role of life history modifications related to habitat conditions as triggers of their size shifts. Here, we test the relationship between hipparionin size and diet analyzing the dental microwear textures of different-sized hipparionins from Vallesian and Turolian circum-Mediterranean localities. Our results show that hipparionins were mainly mixed-feeders and that there was no general link between body size and diet. However, we identified broader feeding spectra in western Mediterranean smaller forms and specialized grazing diets in larger ones, a differentiation not found in the eastern Mediterranean hipparionins. At odds with the notion of more open habitats eastward, we detected a larger browsing component in eastern hipparionin diets. The consumption by extant equids of more woody browse during the dry season us to propose a greater seasonality as a possible cause. Considering the argu-able role of external abrasives on the microwear, another interpretation might involve the presence of more grit in the eastern opener habitats. Interestingly, we found that sympatric hipparionins tend to have similar feeding habits, which points to the fact that their diets were influenced by the local environment. Our results, then, suggest that the small size of some hipparionins resulted from different selective pressures rather than to a general adaptation to increasing habitat opening.
海马皂苷在旧大陆扩散后的适应性辐射与体型变小的趋势有关。小型形式的出现通常与开放环境和放牧饮食有关。此外,最近的一种方法强调了与栖息地条件相关的生活史变化作为其大小变化的触发因素的作用。在这里,我们测试了海马皂苷大小与饮食之间的关系,分析了来自巴列西亚和图罗里安地中海沿岸地区不同大小的海马皂苷的牙齿微磨损结构。我们的研究结果表明,海马素主要是混合食性动物,体型和饮食之间没有普遍的联系。然而,我们在地中海西部较小的形式中发现了更广泛的摄食光谱,在较大的形式中发现了专门的放牧饮食,而在地中海东部的海马中没有发现这种分化。与东部更开放的栖息地的概念不一致,我们在东部海马蛋白饮食中发现了更大的浏览成分。现存的马科动物在旱季消耗更多的木材,我们提出一个更大的季节性可能的原因。考虑到外部磨料对微磨损的作用,另一种解释可能涉及东部揭幕战生境中存在更多的砂砾。有趣的是,我们发现同域海马素倾向于有相似的饮食习惯,这表明它们的饮食受到当地环境的影响。因此,我们的结果表明,一些海马苷的小尺寸是由于不同的选择压力,而不是为了适应不断增加的栖息地开放。
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引用次数: 1
Amphibians and squamate reptiles from the late Miocene of Fălciu (Eastern Romania) 中新世晚期腓利丘(罗马尼亚东部)的两栖动物和有鳞爬行动物
IF 2 4区 地球科学 Q1 Earth and Planetary Sciences Pub Date : 2022-01-01 DOI: 10.26879/1156
V. Codrea, Marian Bordeianu, M. Venczel
,
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引用次数: 1
Chuandianella ovata: An early Cambrian stem euarthropod with feather-like appendages 卵川蝶:寒武纪早期的茎真节肢动物,有羽毛状附属物
IF 2 4区 地球科学 Q1 Earth and Planetary Sciences Pub Date : 2022-01-01 DOI: 10.26879/1172
Dayou Zhai, Mark Williams, D. Siveter, D. Siveter, T. Harvey, Robert S. Sansom, Huijuan Mai, Run-Bang Zhou, X. Hou
Micro-CT scanning reveals unprecedented three-dimensional soft anatomy of the early Cambrian (Epoch 2, Age 3) euarthropod Chuandianella ovata. We interpret the presence of an elongate, antenniform first appendage, and a short uniramous second appendage, followed by 10 homonomous biramous appendages comprising a short paddle-shaped exopod and a unique feather-like limb-branch with at least 27 podomeres each of which bears a long blade-like endite with a short terminal seta: we interpret this as the endopod. Alternative interpretations, that these limbs might represent an epipod+basipod or epipod+exopod arrangement, are unlikely, in that they would require either the complete reduction of the exopod or the endopod. We also find no evidence for head appendage morphologies that would support a more crownward position, for example among pancrustaceans, that has previously been suggested for
