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New association Phragmitetum altissimi ass. nov. (Phragmito-Magnocaricetea Klika in Klika et Novák 1941) from the european part of Russia and Western Siberia 来自俄罗斯欧洲部分和西西伯利亚的新协会Phragmitetum altissimi ass.nov.(Phragmito Magnocaritea Klika in Klika et novák 1941)
Q4 Agricultural and Biological Sciences Pub Date : 2022-01-01 DOI: 10.31111/vegrus/2022.45.74
O. Kapitonova, Т. M. Lysenko
The highest reed (Phragmites altissimus) is a species with Eurasian-North African range, recently expanding its area of distribution in northern direction (Kapitonova, 2016; Golovanov et al., 2019; Tzvelev, Probatova, 2019). It is known that in the forest zone of both the European and Asian parts of Russia, the highest reed is found only as an invasive plant (Tzvelev, 2011). Communities dominated by P. altissimus are known both within its natural range and in the area of invasion. However, in syntaxonomic reviews, cenoses with this species dominanation are traditionally included by the authors in the ass. Phragmitetum australis Savich 1926 (Golub et al., 1991, 2015; Golub, Chorbadze, 1995; Kipriyanova, 2008; Vegetaсе…, 2011; Golovanov, Abramova, 2012; Chepinoga, 2015). The aim of this work is to establish the syntaxonomic status of communities formed by P. altissimus. The work used 65 geobotanical relevés made within the primary range of the P. altissimus (Astrakhan region and the south of the Tyumen region within the forest-steppe zone) and in the area of its secondary range (the Udmurtian Republic and the taiga zone of the Tyumen region). The relevés were introduced into database developed on the basis of the TURBOVEG program (Hennekens, 1996) and processed using the JUICE program (Tichý, 2002). To assess the abundance of species on the sample plots described, the J. Braun-Blanquet abundance scale was used with the following abundance-coverage scores: r — the species is extremely rare with insignificant coverage, + — the species is rare, the degree of coverage is small, 1 — the number of individuals is large, the degree of coverage is small or the individuals are sparse, but the coverage is large, 2 — the number of individuals is large, the projective cover is from 5 to 25 %, 3 — the number of individuals is any, the projective cover is from 25 to 50 %, 4 — the number of individuals is any, the projective cover is from 50 to 75 %, the number of individuals is any, the cover is more than 75 % (Mirkin et al., 1989). Syntaxonomic analysis was performed using the approach suggested by J. Braun-Blanquet (1964). The names of syntaxa are given according to the “International Code of Phytosociological Nomenclature” (Theurillat et al., 2021). The system of higher syntaxa is given in accordance with “Hierarchical floristic classification…” (Mucina et al., 2016). To identify the main factors determining the differentiation and distribution of the studied communities, the NMDS method was used. For each syntaxon, using the IBIS program (Zverev, 2007), the average indicator values were calculated according to the ecological scales of D. N. Tsyganov (Tsyganov, 1983): soil moisture (Hd), soil nitrogen richness (Nt), and illumination-shading (Lc). Processing was carried out in the PC-ORD v. 6.0 (McCune et Mefford, 2011). The studied communities were assigned to the new ass. Phragmitetum altissimi, 4 subassociations, and 7 variants. The nomenclature type of a
最高芦苇(Phragmites altissimus)是一个分布在欧亚-北非的物种,最近在北部方向扩大了其分布区域(Kapitonova, 2016;Golovanov et al., 2019;Tzvelev, Probatova, 2019)。众所周知,在俄罗斯欧洲和亚洲部分的森林地带,最高的芦苇只被发现是一种入侵植物(Tzvelev, 2011)。在其自然分布范围内和入侵区域内,均已知以高山杨为主的群落。然而,在分类学综述中,具有这种种优势的植物通常被作者包括在类群中。Phragmitetum australis Savich 1926 (Golub et al., 1991,2015;Golub, Chorbadze, 1995;Kipriyanova, 2008;贝吉塔се…,2011;Golovanov, Abramova, 2012;Chepinoga, 2015)。本研究的目的是为了确定高山杨群落的分类学地位。这项工作使用了在P. altissimus的主要范围内(阿斯特拉罕地区和秋明地区南部的森林草原地区)和其次要范围内(乌德穆尔特共和国和秋明地区的针叶林地区)进行的65项地球植物学研究。在TURBOVEG程序(Hennekens, 1996)的基础上,将相关的样本导入数据库,并使用JUICE程序(Tichý, 2002)进行处理。为了评估所描述样地上的物种丰度,使用了J. Braun-Blanquet丰度量表,其丰度覆盖分数如下:r -物种与微不足道的报道极为罕见,+ -罕见的物种,覆盖的程度很小,1 -个人的数量大,覆盖面很小的程度或个人稀疏,但覆盖面很大,2 -个人的数量很大,投影覆盖从5到25%,3 -个人的数量,投影覆盖从25到50%,4 -个体数任意,投射覆盖度在50% ~ 75%之间,个体数任意,覆盖度在75%以上(Mirkin et al., 1989)。采用J. Braun-Blanquet(1964)提出的方法进行句法分析。语法群的名称根据“国际植物社会学命名法”(Theurillat et al., 2021)给出。根据“分层区系分类……”(Mucina et al., 2016)给出了高级句法系统。采用NMDS方法确定了影响群落分化和分布的主要因素。采用IBIS程序(Zverev, 2007),根据D. N. Tsyganov (Tsyganov, 1983)的生态尺度:土壤水分(Hd)、土壤氮丰富度(Nt)和光照-遮阳(Lc)计算各句法类的平均指标值。在PC-ORD v. 6.0中进行处理(McCune et Mefford, 2011)。所研究的群落归属于高原芦苇属(Phragmitetum altissimi)新科、4个亚类群和7个变异。关联的命名类型在表3中与n20相关。结果表明,高原芦苇(ass. Phragmitetum altissimi)群落的物种数为1 ~ 15种(平均4种),总投影盖度为20% ~ 100%。牧草的高度为2-5米;它分为四到五个子阶段。在第1亚期,除高原草外,还发现了大型香蒲(Typha austro-orientalis, T. linnaei, T. latifolia, T. tichomirovii),以及高草(Calamagrostis pseudoophragmites, Phalaroides arundinacea)和Scirpus hippolyti。第二亚期由中等高度(可达0.8-1米)的禾草组成,包括:金毛草、竖叶草、水草、水草、水草、水草、杜鹃花、桃根草、小叶草、鸢尾草,以及凤仙花、荨麻等。第三亚阶段并不总是发育,通常非常稀疏,由表面亲水的草(Rorippa amphibia, Galium palustre, Potentilla reptans, Tussilago farfara)形成,通常不超过10-20 (25)cm。第四亚期通常稀疏;它是由漂浮在水面上的有根水生植物(Nymphaea alba)或无根水生植物(Salvinia natans, lena minor, L. turionifera, Spirodela polyrhiza, Hydrocharis morsus-ranae)形成的。第五亚期由完全浸没在水中的非生根水生植物(Lemna trisulca, Ceratophyllum demersum),以及Drepanocladus aduncus和Cladophora sp.形成,通常是层外的藤本植物(Calystegia sepium, Cynanchum acutum);柳树(Salix cinerea, S. alba)的芽也很常见。以P. altissimus为主的群落形成于沿海浅水区,包括水位稳定或微波动的沼泽、停滞或弱流动水体,底部紧密或粉砂质碎屑沉积物层小,有时相当厚。 在潮湿或沼泽的海岸,包括受到干扰的、永久的或暂时干燥的水体上,也会形成群落。在人为生境中,它们发育在有水和潮湿的洼地(沟渠)、浅水和潮湿的池塘、水库、人造水体、挖掘和有水的采石场。这是一艘潜艇。典型水藻(图2)形成于海岸浅水和永久或暂时干燥的水体(包括受干扰的水体)的潮湿海岸。鲈鱼群落。滨水沙蚤主要生长于水位稳定或微波动的沼泽滨海地区和沼泽岸;它们的特点是由高山杨组成的牧草的上亚期相对较低且稀疏。鲈鱼群落。P. a. phalaroidetosum arundinaceae生长在沿海浅水(深5 ~ 10 cm)和水体湿润的岸边,其特点是牧草上亚期较密,生长季节基质暂时干燥。鲈鱼群落。三叶草是在水体中形成的,水体的水位在生长季节会有波动;高原草的群落分布在高原草的主要分布范围内——俄罗斯欧洲部分南部(阿斯特拉罕地区)和西伯利亚西部的森林草原地带(秋明地区)。它们也被发现在最高芦苇入侵的地区-在俄罗斯平原东部(乌德穆尔特共和国),在西伯利亚西部的针叶林地带(秋明地区)(图1)。在最高芦苇的次级范围内,只有被归因于亚鲈鱼的cenoses。记录了典型单胞菌。属于所有四个亚群的群落在俄罗斯欧洲部分领土上广泛分布于P. altissimus的主要范围内。
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The new associations of anthropogenic vegetation of the classes Sisymbrietea Gutte et Hilbig 1975 and Digitario sanguinalis–Eragrostietea minoris Mucina, Lososová et Šilc in Mucina et al. 2016 in the Republic of Bashkortostan Sisymbrietea Gutte et Hilbig 1975和Digitario sanguinalis-Eragrostietea minoris Mucina, lososov<e:1> et Šilc (Mucina et al. 2016)在巴什科尔托斯坦共和国的人为植被新关联
