Pub Date : 2022-12-23DOI: 10.14258/turczaninowia.25.4.9
Ilja N. Timukhin, B. Tuniyev
The description of Galanthus lebedevae from limestone canyons of foothills in south-east Sochi is presented. The species belongs to series Latifolii, subseries Viridifolii (Kem.-Nath.) A. P. Davis. It shows an external morphological similarity with the sympatric Colchian species G. platyphyllus, G. krasnovii, G. panjutini, and G. woronowii, however, it differs sharply from all the listed species in the content of nuclear (2C) DNA, phenology, and ecology. Narrow northern Colchis endemic, which developed under the conditions of the warmest foothill limestone area of the Colchis refugium of tertiary-relict species.
{"title":"Galanthus lebedevae (Amaryllidaceae) – a new species from Sochi","authors":"Ilja N. Timukhin, B. Tuniyev","doi":"10.14258/turczaninowia.25.4.9","DOIUrl":"https://doi.org/10.14258/turczaninowia.25.4.9","url":null,"abstract":"The description of Galanthus lebedevae from limestone canyons of foothills in south-east Sochi is presented. The species belongs to series Latifolii, subseries Viridifolii (Kem.-Nath.) A. P. Davis. It shows an external morphological similarity with the sympatric Colchian species G. platyphyllus, G. krasnovii, G. panjutini, and G. woronowii, however, it differs sharply from all the listed species in the content of nuclear (2C) DNA, phenology, and ecology. Narrow northern Colchis endemic, which developed under the conditions of the warmest foothill limestone area of the Colchis refugium of tertiary-relict species.","PeriodicalId":45595,"journal":{"name":"Turczaninowia","volume":" ","pages":""},"PeriodicalIF":0.5,"publicationDate":"2022-12-23","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"47904422","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":0,"RegionCategory":"","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
Pub Date : 2022-12-23DOI: 10.14258/turczaninowia.25.4.8
Georgiy A. Lazkov, P. Vesselova
A composition of taxa included in Dracocephalum imberbe Bunge s. l. is discussed. Species rank of D. laniflorum Rupr. is reinstated based on the following characters: underground part is represented by a system of branching rhizomes, bracts absent, calyx 12–14 mm long, with triangular or narrowly triangular, pointless teeth, the middle tooth of the upper lip up to 1.5 times wider than the lateral ones and plant being nearly scentless. In contrast, in D. imberbeunderground part is represented by the taproot and developed branches of the caudex, bracts present, with well-developed non-rigid points, calyx 15–18 mm long, with lanceolate or narrowly triangular pointed (though not rigid) teeth, the middle tooth of the upper lip 1.5–2 times wider than the lateral ones, the presence of a smell that persists in herbarium specimens. Dracocephalum laniflorum is firstly reported for flora of Kazakhstan (Almaty Region) and China(Xinjiang Uygur Autonomous Region) based on the study of herbarium materials stored in AA and LE, as well as scanned herbarium specimens stored in MW, HNWP and PE, which were available through the Moscow Digital Herbarium and Chinese Virtual Herbarium (CVH) systems.
{"title":"On two species of Dracocephalum L. (Lamiaceae)","authors":"Georgiy A. Lazkov, P. Vesselova","doi":"10.14258/turczaninowia.25.4.8","DOIUrl":"https://doi.org/10.14258/turczaninowia.25.4.8","url":null,"abstract":"A composition of taxa included in Dracocephalum imberbe Bunge s. l. is discussed. Species rank of D. laniflorum Rupr. is reinstated based on the following characters: underground part is represented by a system of branching rhizomes, bracts absent, calyx 12–14 mm long, with triangular or narrowly triangular, pointless teeth, the middle tooth of the upper lip up to 1.5 times wider than the lateral ones and plant being nearly scentless. In contrast, in D. imberbeunderground part is represented by the taproot and developed branches of the caudex, bracts present, with well-developed non-rigid points, calyx 15–18 mm long, with lanceolate or narrowly triangular pointed (though not rigid) teeth, the middle tooth of the upper lip 1.5–2 times wider than the lateral ones, the presence of a smell that persists in herbarium specimens. Dracocephalum laniflorum is firstly reported for flora of Kazakhstan (Almaty Region) and China(Xinjiang Uygur Autonomous Region) based on the study of herbarium materials stored in AA and LE, as well as scanned herbarium specimens stored in MW, HNWP and PE, which were available through the Moscow Digital Herbarium and Chinese Virtual Herbarium (CVH) systems.","PeriodicalId":45595,"journal":{"name":"Turczaninowia","volume":" ","pages":""},"PeriodicalIF":0.5,"publicationDate":"2022-12-23","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"46917108","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":0,"RegionCategory":"","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
Pub Date : 2022-12-23DOI: 10.14258/turczaninowia.25.4.4
A. Chkalov
From four Alchemilla species described by B. S. Kharitontsev, only one (A. incisa Charitoncev, non Buser) is considered by us as a separate one, for which a new, legitimate name, A. tobolica, is proposed. Type specimens of both A. laxa (nom. illeg., non Plocek) and A. parvula were identified by us as A. micans while that of A. subobtusa was undoubtedly identified by us as A. circularis Juz.