显微ct扫描揭示了早寒武纪(第2世,第3年)真节肢动物川蝶属卵形的三维软解剖结构。我们解释了一个细长的,天线状的第一附属物的存在,一个短的,不知名的第二附属物,然后是10个相同的双名附属物,包括一个短的桨形外足和一个独特的羽毛状肢分支,其中至少有27个足节,每个足节都有一个长刃状的端尖和一个短的端尖:我们将其解释为内足。另一种解释是,这些肢可能代表外足+基足或外足+外足的排列,这是不太可能的,因为它们要么需要完全减少外足,要么需要完全减少内足。我们也没有发现任何证据表明头部附属物的形态可以支持一个更朝上的位置,例如在以前提出的泛甲纲动物中
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引用次数: 1
Diptera of the Middle Eocene Kishenehn Formation II 中始新世Kishenehn组双翅目2
IF 2 4区 地球科学 Q1 Earth and Planetary Sciences Pub Date : 2022-01-01 DOI: 10.26879/1215
D. Greenwalt, D. Amorim, M. Hauser, P. Kerr, Scott M. Fitzgerald, S. Winterton, J. Cumming, N. Evenhuis, B. Sinclair
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引用次数: 2
Pygocephalomorphan crustaceans further emphasise the similarities between the Carboniferous Piesberg quarry in Germany and the Mazon Creek Lagerstätte in North America Pygocephalomorphan甲壳类动物进一步强调了德国石炭纪Piesberg采石场与北美Mazon Creek Lagerstätte之间的相似性
IF 2 4区 地球科学 Q1 Earth and Planetary Sciences Pub Date : 2022-01-01 DOI: 10.26879/1051
P. Pazinato, C. Haug, A. Leipner, J. Haug
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引用次数: 1
Putative Ordovician green alga Krejciella reinterpreted as enteropneust hemichordate tube (Czech Republic) 假定的奥陶纪绿藻Krejciella被重新解释为enteropneust half - hordate tube(捷克)
IF 2 4区 地球科学 Q1 Earth and Planetary Sciences Pub Date : 2022-01-01 DOI: 10.26879/1185
O. Fatka, J. Vodička
Fossil and extant representatives of Enteropneusta play an important role in the interpretation of early evolution of animals, such as echinoderms, hemichordates and chordates. However, remains of fossil Enteropneusta are rare. Re-examination of available specimens of organic tubes of the Ordovician putative green alga Krejciella putzkeri Obrhel 1968 does not show any morphological difference from the Cambrian Margaretia dorus Walcott, 1931. The latter species has been recently interpreted as an organic tube produced and inhabited by the worm-like enteropneust hemichordate Oesia disjuncta Walcott, 1911. However, the absence of the subterranean lateral extension in Ordovician specimens excludes the synonymy of Krejciella and Margaretia . Geographic distribution of Cambrian organic tubes classified as Margaretia Walcott, 1931 indicates a possible latitudinal control, as all occurrences are apparently restricted to tropical and subtropical belts when plotted in Cambrian palaeogeographic maps. In comparison, the occurrence of the herein studied specimens of Krejciella is restricted to cold-water localities of West Gondwana. The micropalaeontological analy-sis of a rock sample bearing one specimen of Krejciella shows the presence of moder-ately preserved chitinozoans, including the zonal species Linochitina pissotensis . This taxon is, for the first time, documented from the Prague Basin and determines the Mid-dle/Late Ordovician boundary interval of the analysed sample. The herein studied specimens of Krejciella extend the record of organic tubes produced by enteropneust hemichordates both stratigraphically and palaeogeographically to the Middle/Late Ordovician cold-water area.