Q4 Agricultural and Biological Sciences Pub Date : 2022-01-01 DOI: 10.31111/vegrus/2022.44.61
Y. Golovanov, G. Biktimerova
Being one of the most characteristic types of plant communities found within settlements, the synanthropic communities of the Republic of Bashkortostan (RB) have been studied since the end of the XX century (Ishbirdin et al., 1988; Ishbirdinа, Anischenko, 1989; Sachapov et al., 1990; Anischenko, 1995; Ryabova, 1996, etc.). At the beginning of the XXI century geobotanical research was actively continued, while significant attention was paid to phytocenoses located within the towns and villages of the republic (Govorov, 2004; Edrenkina, 2005; Yamalov et al., 2008; Yamalov, Suyundukova, 2008; Usmanova et al., 2013, 2014). Urban vegetation is most fully presented in the series of publications on the classification of vegetation of towns of the Southern industrial zone (Golovanov, Abramova, 2012, 2013; Golovanov et al., 2017). The given article presents results of the long-term (2005-2018) research of vegetation of the towns of Bashkir Pre-Urals, as well as rural settlements of Trans-Urals. 63 complete relevés of new communities for RB included in the geobotanical base of synantropic vegetation of the Southern Urals were used. The geobotanical database was implemented using the TURBOVEG software (Hennekens, Schaminée, 2001). The area of relevé depended on the community size and homogeneity. Classification is carried out according to Braun-Blanquet method using the JUICE software (Tichý, 2002). The identification of new syntaxa was carried out in accordance with the “International Code of Phytosociological Nomenclature. 4th edition” (Theurillat et al., 2021). The names of the higher syntaxa are given by “Vegetation of Europe...” (Mucina et al., 2016). The ecological conditions were assessed with use of average values of E. Landolt (1977) ecological scales and processed by IBIS software (Zverev, 2007). Average values are calculated on the following scales: humidification (M), acidity (R), soil richness with mineral nutrition elements (N), humus (H) content, mechanical composition and soil structure (D), illumination (L). At the beginning of the 2010s prodrome of synanthropic communities of RB, included 7 classes, 12 orders, 17 alliances and 61 associations (Yamalov et al., 2012). Currently, the synthaxonomy of the synanthropic vegetation of the study area requires adjustment in accordance with the modern state of European synthaxonomy. Special geobotanical studies of recent years have allowed expanding the synthaxonomic scheme of synanthropic vegetation of the region. Earlier, data have already been published on new associations for the region of the classes Artemisietea vulgaris and Galio-Urticetea (Golovanov, 2017). This article extends our research. This article considers the synanthropic vegetation of two synanthropic classes — Sisymbrietea and Digitario sanguinalis–Eragrostietea minoris. These classes are first given for the territory of the RB. The article contains their regional diagnostic species. Previously, they were a part of the wider cla
作为聚落内发现的最具特征的植物群落类型之一,巴什科尔托斯坦共和国(RB)的共生群落自20世纪末以来一直受到研究(Ishbirdin et al., 1988;伊什伯丁纳,阿尼琴科,1989;Sachapov et al., 1990;Anischenko, 1995;Ryabova, 1996等)。21世纪初,地植物学研究得到了积极的继续,同时对共和国城镇和村庄内的植物群落给予了极大的关注(Govorov, 2004;Edrenkina, 2005;Yamalov et al., 2008;亚马洛夫,苏云杜科娃,2008;Usmanova et al., 2013,2014)。城市植被在南方工业区城镇植被分类系列出版物(Golovanov, Abramova, 2012, 2013;Golovanov et al., 2017)。给定的文章介绍了巴什基尔乌拉尔前城镇以及乌拉尔外农村定居点的长期(2005-2018)植被研究的结果。本文利用乌拉尔南部共向植被地植物学库中包含的63个RB新群落的完整记录。使用TURBOVEG软件实现了地球植物学数据库(Hennekens, schaminsamei, 2001)。相关区域取决于社区规模和同质性。采用JUICE软件(Tichý, 2002),根据布朗-布兰凯法进行分类。根据“国际植物社会学命名法第4版”(Theurillat et al., 2021)对新句法进行鉴定。高等句法群的名称由“欧洲植被”给出。(Mucina et al., 2016)。生态条件评价采用e.l Landolt(1977)生态尺度的平均值,采用IBIS软件(Zverev, 2007)进行处理。在以下尺度上计算平均值:湿度(M)、酸度(R)、土壤矿质营养元素丰富度(N)、腐殖质(H)含量、机械组成和土壤结构(D)、光照(L)。2010年代初,RB共生群落的前期包括7类、12目、17个联盟和61个协会(Yamalov et al., 2012)。目前,研究区共生植被的分类学需要根据欧洲分类学的现代状况进行调整。近年来的特别地植物学研究扩大了该地区共生植被的综合分类方案。早些时候,已经发表了关于青蒿类和Galio-Urticetea类在该地区的新关联的数据(Golovanov, 2017)。本文扩展了我们的研究。本文研究了两种共生植物类——西米草属(Sisymbrietea)和小红花属(Digitario sanguinalis-Eragrostietea)的共生植被。这些类首先是针对RB的领土给出的。本文包含了它们的区域诊断种。在此之前,它们属于更广泛的一类——星形植物,由一年生杂草和原始植被组成。对于茜草纲,给出了3种合纲:合纲是茜草属(Descurainietum sophiae)和蒲草属(Poo annuae-Stellarietum mediae)和亚纲。荨麻疹病。最后两种合胞是新的。它们将低生长一年生植物的亲硝基群落团结在一起,这些植物沿着房屋的墙壁、栅栏、肩膀和其他受干扰的栖息地分布在农村居民点和城镇。该协会将在各种受干扰的栖息地(垃圾填埋场、沉积物以及建筑物附近)发现的群落联合起来。它们主要发生在巴什基尔-乌拉尔地区的小定居点,很少发生在巴什基尔-乌拉尔地区。对RB和欧洲其他地区所描述的群落群落区系的比较(表2)表明,诊断种群的组成非常相似。最大的相似之处是位于捷克共和国的社区。Association Malvetum pusillae, subbass m.p. urticettosum urentis(表3). holotypus:表3,相关资料2,Bashkortostan共和国,Stepnoe Khaibullinsky区,沿着建筑物的墙壁,51.81988°N, 58.15081°E, 2005年6月14日,作者- G. Ya。Suyundukova。我们比较了在RB领土上描述的以Urtica urens为主的群落与东欧最密切的Malvion忽视联盟的群落(表4)。在几乎完全没有主要诊断种和群落区系特征上都可以看到显著的差异。在捷克共和国描述的cenoses的特点是高度稳定的Malva忽视,在乌拉尔南部完全没有,以及显著存在抗践踏的物种。这表明在比较的句法群中环境条件的性质不同。 作为聚落内发现的最具特征的植物群落类型之一,巴什科尔托斯坦共和国(RB)的共生群落自20世纪末以来一直受到研究(Ishbirdin et al., 1988;伊什伯丁纳,阿尼琴科,1989;Sachapov et al., 1990;Anischenko, 1995;Ryabova, 1996等)。21世纪初,地植物学研究得到了积极的继续,同时对共和国城镇和村庄内的植物群落给予了极大的关注(Govorov, 2004;Edrenkina, 2005;Yamalov et al., 2008;亚马洛夫,苏云杜科娃,2008;Usmanova et al., 2013,2014)。城市植被在南方工业区城镇植被分类系列出版物(Golovanov, Abramova, 2012, 2013;Golovanov et al., 2017)。给定的文章介绍了巴什基尔乌拉尔前城镇以及乌拉尔外农村定居点的长期(2005-2018)植被研究的结果。本文利用乌拉尔南部共向植被地植物学库中包含的63个RB新群落的完整记录。使用TURBOVEG软件实现了地球植物学数据库(Hennekens, schaminsamei, 2001)。相关区域取决于社区规模和同质性。采用JUICE软件(Tichý, 2002),根据布朗-布兰凯法进行分类。根据“国际植物社会学命名法第4版”(Theurillat et al., 2021)对新句法进行鉴定。高等句法群的名称由“欧洲植被”给出。(Mucina et al., 2016)。生态条件评价采用e.l Landolt(1977)生态尺度的平均值,采用IBIS软件(Zverev, 2007)进行处理。在以下尺度上计算平均值:湿度(M)、酸度(R)、土壤矿质营养元素丰富度(N)、腐殖质(H)含量、机械组成和土壤结构(D)、光照(L)。2010年代初,RB共生群落的前期包括7类、12目、17个联盟和61个协会(Yamalov et al., 2012)。目前,研究区共生植被的分类学需要根据欧洲分类学的现代状况进行调整。近年来的特别地植物学研究扩大了该地区共生植被的综合分类方案。早些时候,已经发表了关于青蒿类和Galio-Urticetea类在该地区的新关联的数据(Golovanov, 2017)。本文扩展了我们的研究。本文研究了两种共生植物类——西米草属(Sisymbrietea)和小红花属(Digitario sanguinalis-Eragrostietea)的共生植被。这些类首先是针对RB的领土给出的。本文包含了它们的区域诊断种。在此之前,它们属于更广泛的一类——星形植物,由一年生杂草和原始植被组成。对于茜草纲,给出了3种合纲:合纲是茜草属(Descurainietum sophiae)和蒲草属(Poo annuae-Stellarietum mediae)和亚
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引用次数: 0
Syntaxonomy of xeropetrophytic vegetation of Southern Urals: alliance Elytrigion pruiniferae all. nov. 乌拉尔南部旱生植被的分类学:鞘翅藓属。11月。
Q4 Agricultural and Biological Sciences Pub Date : 2022-01-01 DOI: 10.31111/vegrus/2022.43.88