{"title":"On the Alchemilla species described by B. S. Kharitontsev","authors":"A. Chkalov","doi":"10.14258/turczaninowia.25.4.4","DOIUrl":"https://doi.org/10.14258/turczaninowia.25.4.4","url":null,"abstract":"From four Alchemilla species described by B. S. Kharitontsev, only one (A. incisa Charitoncev, non Buser) is considered by us as a separate one, for which a new, legitimate name, A. tobolica, is proposed. Type specimens of both A. laxa (nom. illeg., non Plocek) and A. parvula were identified by us as A. micans while that of A. subobtusa was undoubtedly identified by us as A. circularis Juz.","PeriodicalId":45595,"journal":{"name":"Turczaninowia","volume":" ","pages":""},"PeriodicalIF":0.5,"publicationDate":"2022-12-23","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"43848513","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":0,"RegionCategory":"","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
Pub Date : 2022-12-23DOI: 10.14258/turczaninowia.25.4.2
D. German, P. Vesselova
Taxonomy and species diversity of the mustard family (Cruciferae or Brassicaceae) in Kazakhstan is updated. Draba bajtenovii described and hitherto known from Kungei Alatau is synonymized with D. fladnizensis for which it is a southern limit of distribution in the Eurasia, while Trans-Ili Alatau endemic D. microcarpella is found conspecific with a montane Asian D. lanceolata. Noccaea thlaspidioides (N. cochleariformis) is confirmed to occur in the country based on collections from Altai. Finally, Isatis trachycarpa and Smelowskia annua are shown to be absent in the flora of Kazakhstan due to the lack of gatherings confirming their occurrence in the country.
{"title":"On the taxonomy and diversity of Cruciferae in Kazakhstan","authors":"D. German, P. Vesselova","doi":"10.14258/turczaninowia.25.4.2","DOIUrl":"https://doi.org/10.14258/turczaninowia.25.4.2","url":null,"abstract":"Taxonomy and species diversity of the mustard family (Cruciferae or Brassicaceae) in Kazakhstan is updated. Draba bajtenovii described and hitherto known from Kungei Alatau is synonymized with D. fladnizensis for which it is a southern limit of distribution in the Eurasia, while Trans-Ili Alatau endemic D. microcarpella is found conspecific with a montane Asian D. lanceolata. Noccaea thlaspidioides (N. cochleariformis) is confirmed to occur in the country based on collections from Altai. Finally, Isatis trachycarpa and Smelowskia annua are shown to be absent in the flora of Kazakhstan due to the lack of gatherings confirming their occurrence in the country.","PeriodicalId":45595,"journal":{"name":"Turczaninowia","volume":" ","pages":""},"PeriodicalIF":0.5,"publicationDate":"2022-12-23","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"44274639","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":0,"RegionCategory":"","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
Pub Date : 2022-11-30DOI: 10.14258/turczaninowia.25.4.1
Alexsey V. Vaganov, T. Sinitsyna, Maxim G. Kutsev, M. V. Skaptsov, Yelizaveta A. Zholnerova, P. Kosachev, A. Kechaykin, S. V. Smirnov, A. Shmakov
The article presents first data of the work on DNA barcoding of type specimens of ALTB Herbarium (Barnaul, Russia). Obtained sequences of ITS and trnL-trnF, trnH-psbA markers of DNA were deposited in NCBI GenBank, and corresponding dataset was published in the GBIF (Global Biodiversity Information Facility).