Enteropneusta的化石和现存代表在解释棘皮动物、半足动物和脊索动物的早期进化中起着重要作用。然而,化石enteropnesta的遗迹是罕见的。对奥陶纪推定绿藻Krejciella putzkeri Obrhel 1968的有机管状标本的重新检查显示,与寒武纪Margaretia dorus Walcott, 1931的形态没有任何差异。后一种最近被解释为一种有机管,由蠕虫状的半翅虫Oesia disjuncta Walcott, 1911产生并居住。然而,奥陶纪标本中缺乏地下侧向伸展,排除了Krejciella和Margaretia的同义种。分类为Margaretia Walcott, 1931的寒武系有机管的地理分布可能受纬度控制,因为在寒武系古地理图中,所有有机管的分布明显局限于热带和亚热带带。相比之下,本文研究的克雷希氏菌标本的出现仅限于西冈瓦纳的冷水地区。对含有一个Krejciella标本的岩石样本进行的微体古生物学分析表明,存在中度保存的几丁质动物,包括地带性物种Linochitina pissotensis。该分类群首次在布拉格盆地中发现,并确定了样品的中晚奥陶世边界区间。本文所研究的Krejciella标本,在地层和古地理上都将半长尾藻(enteropnest半长尾藻)产生的有机管的记录扩展到了中晚奥陶世冷水区。
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引用次数: 2
Designing scientifically-grounded paleoart for augmented reality at La Brea Tar Pits 在拉布雷亚沥青坑为增强现实设计科学基础的古艺术
IF 2 4区 地球科学 Q1 Earth and Planetary Sciences Pub Date : 2022-01-01 DOI: 10.26879/1191
Matthew M. Davis, Benjamin D. Nye, G. Sinatra, W. Swartout, Molly Sjӧberg, Molly A. Porter, David Nelson, Alana A. U. Kennedy, Imogen Herrick, Danaan DeNeve Weeks, Emily Lindsey
Paleoart is an important medium that communicates scientific understanding about prehistoric life to both the public and researchers. However, despite its broad influence, the scientific and aesthetic decisions that go into paleoart are rarely described in formal academic literature or subjected to peer review. This is unfortunate, as paleoart can easily create and perpetuate misconceptions that are carried through generations of iterative popular media. As an example of what we hope will become a standard article type in paleontological journals, we describe the process and latest scientific research used to develop 13 new paleoart reconstructions of Ice Age animals found in the La Brea Tar Pits, including the saber-toothed cat, dire wolf, and teratorn. We adopted a stylized low polygon aesthetic for these three-dimensional (3D), animated virtual models both to support learning objectives and to optimize performance for smartphone based augmented reality (AR) experiences. We encourage all researchers to follow the example of this article by publishing paleoart descriptions for any major new work that, at a minimum, reference the aesthetic and scientific reasoning behind general posture and proportions, gross appearance of soft tissues, coloration, and behavior.
古艺术是向公众和研究人员传达关于史前生活的科学理解的重要媒介。然而,尽管古艺术有着广泛的影响,但它所涉及的科学和美学决策很少在正式的学术文献中被描述,也很少受到同行评审的影响。这是很不幸的,因为古艺术很容易创造和延续误解,而这些误解会被一代又一代的流行媒体所传播。作为一个我们希望成为古生物学期刊标准文章类型的例子,我们描述了在拉布雷亚沥青坑中发现的13种新的冰河时代动物的古艺术重建的过程和最新的科学研究,包括剑齿虎、可怕的狼和畸形兽。我们对这些三维(3D)动画虚拟模型采用了风格化的低多边形美学,以支持学习目标并优化基于智能手机的增强现实(AR)体验的性能。我们鼓励所有的研究人员效仿这篇文章,为任何重要的新作品发表古艺术描述,至少参考一般姿势和比例、软组织的大体外观、颜色和行为背后的美学和科学推理。
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引用次数: 2
期刊
Palaeontologia Electronica
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