A. Korolyuk, S. Yamalov, M. Lebedeva, Y. Golovanov, N. Dulepova, N. Zolotareva
The dominance of xeropetrophytic plant communities with original species composition along with high phytocenotic diversity is the first to determine the distinctness of the South Ural low-mountain steppe province within the Ural Mountains. As the zonal steppe ecosystems are totally destroyed and transformed, petrophytic cenoses represent the predominant type of natural vegetation. They are of high conservation importance due to preserving unique biodiversity with a large number of rare and endangered species, relics and endemics (Ryabinina, 2003; Yamalov et al., 2019). Stony steppes in the Southern Urals have been studied for more than 100 years. Initiated in the 1980s, the syntaxonomical research covers now only the limited territories, mainly within the protected areas (Unikalnye ..., 2014; Prirodnye ..., 2018). The recent studies are focused on the relationship between the petrophytic vegetation and macroecological gradients in the context of geographical zonation (Zolotareva et al., 2019; Korolyuk et al., 2020). Our study presents the results of the analysis of 265 relevés of petrophytic communities of the steppe zone, carried out by the authors on the territory of the Gaysky, Kuvandyksky, Saraktashsky and Belyaevsky administrative districts of the Orenburg region, as well as the Khaibullinsky district of the Bashkortostan Republic in 2014–2018 (Fig. 1). Six phytocoenons were identified using the TWINSPAN algorithm followed by manual sorting. They represent the main diversity of petrophytic vegetation in the studied region and differ in the groups of species identified by formal criteria (Korolyuk et al., 2016; Zolotareva et al., 2019). We conducted the cluster analysis to compare these with the petrophytic associations of the Urals: 26 associations from the classes Festuco-Brometea Br.-Bl. et Tx. ex Soó 1947 and Anabasietea cretaceae Golovanov et al. 2021 (Fig. 2, 3). Five communities were assigned to the steppe vegetation, and their position in the syntaxonomical system was determined. Communities 1–3 regarded as typical steppes have similar floristic composition and dominant plants; community 4 is closer to the rich forb-bunchgrass steppes; community 5 and the ass. Anthemido trotzkianae–Thymetum guberlinensis have transitional features to desert steppes. We identified the new alliance Elytrigion pruiniferae all. nov. D. s.: Allium tulipifolium, Alyssum turkestanicum, Artemisia salsoloides, Centaurea turgaica, Elytrigia pruinifera, Ferula tatarica, Linaria uralensis, Poa bulbosa s. l., Polycnemum arvense, Spiraea hypericifolia. Holotypus — the association Elytrigietum pruiniferae Lebedeva ass. nov. Alliance represents xeropetrophytic vegetation of the steppe zone of the South Urals. Its area covers the watersheds of the Sakmara, Ural and Ilek rivers. Its communities were also found on the right bank of the Sakmara river (the Guberlya and Kuragan basins), as well as in the lower part of the Bolshoy Ik river basin. The communities occupy
具有原始物种组成和高度植物多样性的旱生植物群落优势是乌拉尔山脉内南乌拉尔低山草原省独特性的第一个决定因素。随着地带性草原生态系统的全面破坏和改造,岩生苔藓是天然植被的主要类型。由于保存了大量稀有和濒危物种、遗迹和特有物种的独特生物多样性,它们具有很高的保护重要性(Ryabinina, 2003;Yamalov et al., 2019)。人们对乌拉尔南部的石质草原进行了100多年的研究。分类学研究始于20世纪80年代,目前仅覆盖有限的领土,主要是在保护区内(尤卡尼耶…, 2014;Prirodnye……, 2018)。最近的研究主要集中在地理区划背景下岩石植被与宏观生态梯度的关系(Zolotareva et al., 2019;Korolyuk et al., 2020)。我们的研究展示了作者在2014-2018年在奥伦堡地区的Gaysky、Kuvandyksky、Saraktashsky和Belyaevsky行政区以及巴什科尔托斯坦共和国的Khaibullinsky地区对草原带岩石植物群落的265个相关数据进行分析的结果(图1)。使用TWINSPAN算法识别了6个植物群落,然后进行了人工分类。它们代表了研究区域岩石植物植被的主要多样性,并且根据正式标准确定的物种群有所不同(Korolyuk等人,2016;Zolotareva et al., 2019)。通过聚类分析,将这些组合与乌拉尔地区的岩生组合进行了比较:Festuco-Brometea Br.-Bl类中有26个组合。et txt . ex Soó 1947和Anabasietea白垩纪Golovanov et al. 2021(图2,3)。将5个群落划分为草原植被,并确定其在分系统中的位置。典型草原群落1 ~ 3具有相似的区系组成和优势植物;4号社区更靠近茂密的束草草原;群落5和牛头草-百里香草具有向荒漠草原过渡的特征。我们确定了新联盟Elytrigion priiniferae all。11月5日:郁金香葱、土耳其花、黄花蒿、半人马草、黄花蒿、乌拉尔阿魏、乌拉尔丽娜、金盏花、金盏花、绣线菊。Holotypus - Elytrigietum pruiniferae Lebedeva ass11 . Alliance代表了南乌拉尔草原地带的旱生植被。它的区域覆盖了萨克马拉河、乌拉尔河和伊莱克河的流域。在Sakmara河的右岸(Guberlya和Kuragan流域)以及Bolshoy Ik河流域的下游也发现了它的群落。这些群落位于凹凸的斜坡和未成熟砾石土的山脊顶部。在联盟分布区的南部,它们被限制在各种形状和暴露的斜坡上,而在北部,它们主要是凸的南斜坡或山脊的顶部。该联盟代表了枯毛目的旱生部分。在水分梯度上,它取代了更多的中生联盟——荒漠Helictotricho - orostachyion spinosae (Korolyuk, 2017)。联盟包括3个协会和1个社区类型。(表4),盲蝽1型(田号:1)。15-086):库万德克地区奥伦堡地区,Yalnair村西南1.5公里处,岩石斜坡的近峰顶凸起部分,51.28635°N, 57.81782°E, 2015年6月10日,作者- a . Yu。Korolyuk。主要品种:土耳其香、土耳其半马菊、乌拉尔花、乌拉尔花、金莲花、金莲花、绣线菊、百里香、郁金香。该协会将奥伦堡地区的萨拉克塔什、库万代克和盖斯基地区的丘陵地带和低山的岩石植物草原以及巴什基里亚的哈布林斯基地区的图拉特卡山上的岩石植物草原联合起来。这种植物局限于凸坡和山脊的顶部。它们发育在基性岩和变质岩露头的砾石土壤上,较少发育在沉积岩上。在关联中有4个变体。百里香(Thymo guberlinensis-Galatelletum villosae Yamalov), 11 .(表5),holotypus - rel. 2(田号)。16-015):库万德克地区奥伦堡地区,扎鲁泽村西南19公里,北纬51.18268°,东经56.91858°,作者- A. Yu。Korolyuk。研究对象:黄芪、麻黄、青花蒿、黄芪、黄芪、黄芪、黄芪、黄芪、黄芪。该协会代表旱生石质草原。它们主要发生在南坡和沉积岩上有细砾质土壤的山脊顶部的所有地区。 具有原始物种组成和高度植物多样性的旱生植物群落优势是乌拉尔山脉内南乌拉尔低山草原省独特性的第一个决定因素。随着地带性草原生态系统的全面破坏和改造,岩生苔藓是天然植被的主要类型。由于保存了大量稀有和濒危物种、遗迹和特有物种的独特生物多样性,它们具有很高的保护重要性(Ryabinina, 2003;Yamalov et al., 2019)。人们对乌拉尔南部的石质草原进行了100多年的研究。分类学研究始于20世纪80年代,目前仅覆盖有限的领土,主要是在保护区内(尤卡尼耶…, 2014;Prirodnye……, 2018)。最近的研究主要集中在地理区划背景下岩石植被与宏观生态梯度的关系(Zolotareva et al., 2019;Korolyuk et al., 2020)。我们的研究展示了作者在2014-2018年在奥伦堡地区的Gaysky、Kuvandyksky、Saraktashsky和Belyaevsky行政区以及巴什科尔托斯坦共和国的Khaibullinsky地区对草原带岩石植物群落的265个相关数据进行分析的结果(图1)。使用TWINSPAN算法识别了6个植物群落,然后进行了人工分类。它们代表了研究区域岩石植物植被的主要多样性,并且根据正式标准确定的物种群有所不同(Korolyuk等人,2016;Zolotareva et al., 2019)。通过聚类分析,将这些组合与乌拉尔地区的岩生组合进行了比较:Festuco-Brometea Br.-Bl类中有26个组合。et txt . ex Soó 1947和Anabasietea白垩纪Golovanov et al. 2021(图2,3)。将5个群落划分为草原植被,并确定其在分系统中的位置。典型草原群落1 ~ 3具有相似的区系组成和优势植物;4号社区更靠近茂密的束草草原;群落5和牛头草-百里香草具有向荒漠草原过渡的特征。我们确定了新联盟Elytrigion priiniferae all。11月5日:郁金香葱、土耳其花、黄花蒿、半人马草、黄花蒿、乌拉尔阿魏、乌拉尔丽娜、金盏花、金盏花、绣线菊。Holotypus - Elytrigietum pruiniferae Lebedeva ass11 . Alliance代表了南乌拉尔草原地带的旱生植被。它的区域覆盖了萨克马拉河、乌拉尔河和伊莱克河的流域。在Sakmara河的右岸(Guberlya和Kuragan流域)以及Bolshoy Ik河流域的下游也发现了它的群落。这些群落位于凹凸的斜坡和未成熟砾石土的山脊顶部。在联盟分布区的南部,它们被限制在各种形状和暴露的斜坡上,而在北部,它们主要是凸的南斜坡或山脊的顶部。该联盟代表了枯毛目的旱生部分。在水分梯度上,它取代了更多的中生联盟——荒漠Helictotricho - orostachyion spinosae (Korolyuk, 2017)。联盟包
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引用次数: 1
International Code of Phytosociological Nomenclature. 4th ed. 国际植物社会学命名规则。第4版。
Q4 Agricultural and Biological Sciences Pub Date : 2022-01-01 DOI: 10.31111/vegrus/2005.07.3
J. Theurillat, W. Willner, F. Fernández‐González, H. Bültmann, A. Čarni, D. Gigante, L. Mucina, H. Weber
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引用次数: 9