{"title":"DNA barcodes of the vascular flora of the Altai Mountain Country: type material of the Herbarium ALTB","authors":"Alexsey V. Vaganov, T. Sinitsyna, Maxim G. Kutsev, M. V. Skaptsov, Yelizaveta A. Zholnerova, P. Kosachev, A. Kechaykin, S. V. Smirnov, A. Shmakov","doi":"10.14258/turczaninowia.25.4.1","DOIUrl":"https://doi.org/10.14258/turczaninowia.25.4.1","url":null,"abstract":"\u0000 \u0000 \u0000The article presents first data of the work on DNA barcoding of type specimens of ALTB Herbarium (Barnaul, Russia). Obtained sequences of ITS and trnL-trnF, trnH-psbA markers of DNA were deposited in NCBI GenBank, and corresponding dataset was published in the GBIF (Global Biodiversity Information Facility). \u0000 \u0000 \u0000","PeriodicalId":45595,"journal":{"name":"Turczaninowia","volume":" ","pages":""},"PeriodicalIF":0.5,"publicationDate":"2022-11-30","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"49276427","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":0,"RegionCategory":"","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
Pub Date : 2022-09-30DOI: 10.14258/turczaninowia.25.3.1
R. Kamelin
East Asia is a very integral region, the East Asian sub-kingdom of the Holarctic, in which two large areas stand out: the Sino-Himalayan and Sino-Japanese. The flora of East Asia is rich and unique. About 25 000 species of vascular plants grow in this area. There are about 23 endemic families and over 350 endemic genera. The number of endemic species is about 15 000. In East Asia, there are modern centers of diversity for many plants, but there are also many relict groups that have survived in this territory from past richly developed types.
{"title":"Flora of East Asia and its genesis","authors":"R. Kamelin","doi":"10.14258/turczaninowia.25.3.1","DOIUrl":"https://doi.org/10.14258/turczaninowia.25.3.1","url":null,"abstract":"East Asia is a very integral region, the East Asian sub-kingdom of the Holarctic, in which two large areas stand out: the Sino-Himalayan and Sino-Japanese. The flora of East Asia is rich and unique. About 25 000 species of vascular plants grow in this area. There are about 23 endemic families and over 350 endemic genera. The number of endemic species is about 15 000. In East Asia, there are modern centers of diversity for many plants, but there are also many relict groups that have survived in this territory from past richly developed types.","PeriodicalId":45595,"journal":{"name":"Turczaninowia","volume":" ","pages":""},"PeriodicalIF":0.5,"publicationDate":"2022-09-30","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"44511850","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":0,"RegionCategory":"","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
Pub Date : 2022-09-30DOI: 10.14258/turczaninowia.25.3.12
L. Kalinina
Information on 18 species recorded for the first time in the Leningrad Region is given, including seven (Atheniella leptophylla, Cystoagaricus sylvestris, Lepiota subalba, Mycena mirata, Pluteus ephebeus, Psathyrella longicauda and Psathyrella pygmaea) previously not known from the north-west of European Russia. The species are annotated with comments on ecology, and colour illustrations of basidiomata are provided. Three species are recommended for the next edition of Red Data Book of the Leningrad Region: Atheniella leptophylla as confined to limestone outcrops, rare in the region, and Cystoagaricus sylvestris and Pluteus aurantiorugosus vulnerable due to substrate confinement to elm wood.