Land typology by L. G. Ramenskiy and eunis habitat classification (retrospective view) L. G. Ramenskiy的土地类型学与生境分类(回顾)
Q4 Agricultural and Biological Sciences Pub Date : 2021-12-01 DOI: 10.31111/vegrus/2021.41.150
V. Golub
The rapid rate of decline in the Earth’s biodiversity under the influence of direct and indirect anthropogenic pressure makes it necessary to develop the scientific foundations for its conservation at all levels of life. Ecologists have come to understand that the best way to ensure the conservation of populations of organisms and their communities is to preserve the environment in which they live. The countries of the European Community, where special programs have been developed since mid 1980s, have shown the greatest activity in preserving environmental conditions. Currently, the «European Union Nature Information System» (EUNIS) has become the most popular among such programs. Habitat is a central concept in EUNIS. For the purposes of EUNIS, habitat is defined asa place where plants or animals normally live, characterized primarily by its physical features (topography, plant or animal physiognomy, soil characteristics, climate, water quality etc.) and secondarily by the species of plants and animals that live there (Davies et al., 2004). Most often, habitat is considered to be synonym of the term biotope. The EUNIS biotope classification would correspond to the ecosystem classification if heterotrophic components were largely present in it. However, at present, these organisms, are not used for classification of terrestrial ecosystems. The latter (especially benthos) are important in the characterization of marine habitat types.The author does not deny the extreme importance of the EUNIS habitat classification for ecological science and solving problems of nature conservation. He is only sure that the concept of habitat classification began to be developed in the Soviet Union as early as 1920–1930th in the papers by L. G. Ramenskiy who in 1927 published the definition of habitat type: The type of habitat or natural area is determined by a combination of climate conditions, relief, irrigation, and the nature of the soil and subsoil. The same type can be covered by a meadow, or a forest, or plowed up, etc.: these are its transitional states (in virgin untouched nature, each type is inhabited by a completely definite combination of plants - steppe, forest, meadow, etc.). Afterwards L. G. Ramenskiy began to use the term land type instead of habitat type.In the 1930s, by the land type he meant an ecosystem unit in which plant community would exist without human influence. The land type in nature is represented by a set of various modifications that arise, as a rule, under man pressure. Modifications can transform into each other and revert to the original state of the type. Later, such plant community was called potential vegetation (Tüxen, 1956). In 1932–1935, L. G. Ramenskiy supervised the inventory of natural forage lands in the USSR, which used this concept of land type (Golub, 2015). The inventory of natural forage lands in the USSR resulted in their hierarchical classification: 19 classes and 43 subclasses were established. The exact numb
在直接和间接的人为压力的影响下,地球的生物多样性迅速减少,因此有必要为在生命的各个层次上保护生物多样性发展科学基础。生态学家已经认识到,保护生物种群及其群落的最好方法是保护它们生存的环境。欧共体国家自1980年代中期以来制定了特别方案,在保护环境条件方面表现出最大的活动。目前,“欧盟自然信息系统”(EUNIS)已成为这些项目中最受欢迎的。生境是EUNIS的核心概念。就EUNIS而言,栖息地被定义为植物或动物正常生活的地方,其主要特征是其物理特征(地形、动植物地貌、土壤特征、气候、水质等),其次是生活在那里的动植物物种(Davies et al., 2004)。通常,生境被认为是生物圈的同义词。如果异养成分大量存在,则EUNIS的生物群落分类将与生态系统分类相对应。然而,目前,这些生物还没有被用于陆地生态系统的分类。后者(尤其是底栖动物)在海洋生境类型的表征中很重要。作者不否认EUNIS生境分类对生态科学和解决自然保护问题的极端重要性。他只确信,栖息地分类的概念早在1920 - 1930年代就在苏联的L. G. Ramenskiy的论文中开始发展起来,Ramenskiy在1927年发表了栖息地类型的定义:栖息地或自然区域的类型是由气候条件、地形、灌溉以及土壤和底土的性质综合决定的。同一类型可以被草甸、森林或耕地覆盖,等等:这些是它的过渡状态(在原始的未受破坏的自然中,每种类型都有完全确定的植物组合——草原、森林、草甸等)。后来L. G. Ramenskiy开始使用土地类型这个术语来代替栖息地类型。在20世纪30年代,他所说的土地类型指的是一种生态系统单元,在这种生态系统单元中,植物群落可以在没有人类影响的情况下生存。自然界中的土地类型通常由一系列在人类压力下出现的各种变化来表示。修改可以相互转换,并恢复到类型的原始状态。后来,这种植物群落被称为潜在植被(txen, 1956)。1932-1935年,L. G. Ramenskiy监督了苏联天然牧草地的清查,其中使用了这种土地类型的概念(Golub, 2015)。通过对苏联天然牧草地的调查,将其划分为19个类和43个亚类。区分类型的确切数量没有计算,根据拉门斯基的粗略估计,有一千多种。在大多数情况下,这些类型的潜在植被无法被识别。这次清查的记录没有公布。然而,L. G. Ramenskiy前研究生N. V. Kuksin,他参加了乌克兰的调查,写了一本关于苏联共和国牧草类型土地的书(Kuksin, 1935)。这本书中提出的干草和牧场的类型学与根据EUNIS系统原则开发的栖息地分类非常相似(Kuzemko等人,2018)。到20世纪40年代末,拉曼斯基得出结论,现代科学无法为许多栖息地类型建立潜在的植被。因此,他建议将他之前归因于修改的土地类型称为土地类型。出于实际原因和简洁的考虑,也可以将土地类型(森林、草甸、可耕地)的主要修改组称为类型(Ramenskiy, 1950, p. 489)。因此,他对土地类型的理解与后来在EUNIS系统中对生境的解释是一致的。Ramenskiy土地类型学与EUNIS生境分类具有相同的本质和依据,但人类社会的不同群体提出了前者,后者试图保护土地资源。基于Ramenskiy类型学,对生物群落的利用提出了建议,以获得可持续的最大经济产量。根据EUNIS系统的分类,就保护植物和动物种群及其特定生物群落的特征提出了建议。如果社会有这样的需求,拉曼斯基的土地类型学可以很好地用于保护生物群落。在拉门斯基的一生中,从未有过像现在这样对自然保护如此浓厚的兴趣。 目前,前苏联境内已开始使用EUNIS生物类群分类,而拉曼斯基的土地类型学已被遗忘。造成这种现象的原因有两个:1)苏联科学的孤立主义,使国内科学家与西方同行分离;拉门斯基的思想太超前了,其深度、本质和重要性直到几十年后才为生物学家所理解。拉门斯基关于生境类型学观点的形成可能受到俄罗斯森林科学家克鲁德纳思想的影响。
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引用次数: 0
Vegetation of chalk outcrops of Sub-Ural plateau and adjacent territories 乌拉尔高原及其邻近地区白垩露头植被
Q4 Agricultural and Biological Sciences Pub Date : 2021-01-01 DOI: 10.31111/vegrus/2021.40.3
Y. Golovanov, S. Yamalov, M. Lebedeva, A. Korolyuk, L. Abramova, N. Dulepova
The results of long-term studies of the vegetation of chalk outcrops of the Orenburg region (Russian Federation) and North-West Kazakhstan on Sub-Ural plateau and adjacent territories are presented. Chalk outcrops are unique botanical-geographical sites located in steppe and desert zones of Eurasia. Specific communities of calcephyte plant species have spread in these areas, in places of outcrops or close occurrence from the surface of upper-Cretaceous carbonate rocks. The flora of chalk outcrops is characterized by a great amount of rare species, mainly ende­mic, associated with peculiar substrates, the locality of habitats, and the historical past of the area of outcrops location (Matyshenko, 1985) The history of the study of flora and vegetation of chalk outcrops is given. Synthaxonomic studies of chalk vegetation as part of the ecological-floristic approach cover only territories west of the Volga river (Poluyanov, 2009; Averinova, 2011, 2016; Demina, 2014; Demina et al., 2016; Didukh et al., 2018). Chalk highlands of the North-West Kazakhstan and adjacent regions of the Russian Federation occupy quite large areas. However, up to date, there is no data on the vegetation diversity of these territories based on complete geobotanical relevés, that is why their synthaxonomy remains undeveloped. The study area with 15 massifs of chalk outcrops (Fig. 1) includes the Orenburg region (Novosergievsky, Perevolotsky, Sol-Iletskiy, Akbulak and Gaysky districts), and Aktobe (Hobdinsky, Uilsky and Bayganinsky district) and Atyrau (Zhylyoysky district) regions of the Republic of Kazakhstan. The largest massifs in the Orenburg region of the Russian Federation are: Starobelogorskie (Fig. 2), Chesnokov­skie (Fig. 3), Verkhnechibendinskie (Fig. 6), Troits­kie (Fig. 7), Pokrovskie Chalk Mountains (Fig. 4) and Durtel mountain (Fig. 5). Chalk massif Akshatau (Fig. 8) and the range Aktolagai (Fig. 9) are the largest within Aktobe region. The investigated sites are mostly located on the Sub-Ural Plateau, which extended from the southern regions of the Orenburg region to the Emba River in the territory of Aktobe region. They are less common within the Obschiy Syrt and sporadic in the Guberlinskie mountains. The study area covers a wide range of zonal vegetation from dry steppes in the northern part of the gradient to northern deserts in the southern one. The dataset includes 270 relevés of chalk outcrops communities performed by the authors in 2014–2019. The primary classification was carried out using TWINSPAN algorithm. As a result three groups of communities are established. The first group is communities of the Emben Plateau, the most southern area; second is communities on relatively developed soils in the slopes bases, depressions between chalk ridges and on their flat tops; third is widespread communities on most of the Podural Plateau and Obschy Syrt, excluding the Emben Plateau. Comparison with associations of calcephyte, semidesert and steppe vegetation
本文介绍了对奥伦堡地区(俄罗斯联邦)和哈萨克斯坦西北部乌拉尔高原及其邻近地区白垩露头植被的长期研究结果。白垩露头是位于欧亚大陆草原和沙漠地带的独特的植物地理遗址。在这些地区,在上白垩统碳酸盐岩表面的露头或近产地,分布着特定的钙生植物群落。白垩露头植物区系的特点是有大量的稀有物种,主要是终末种,与特殊的底物、生境的位置和露头所在地区的历史有关(Matyshenko, 1985)。作为生态-植物区系研究的一部分,白垩植被的分类学研究只覆盖了伏尔加河以西的地区(Poluyanov, 2009;Averinova, 2011, 2016;Demina, 2014;Demina et al., 2016;Didukh et al., 2018)。哈萨克斯坦西北部的白垩高地和俄罗斯联邦的邻近地区占据了相当大的面积。然而,到目前为止,还没有根据完整的地学资料汇编的关于这些领土的植被多样性的数据,这就是为什么它们的分类学仍然没有发展的原因。研究区有15块白垩露头(图1),包括哈萨克斯坦共和国的Orenburg地区(Novosergievsky、Perevolotsky、Sol-Iletskiy、Akbulak和Gaysky地区)、Aktobe地区(Hobdinsky、Uilsky和Bayganinsky地区)和Atyrau地区(Zhylyoysky地区)。俄罗斯联邦奥伦堡地区最大的山体是:Starobelogorskie(图2)、Chesnokov-skie(图3)、verkhnechibendinsky(图6)、troitskie(图7)、pokrovsky白垩山(图4)和Durtel山(图5)。白垩山Akshatau(图8)和Aktolagai山脉(图9)是阿克托别地区最大的山体。调查的地点大多位于乌拉尔高原以南,从奥伦堡地区的南部地区延伸到阿克托别地区的伊巴河。它们在叙利亚境内不太常见,在古伯林斯基山脉零星分布。研究区地带性植被分布广泛,从北部的干草原到南部的北部沙漠。该数据集包括作者在2014-2019年对白垩露头群落进行的270次相关采样。采用TWINSPAN算法进行初步分类。因此,建立了三个社区群体。第一组是最南部恩本高原的群落;二是在相对发达的斜坡基部、白垩脊之间的洼地及其平顶上的群落;三是除恩本高原外,豆瓣高原和叙利亚北部大部分地区都有广泛的群落。与钙质植物、半沙漠和草原植被关联的比较(Golub, 1994;科洛米丘克,维诺库罗夫,2016;Lysenko, Yamalov, 2017;Didukh等人,2018;Korolyuk, 2017),以确定所研究的群落在欧亚大陆草本植被生态-区系分类系统中的位置。聚类分析结果(图10)显示,与东欧的钙化群落以及沙漠和草原地带性植被相比,亚乌拉尔高原的白垩露头具有显著的特异性。这就是为研究的白垩露头植被描述一个新类别的原因。白垩纪Anabasietea白垩纪Golovanov类11 . hoc loco。诊断种:白垩木、黄杨、黄花蒿、白垩木、白垩木、吉林菊树、白垩木、黑桫椤、白垩木、白桫椤、白桫椤、白桫椤、白桫椤、白桫椤、白桫椤、白桫椤、白桫椤、白桫椤、白桫椤;类结合了奥伦堡地区南部和哈萨克斯坦西北部的白垩和泥灰岩露头上的钙化,主要是半灌木群落,位于草原(真正的草原和沙漠草原的亚带)和沙漠地带。描述了中心目Anabasietalia白垩纪Golovanov ord. nov. hoc loco;holotypus是Anthemido trotzkianae-Artemision salsoloidis all联盟。11月特别的火线。群落内的3个联盟反映了群落沿纬度梯度的分布和演替阶段。Sileno fruticulose - nanophytion erinacei Lebedeva all联盟。特别生境是物种贫乏的群落,主要分布在乌拉尔高原南部(恩本高原)和邻近地区的白垩地块上。该联盟的Holotypus为ass. Onosmo staminei-Anabasietum白垩纪ass. nov。 本文介绍了对奥伦堡地区(俄罗斯联邦)和哈萨克斯坦西北部乌拉尔高原及其邻近地区白垩露头植被的长期研究结果。白垩露头是位于欧亚大陆草原和沙漠地带的独特的植物地理遗址。在这些地区,在上白垩统碳酸盐岩表面的露头或近产地,分布着特定的钙生植物群落。白垩露头植物区系的特点是有大量的稀有物种,主要是终末种,与特殊的底物、生境的位置和露头所在地区的历史有关(Matyshenko, 1985)。作为生态-植物区系研究的一部分,白垩植被的分类学研究只覆盖了伏尔加河以西的地区(Poluyanov, 2009;Averinova, 2011, 2016;Demina, 2014;Demina et al., 2016;Didukh et al., 2018)。哈萨克斯坦西北部的白垩高地和俄罗斯联邦的邻近地区占据了相当大的面积。然而,到目前为止,还没有根据完整的地学资料汇编的关于这些领土的植被多样性的数据,这就是为什么它们的分类学仍然没有发展的原因。研究区有15块白垩露头(图1),包括哈萨克斯坦共和国的Orenburg地区(Novosergievsky、Perevolotsky、Sol-Iletskiy、Akbulak和Gaysky地区)、Aktobe地区(Hobdinsky、Uilsky和Bayganinsky地区)和Atyrau地区(Zhylyoysky地区)。俄罗斯联邦奥伦堡地区最大的山体是:Starobelogorskie(图2)、Chesnokov-skie(图3)、verkhnechibendinsky(图6)、troitskie(图7)、pokrovsky白垩山(图4)和Durtel山(图5)。白垩山Akshatau(图8)和Aktolagai山脉(图9)是阿克托别地区最大的山体。调查的地点大多位于乌拉尔高原以南,从奥伦堡地区的南部地区延伸到阿克托别地区的伊巴河。它们在叙利亚境内不太常见,在古伯林斯基山脉零星分布。研究区地带性植被分布广泛,从北部的干草原到南部的北部沙漠。该数据集包括作者在2014-2019年对白垩露头群落进行的270次相关采样。采用TWINSPAN算法进行初步分类。因此,建立了三个社区群体。第一组是最南部恩本高原的群落;二是在相对发达的斜坡基部、白垩脊之间的洼地及其平顶上的群落;三是除恩本高原外,豆瓣高原和叙利亚北部大部分地区都有广泛的群落。与钙质植物、半沙漠和草原植被关联的比较(Golub, 1994;科洛米丘克,维诺库罗夫,2016;Lysenko, Yamalov, 2017;Didukh等人,2018;Korolyuk, 2017),以确定所研究的群落在欧亚大陆草本植被生态-区系分类系统中的位置。聚类分析结果(图10)显示,与东欧的钙化群落以及沙漠和草原地带性植被相比,亚乌拉尔高原的白垩露头具有显著的特异性。这就是为研究的白垩露头植被描述一个新类别的原因。白垩纪Anabasietea白垩纪Golovanov类11 . hoc loco。诊断种:白垩木、黄杨、黄花蒿、白垩木、白垩木、吉林菊树、白垩木、黑桫椤、白垩木、白桫椤、白桫椤、白桫椤
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引用次数: 4
Diversity of larch forests and woodlands of the western macroslope of the Subpolar and Northern Urals 亚极地西部和北乌拉尔大斜坡落叶松林和林地的多样性
Q4 Agricultural and Biological Sciences Pub Date : 2021-01-01 DOI: 10.31111/vegrus/2021.41.3
S. Degteva, Y. Dubrovskiy
The study of the diversity of plant species and communities on several mountain ridges of Subpolar and Northern Urals (Fig. 1) in the basins of the rivers Kozhym, Kosyu, Bolshaya Synya, Vangyr, Schugor, and Ilych was carried out in 2007–2018 by researches of the Institute of Biology of Federal Research Centre “Komi Science Centre Ural Branch Russian Academy of Sciences”. Special attention was paid to fir (Abies sibirica) forests as well as larch (Larix sibirica) forests and woodlands due to the luck of data on their diversity. The study following both traditional (Polevaya…, 1964) and modern (Ipatov, Mirin, 2008) approaches of geobotanical and floristic researches is based on 168 original relevés (on sample plots of 400 m2 or within the community limits). The geobotanical data set which contains 184 relevés is stored in the archive (phytocoenarium) of the Institute of Biology (above). The community vertical and horizontal structure as well as the diversity and abundance of vascular plants, bryophytes, and lichens were under study. Ecological-phytocoenotical (dominant) approach was used for classification of larch forests and woodlands in the study area using both the author’s and literature data (Yudin, 1954; Gorchakovskiy, 1966; Nepomilueva, 1974; Martynenko, 1999; Neshataeva, Neshataev, 2005; Rysin, 2010; Kucherov, 2019). Larch forests and woodlands of the study area belong to the Montano-Lariceta subformation of the Lariceta sibiricae formation, which belongs to the Therhodendrosa vegetation subtype of the Aciculilignosa vegetation type (Bykov, 1960). The list of syntaxa for subformation Montano-Lariceta (M.-L.) includes 20 associations, 2 subassociations, 23 variants, and 3 community types from 5 forest types — lichen, green moss, herbaceous, hair cap moss and sphagnum ones. Three associations are transitional between various types of forest (Table 2–6, Fig. 2–10). Forest types of this subformation are allocated in different ordination areas of ecological space according to vectors of soil nitrogen content and light (Table 7, Fig. 11). Larch forests and woodlands of type M.-L. cladinosa occur in dry habitats with poor acidic soils, while phytocenoses of M.-L. polytrichosa and M.-L. sphagnosa are common on wet poor and acidic soils and those of M.-L. hylocomiosa on more fertile soils. In the study area, they do not occupy large areas. The communities of type M.-L. herbosa are common in low elevated mesophyte habitats with more fertile soils. Communities of M.-L. hylocomiosa and M.-L. herbosa types widely occur both in mountain forest and woodland altitudinal belts at the western macroslope of the Subpolar Urals northward N 64°. The use of environmental scales and statistical methods to identify classification units of lower rank did not give well-interpreted results. Analysis of the cenotic significance of species in various forest types of Montano-Lariceta revealed the stable and compact “core” of the most frequent species: trees Betula pubes
联邦研究中心生物研究所“科米科学中心-俄罗斯科学院乌拉尔分院”于2007-2008年对Kozhym河、Kosyu河、Bolshaya Synya河、Vangyr河、Schugor河和Ilych河流域的亚极地和北乌拉尔山脉(图1)的植物物种和群落多样性进行了研究。由于其多样性数据的运气,人们特别关注冷杉林、落叶松林和林地。这项研究遵循了传统(Polevaya…,1964)和现代(Ipatov,Mirin,2008)的地植物学和区系研究方法,基于168 原始相关(在400平方米的样地上或在社区范围内)。地球植物学数据集包含184个相关信息,存储在生物研究所的档案(植物群落)中(上图)。群落的垂直和水平结构以及维管植物、苔藓植物和地衣的多样性和丰度正在进行研究。利用作者和文献数据,采用生态植物群落(优势)方法对研究区域的落叶松森林和林地进行分类(Yudin,1954;Gorchakovskiy,1966;Nepomileuva,1974;Martynenko,1999;Neshataeva,Neshataev,2005;Rysin,2010;Kucherov,2019)。研究区的落叶松森林和林地属于西伯利亚落叶松组的Montano落叶松亚组,属于Aciculinsa植被类型的Theropendrosa植被亚型(Bykov,1960)。Montano Lariceta(M.-L.)亚组的句法结构列表包括20个组合、2个亚组合、23个变体和3个群落类型,它们来自5种森林类型——地衣、绿苔藓、草本、毛帽苔藓和泥炭藓。三种关联是不同类型森林之间的过渡(表2-6,图2-10)。根据土壤含氮量和光照的载体,该亚组的森林类型被分配在生态空间的不同排序区域(表7,图11)。M.-L.枝状落叶松的落叶松林和林地分布在酸性土壤较差的干燥栖息地,而多毛M.-L.和球形M.-L.在潮湿的贫瘠和酸性土壤上常见,而hylocomiosa在较肥沃的土壤上常见。在研究区域,它们并不占据很大的面积。草本M.-L.群落常见于土壤肥沃的低海拔中生植物栖息地。M.-L.hylocomiosa和M.-L.herbosa类型的群落广泛分布在北纬64°以北的亚极地乌拉尔山脉西部宏观斜坡的山林和林地海拔带。使用环境尺度和统计方法来识别较低级别的分类单元并没有得到很好的解释结果。对Montano Lariceta不同森林类型中物种的群落意义的分析揭示了最常见物种的稳定和紧凑的“核心”:乔木Betula pubescens、Larix sibirica、Picea obovata、Sorbus sibirica,灌木Betula nana和Juniperus sibirica;矮生灌木越橘属(Vaccinium myrtillus)、紫丁香属(V.uliginosum)、葡萄属(V.vitis idaea);草本Avenella flexuosa,Bistorta major,Calamagrostis purpurea,Carex arctisibrica,Empetrum hermachroditum,Festuca ovina,Trientalis europaea;苔藓Dicranum flexicaule、D.scoparium、Pleurozium schreberi、Polytrichum community(表2、8、9)。分枝杆菌森林类型的特征种是地衣Cladonia arbuscula、C.stellaris、Flavocetraria nivalis,差异种是草本Silene acaulis、S.pauciflora、Tephroseris residifolia和地衣Alectoria nigricans、Ashinea chrysantha、Bryocoulon diversions、Cetraria nigricans。最丰富的是矮灌木Arctous alpina、Empetrum hermachroditum、Dryas octopala、Ledum decumbens、Vaccinium uliginosum,以及草本植物Carex arctisibrica和C.globilaris。苔藓地衣层主要为岛状Cetraria islandica、丛枝Cladonia arbuscula、兰枝C.rangiferina、星状C.stellaris、雪花Flavocetraria nivalis和Stereocaulon paschale;同样丰富的还有地衣Cladonia gracilis和C.uncialis,以及苔藓Dicranum flexicaule和Pleurozium schreberi。M.-L.hylocomiosa的特征种是Avenella flexuosa,差异种是地衣Cetrariella delisei、C.laevigata、Cladonia subfurcata、Cladona crispata以及苔藓Dicranum polystum和Polytrichum piliferinum;恒定且高度丰富的物种有弯曲Avenella flexuosa、南桦Betula nana、两性Empetrum hermachroditum、越橘Vaccinium myrtillus、V.uliginosum和苔藓Pleurozium schreberi;具有平均恒常性的物种——越橘属、地衣Cladonia arbuscula、C.rangiferina和苔藓Dicranum flexicaule、Hylocomeum splines、Polytrichum community。有时矮生灌木Ledum decumbens和Rubus arcticus,以及苔藓植物Barbilophozia hatcheri、B.lycopdioides、Dicranum polysitem、D.scoparium和Sphagnum angustifolium占主导地位或共同占主导地位(表8)。
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引用次数: 0
Scientific workshop «Theoretical problems of the classification of the plant communities of Russia» (Yalta, November 1–3, 2021) 科学研讨会«俄罗斯植物群落分类的理论问题»(雅尔塔,2021年11月1日至3日)
Q4 Agricultural and Biological Sciences Pub Date : 2021-01-01 DOI: 10.31111/vegrus/2021.42.165
L. Kipriyanova, Y. Plugatar, N. V. Matveyeva, N. Ermakov
Thirty years after the publication of the USSR vegetation syntaxa prodromus (Korotkov et al., 1991), the active work on the preparation of the Prodromus of the Russian Federation vegetation (Plugatar et al., 2020) has started. 29 contributions (incl. 5 via Skype), were represented at the Workshop “Theoretical problems of the classification of plant communities of Russia”, that took place in Yalta, Republic of Crimea, on November 1–3, 2021. Twenty six scientists (Fig. 1, 2) from St. Petersburg, Moscow, Novosibirsk, Ufa, Vladivostok, Yakutsk, Syktyvkar, Bryansk, Rostov on Don, Khanty-Mansiysk, Ekaterinburg, Nalchik, Kursk, Yalta) participated this meeting. The reviews of syntaxonomic systems of forests, steppes, meadows, and aquatic, halophytic, synantropic, alpine, arctic, petrophytic, see-shore vegetation and wetlands were represented. The problems, that have appeared after beginning the syntaxa checklists development, such as the validation of units, the dubiousness of some accepted syntaxonomic decisions, poor syntaxonomic knowledge of some regions were discussed. The directors of three Botanical Gardens (from Yalta, Novosibirsk and Vladivostok) of the Russian Academy of Sciences signed documents (Fig. 3) on the starting of a scientific consortium «National Network of Botanical Gardens of the Russian Federation», focused on joint fundamental researches in the vegetation science, ecology, bioclimatology and biogeography; on the development of the National classification of vegetation and National typology of habitats as a scientific basis for assessing the carbon balance, rational use of natural resources and vegetation conservation in the country. Resolution of the Workshop emphasizes the necessity to consider the development of the Russian national vegetation classification as a priority task for phytosociologists in Russian Federation for the next five years.
《苏联植被句法》(Korotkov et al., 1991)出版三十年后,《俄罗斯联邦植被句法》(Plugatar et al., 2020)的积极编制工作已经开始。在2021年11月1日至3日在克里米亚共和国雅尔塔举行的“俄罗斯植物群落分类的理论问题”研讨会上,代表了29份贡献(其中5份通过Skype)。来自圣彼得堡、莫斯科、新西伯利亚、乌法、符拉迪沃斯托克、雅库茨克、锡克特夫卡尔、布良斯克、顿河畔罗斯托夫、汉特-曼西斯克、叶卡捷琳堡、纳尔奇克、库尔斯克、雅尔塔的26位科学家(图1、2)参加了本次会议。对森林、草原、草甸、水生、盐生、共向、高寒、北极、岩生、滨海植被和湿地的分类系统进行了综述。讨论了句法检查表开发开始后出现的一些问题,如单位的验证问题、一些公认的句法决定的不确定性问题、某些地区的句法知识不充分等。俄罗斯科学院三个植物园(分别来自雅尔塔、新西伯利亚和符拉迪沃斯托克)的负责人签署了关于启动“俄罗斯联邦国家植物园网络”科学联盟的文件(图3),重点是在植被科学、生态学、生物气候学和生物地理学方面进行联合基础研究;发展国家植被分类和国家生境类型学,为评估我国碳平衡、自然资源合理利用和植被保护提供科学依据。讲习班的决议强调有必要考虑将俄罗斯国家植被分类的发展作为俄罗斯联邦植物社会学家未来五年的优先任务。
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引用次数: 1
Carici supinae–Betuletea pendulae — new forest vegetation class in steppe zone of West Siberian plain and Transural Plateau 西西伯利亚平原和越冬高原草原带森林植被新分类
Q4 Agricultural and Biological Sciences Pub Date : 2021-01-01 DOI: 10.31111/vegrus/2021.40.65
N. Lashchinskiy, N. Makunina
The steppe zone occupies only the southernmost part of West Siberian plain and Transural Plateau, approximately between 54° and 52° N. Zonal communities are species-rich grasslands (Isachenko, Rachkovskaya, 1961; Lapshina, Lavrenko, 1985), but due to relief and geological substrates there are numerous isolated saucer-shaped suffosion depression among flat steppe landscape occupied by small (0.5–1.5 ha) forest massifs. Previously such landscapes were described as “false forest-steppe” or “kolok steppe” following local term “kolok” that is the forest “island”. In some areas such “forest islands” occupy up to 20 % of whole area that makes difficult to identify the border between steppe and forest-steppe zones, in particular in anthropogenically transformed landscape. Until now, there is no information on the structure and floristic composition of such forests, except for brief mentiones in papers on steppe vegetation. The main aim of this paper is to assess the diversity of the deciduous forests in the steppe zone within the study area and establish their syntaxonomical position in the floristic classification. The research is based on 376 original relevés performed by authors in 2007–2016 years in steppe zone from Urals on the west to the Ob river valley on the east. Small intrazonal forest massifs are described as a part of the new class wich contains one order, two alliances and seven associations with four subassociations. Class Carici supinae–Betuletea pendulae class nov. hoc loco. Nomenclature type (holotypus hoc loco) is the order Carici supinae–Betuletalia pendulae ord. nov hoc loco. Diagnostic species: Artemisia austriaca, A. pontica, A. sericea, Carex praecox, C. supina, Festuca rupicola, Medicago falcata, Phleum phleoides, Spiraea crenata, Thymus marschallianus, Veronica spicata, V. spuria; all are mesoxerophytes, most are typical for the steppe class Festuco–Brometea. Order Carici supinae–Betuletalia pendulae ord. nov. hoc loco. Nomenclature type (holotypus hoc loco) is the alliance Sileno nutantis–Betulion pendulae all. nov. hoc loco. Diagnostic species are the same as for class. There are two alliances differing in geographic distribution and habitats within the order. Alliance Sileno nutantis–Betulion pendulae all. nov. hoc loco. Nomenclature type (holotypus hoc loco) is the ass. Sileno nutantis–Betuletum pendulae ass. nov. hoc loco. Diagnostic species: Adenophora lilifolia, Campanula bononiensis, C. wolgensis, Euphorbia subtilis, Galium boreale, Plantago urvillei, Xanthoselinum alsaticum. Association includes deciduous, mostly birch, forests which occur on well-drained plains as small massifs in river valleys and in suffosion depressions. There are four associations differing by habitat moistening within the alliance. Ass. Sileno nutantis–Betuletum pendulae ass. nov. hoc loco (nomenclature type (holotypus hoc loco) — Table 1, relevé 5 (field number L13-035), N 53.09602°, E 60.91305°) unites forests at the base of the northoriented sl
草原区仅占西西伯利亚平原和越界高原的最南端,大约在北纬54°至52°之间,地带性群落是物种丰富的草原(Isachenko, Rachkovskaya, 1961;Lapshina, Lavrenko, 1985),但由于地形起伏和地质底物的作用,在被小(0.5-1.5公顷)森林块体占据的平坦草原景观中存在许多孤立的碟状洼地。以前,这样的景观被描述为“假森林草原”或“kolok草原”,当地术语“kolok”是森林“岛”。在一些地区,这种“森林岛屿”占据了整个地区的20%,这使得很难确定草原和森林草原带之间的边界,特别是在人为改变的景观中。到目前为止,除了在关于草原植被的论文中简要提到外,没有关于这种森林的结构和植物区系组成的信息。本文的主要目的是评价研究区内草原带落叶森林的多样性,确定其在区系分类中的分类学地位。该研究基于作者2007-2016年在西乌拉尔至东鄂布河谷的草原地带进行的376次原始相关研究。小的带内森林块体被描述为新类的一部分,其中包含一个阶,两个联盟和七个协会,四个亚协会。长尾蕨纲:长尾蕨类:长尾蕨类:长尾蕨类。命名类型(holotypus hoc loco)是目Carici supinae-Betuletalia pendulae order . nov hoc loco。诊断种:奥地利黄花蒿、蓬花蒿、绢花蒿、早花蒿、穗花蒿、羊蹄草、falcatago falcata、phleoides、绣线菊、胸腺草、Veronica spicata、V. spuria;均为中叶植物,多数为典型的草原类羊茅属。金鱼目-金鱼目钟摆单。命名类型(holotypus hoc loco)是联盟Sileno nutantis-Betulion pend钟摆所有。11月特别的火线。诊断种与分类相同。在该目中有两个不同的地理分布和栖息地的联盟。联盟无声无息-钟摆之间的所有。11月特别的火线。命名类型(holotypus hoc loco)为Sileno nutantis-Betuletum pendulae ass. nov. hoc loco。诊断种:百合腺苷、风铃、狼耳草、枯草大戟、北风草、车前草、阿尔萨特黄花。关联包括落叶林,主要是白桦林,在排水良好的平原上以河谷和洼地的小块状出现。根据联盟内栖息地的湿润程度不同,有四个协会。as . Sileno nutantis-Betuletum pendulae Ass. nov. hoc loco(命名类型(holotypus hoc loco) -表1,相关资料5(野外编号L13-035),北纬53.09602°,东纬60.91305°)将整个类分布区的北向斜坡底部和大型洼地外围的森林统一起来。诊断种=联合诊断种。a . Pulsatillo patentis-Betuletum pendulae Ass. 11 . hoc loco(命名类型(holotypus hoc loco)) -表2,相关文献6(野外编号L12-134),北纬52.46126°,东经60.32398°)包括该类分布区域西南部靠近西西伯利亚平原和哈萨克高原交界处最干燥生境的森林块体。诊断种:大跟虫、天线虫、高背蝇、沙漠旋毛虫、针叶树、单叶树、白头翁、白头翁。Subass。白头山-典型白头山(表2,相关1-13)。外形、无月经型和诊断种与该协会相同。Subass。白头翁-白头翁-白头翁-白头翁。11 . hoc loco(命名类型(holotypus hoc loco) -表2,相关文献17(野外编号L15-074),北纬52.05014°,东经59.14092°)包括类分布区西部边界的小块白杨林。诊断种:亚美尼亚蒿、芦笋、黑贝母、水杨、木犀草、木犀草、木犀草、木犀草。Ass. Brachypodio pinnatim - betuletum pendulae Ass. nov. hoc loco(命名类型(holotypus hoc loco) -表3,相关文献8(场号L13-130), N 55.32487, E 64.62582)可以被认为是在Carici supinae-Betuletea pendulae和Brachypodio pinnatim - betuletea pendulae类之间的过渡分类。诊断种:短竹、菖蒲、玉竹。属群包括两个亚属群:典型羽状矮脚草-羽状矮脚草和地理分布不同的羽状矮脚草-羽状矮脚草。附属物:散形附属物:附属物:钟摆附属物。 草原区仅占西西伯利亚平原和越界高原的最南端,大约在北纬54°至52°之间,地带性群落是物种丰富的草原(Isachenko, Rachkovskaya, 1961;Lapshina, Lavrenko, 1985),但由于地形起伏和地质底物的作用,在被小(0.5-1.5公顷)森林块体占据的平坦草原景观中存在许多孤立的碟状洼地。以前,这样的景观被描述为“假森林草原”或“kolok草原”,当地术语“kolok”是森林“岛”。在一些地区,这种“森林岛屿”占据了整个地区的20%,这使得很难确定草原和森林草原带之间的边界,特别是在人为改变的景观中。到目前为止,除了在关于草原植被的论文中简要提到外,没有关于这种森林的结构和植物区系组成的信息。本文的主要目的是评价研究区内草原带落叶森林的多样性,确定其在区系分类中的分类学地位。该研究基于作者2007-2016年在西乌拉尔至东鄂布河谷的草原地带进行的376次原始相关研究。小的带内森林块体被描述为新类的一部分,其中包含一个阶,两个联盟和七个协会,四个亚协会。长尾蕨纲:长尾蕨类:长尾蕨类:长尾蕨类。命名类型(holotypus hoc loco)是目Carici supinae-Betuletalia pendulae order . nov hoc loco。诊断种:奥地利黄花蒿、蓬花蒿、绢花蒿、早花蒿、穗花蒿、羊蹄草、falcatago falcata、phleoides、绣线菊、胸腺草、Veronica spicata、V. spuria;均为中叶植物,多数为典型的草原类羊茅属。金鱼目-金鱼目钟摆单。命名类型(holotypus hoc loco)是联盟Sileno nutantis-Betulion pend钟摆所有。11月特别的火线。诊断种与分类相同。在该目中有两个不同的地理分布和栖息地的联盟。联盟无声无息-钟摆之间的所有。11月特别的火线。命名类型(holotypus hoc loco)为Sileno nutantis-Betuletum pendulae ass. nov. hoc loco。诊断种:百合腺苷、风铃、狼耳草、枯草大戟、北风草、车前草、阿尔萨特黄花。关联包括落叶林,主要是白桦林,在排水良好的平原上以河谷和洼地的小块状出现。根据联盟内栖息地的湿润程度不同,有四个协会。as . Sileno nutantis-Betuletum pendulae Ass. nov. hoc loco(命名类型(holotypus hoc loco) -表1,相关资料5(野外编号L13-035),北纬53.09602°,东纬60.91305°)将整个类分布区的北向斜坡底部和大型洼地
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引用次数: 0
To the syntaxonomy of the Middle Urals weeding vegetation 中乌拉尔地区除草植被的分类学
Q4 Agricultural and Biological Sciences Pub Date : 2021-01-01 DOI: 10.31111/vegrus/2021.40.95
G. Khasanova, S. Yamalov, M. Lebedeva, A. Tretyakova, P. Kondratkov, N. Grudanov
The decrease in the crop area and changes in their ratio have led to changes in weed vegetation in the Middle Urals, where its cenoflora was studied relatively well (Tretyakova, 2006; Tretyakova, Kondratkov, 2018; Kondratkov, Tretyakova, 2018; 2019 a, b), while community diversity so far has not yet been assessed. The first results of the ecological and floristic classification of basic crop communities (wheat, barley, oats, corn, potatoes, peas, rapeseed etc.) in six botanical and geographical regions of the taiga and forest-steppe zones (Kulikov et al., 2013) within the Sverdlovsk region, based on the analysis of 160 relevés performed by A. S. Tretyakova, P. V. Kondratkov and N. Yu. Grudanov in 2019, are presented. Communities are classified within the order Aperetalia spicae-venti J. Tx. et Tx. in Malato-Beliz et al. 1960 of the class Papaveretea rhoeadis S. Brullo et al. 2001 nom. conserv. propos. and assigned to the alliance Scleranthion annui (Kruseman et Vlieger 1939 nom. conserv. propos.) Sissingh in Westhoff et al. 1946), which represents the most mesophytic types of weed vegetation in the Urals. Three associations allocated in alliance (two for the first time): Solano nigri–Erodietum cicutarii ass. nov. hoc loco (Table 2, syntaxon 7; Table 4); nomenclature type (holotypus hoc loco) — Table 4, rel. 11 (number in database — 25): Sverdlovsk Region, Sysertskiy district, Kunarskoe village environs, 23.07.2019, authors — A. S. Tretyakova, P. V. Kondratkov);Lamio amplexicaulis–Stellarietum mediae ass. nov. hoc loco (Table 2, syntaxon 8; table 5; nomenclature type (holotypus hoc loco) — Table 5, rel. 10 (number in database — 27): Sverdlovsk Region, Sysertskiy district, Shaidurovo village environs, 22.07.2019, authors — A. S. Tretyakova, P. V. Kondratkov); ass. Euphorbio helioscopiae–Fumarietum officinalis Khasanova et al. 2018. The last one includes two variants. Two unranked communities are also described. Distribution areas of syntaxa cover the southern taiga and pre-forest-steppe pine-birch forests subzones of the taiga zone and the northern forest-steppe subzone of the forest-steppe zone. Habitats are confined mainly to gray forest soils, in rare cases — leached chernozems. The mostly wide distributed is the ass. Euphorbio helioscopiae-Fumarietum officinalis, also common to the south of the study area, in the Republic of Bashkortostan. Аss. Lamio amplexicaulis–Stellarietum mediae is identified in five botanical and geographical districts within the Sverdlovsk region. The confluence with crops and the accompanying system of agriculturе is different. The species composition of the established syntaxa was compared with the associations of the alliance Scleranthion annui described earlier in the Republic of Bashkortostan. The species richness of the latter is generally higher. Differences in species composition of diagnostic groups were revealed. There is decreasing of constancy of Tripleurospermum inodorum, Centaurea cyanus, Polygonum avicu
作物面积的减少及其比例的变化导致乌拉尔中部杂草植被的变化,在那里对其新植物区系的研究相对较好(Tretyakova, 2006;特列季亚科娃,康德拉科夫,2018;康德拉特科夫,特列季亚科娃,2018;2019年a, b),而迄今为止尚未对社区多样性进行评估。基于A. S. Tretyakova、P. V. Kondratkov和N. Yu对160个相关的数据进行的分析,在斯维尔德洛夫斯克地区针叶林和森林草原带的六个植物和地理区域(小麦、大麦、燕麦、玉米、土豆、豌豆、油菜籽等)的基本作物群落(小麦、大麦、燕麦、土豆、豌豆、油菜籽等)的生态和区系分类的第一批结果。格鲁达诺夫在2019年,都提出了。在Malato-Beliz etal . 1960的Papaveretea rhoeadis S. Brullo etal . 2001中,群落被分类为Aperetalia spicae-venti J. Tx. et Tx.。言之凿凿。并被分配到联盟基金会(Kruseman et Vlieger 1939)。提议)。Sissingh in Westhoff et al. 1946),它代表了乌拉尔地区杂草植被中最多的中植物类型。3个组合(其中2个为首次):Solano nigri-Erodietum cicutarii ass. nov. hoc loco(表2,句法7;表4);命名类型(holotypus hoc loco) -表4,rel. 11(数据库中数量- 25):Sverdlovsk地区,Sysertskiy区,Kunarskoe村周边,2019年7月23日,作者- A. S. Tretyakova, P. V. Kondratkov);Lamio amplexicaulis-Stellarietum mediae ass. nov. hoc loco(表2,句法8;表5;命名类型(holotypus hoc loco) -表5,rel. 10(数据库数量- 27):斯维尔德洛夫斯克州,Sysertskiy区,Shaidurovo村周边,2019年7月22日,作者- A. S. Tretyakova, P. V. Kondratkov);[j] . Euphorbio helioscopiae-Fumarietum officinalis Khasanova等。2018。最后一个包括两个变体。还描述了两个未排名的社区。其区系主要分布在针叶林带的南部针叶林和森林-草原前松桦林亚区以及森林-草原带的北部森林-草原亚区。栖息地主要局限于灰色森林土壤,在极少数情况下-淋溶黑钙土。分布最广的是Euphorbio helioscopae - fumarietum officinalis,也常见于研究区南部的巴什科尔托斯坦共和国。А党卫军。在斯维尔德洛夫斯克地区的5个植物学和地理区域中鉴定出了大叶星。与作物的融合和农业配套制度是不同的。建立的句法群的物种组成与先前在巴什科尔托斯坦共和国描述的Scleranthion annui联盟的关联进行了比较。后者的物种丰富度一般较高。揭示了诊断组物种组成的差异。在第一个群落的新植物区系中,三胸草、半毛菊、木犀蓼、Raphanus raphanstrum的稳定性有所下降,而旱生性较强的群落群Caucalidion lappulae von Rochow 1951的诊断种(Galeopsis bifida、Persicaria lapathifolia、Silene noctiflora、Galeopsis ladanum、Melilotus officinalis)、草牧场和森林边缘的植物(Medicago lupina、Trifolium hybridum、Trifolium medium、Achillea millefolium等)的稳定性也有所下降。旱生种(天葵,苋,狗尾草,大戟)。后者可能与某些作物(向日葵、亚麻等)的田间面积显著减少有关。排序分析的结果说明了比较类群在物种组成和生态方面的差异。乌拉尔南部的杂草群落位于dca排序图的左侧,乌拉尔中部的杂草群落位于右侧。在第一个轴上,从旱生植物最多的Linario vulgari-Lactucetum serriolae群落向中生植物最多的Lamio amplexicaulum - stellarietum mediae和Euphorbio helioscopium - fumarietum officalinae群落发生了变化。第二个轴显示了在农艺因素-栽培及其相应的农业系统的影响下物种组成的变化,冬季谷物群落(Consolido regi - centaureetum cyanae和Galeopsetum bifidae)被春季谷物和行作物群落(玉米、土豆、豌豆、油菜籽)所取代,它们被分配给Linario vulgari-Lactucetum serriolae和solan nigri-Erodietum cicutarii。
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