{"title":"Agaricoid fungi new to Leningrad Region, Russia","authors":"L. Kalinina","doi":"10.14258/turczaninowia.25.3.12","DOIUrl":"https://doi.org/10.14258/turczaninowia.25.3.12","url":null,"abstract":"Information on 18 species recorded for the first time in the Leningrad Region is given, including seven (Atheniella leptophylla, Cystoagaricus sylvestris, Lepiota subalba, Mycena mirata, Pluteus ephebeus, Psathyrella longicauda and Psathyrella pygmaea) previously not known from the north-west of European Russia. The species are annotated with comments on ecology, and colour illustrations of basidiomata are provided. Three species are recommended for the next edition of Red Data Book of the Leningrad Region: Atheniella leptophylla as confined to limestone outcrops, rare in the region, and Cystoagaricus sylvestris and Pluteus aurantiorugosus vulnerable due to substrate confinement to elm wood.","PeriodicalId":45595,"journal":{"name":"Turczaninowia","volume":" ","pages":""},"PeriodicalIF":0.5,"publicationDate":"2022-09-30","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"48288599","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":0,"RegionCategory":"","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
Pub Date : 2022-09-30DOI: 10.14258/turczaninowia.25.3.15
I. Schanzer, A. Fedorova, O. I. Kuznetsova, I. Belyaeva, O. Razumova
Salix fursaevii, a species endemic to the Volga River flood plain, was described by E. Mavrodiev and co-authors in Byull. Moskovsk. Obshch. Isp. Prir. in 2012. It differs from the closely related and widespread S. triandra by a few quantitative morphological characters and, most of all by its much later flowering time. This late-flowering willow was known and studied before as an ecotype of S. triandra. Our study of a large sample of S. triandra s. l. revealed that 1) S. triandra s. str. and S. fursaevii could not be clearly distinguished using several morphological characters and the time of flowering; 2) specimens morphologically corresponding to S. fursaevii are with a single exception strictly confined to the Volga River flood plain; 3) the populations of S. triandra and S. fursaevii do not differ by nrITS sequences; 4) the populations of S. triandra and S. fursaevii are poorly differentiated by chloroplast atpB-rbcL sequences; 5) representatives of both taxa are tetraploids 2n = 4x = 76. Poor differentiation of populations by chloroplast sequences is indicative of the intraspecific variability, the geographic pattern of which suggests that the Volga River valley was a probable glacial refugium and a source of post glacial spread of S. triandra.
{"title":"Is Salix fursaevii Mavrodiev (Salicaceae) a palaeoendemic of the Volga River valley or an ecotype of S. triandra L.?","authors":"I. Schanzer, A. Fedorova, O. I. Kuznetsova, I. Belyaeva, O. Razumova","doi":"10.14258/turczaninowia.25.3.15","DOIUrl":"https://doi.org/10.14258/turczaninowia.25.3.15","url":null,"abstract":"Salix fursaevii, a species endemic to the Volga River flood plain, was described by E. Mavrodiev and co-authors in Byull. Moskovsk. Obshch. Isp. Prir. in 2012. It differs from the closely related and widespread S. triandra by a few quantitative morphological characters and, most of all by its much later flowering time. This late-flowering willow was known and studied before as an ecotype of S. triandra. Our study of a large sample of S. triandra s. l. revealed that 1) S. triandra s. str. and S. fursaevii could not be clearly distinguished using several morphological characters and the time of flowering; 2) specimens morphologically corresponding to S. fursaevii are with a single exception strictly confined to the Volga River flood plain; 3) the populations of S. triandra and S. fursaevii do not differ by nrITS sequences; 4) the populations of S. triandra and S. fursaevii are poorly differentiated by chloroplast atpB-rbcL sequences; 5) representatives of both taxa are tetraploids 2n = 4x = 76. Poor differentiation of populations by chloroplast sequences is indicative of the intraspecific variability, the geographic pattern of which suggests that the Volga River valley was a probable glacial refugium and a source of post glacial spread of S. triandra.","PeriodicalId":45595,"journal":{"name":"Turczaninowia","volume":" ","pages":""},"PeriodicalIF":0.5,"publicationDate":"2022-09-30","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"45846460","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":0,"RegionCategory":"","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
Pub Date : 2022-09-30DOI: 10.14258/turczaninowia.25.3.21
A. P. Shalimov
Based on examination of herbarium specimen that was previously misidentified as Selaginella helvetica (L.) Spring, we report the first voucher-based record of S. kraussiana (Kunze) A. Braun from the eastern Carpathian Mountains in West Ukraine. A morphological comparison of S. kraussiana with S. helvetica and S. selaginoides occurring in this region is given. Selaginella kraussiana differs from them in habit, vegetative leaves and strobili structure.
基于对以前被误认为helvetica卷柏(Selaginella helvetica, L.)的植物标本馆标本的检验春天,我们报道了来自乌克兰西部喀尔巴阡山脉东部的S. kraussiana (Kunze) A. Braun的第一个凭证记录。本文对发生在该地区的kraussiana与S. helvetica和S. selaginoides进行了形态学比较。柏卷草在习性、营养叶和球团结构上与它们不同。
{"title":"First record of Selaginella kraussiana (Kunze) A. Braun in Western Ukraine (Eastern Europe)","authors":"A. P. Shalimov","doi":"10.14258/turczaninowia.25.3.21","DOIUrl":"https://doi.org/10.14258/turczaninowia.25.3.21","url":null,"abstract":"Based on examination of herbarium specimen that was previously misidentified as Selaginella helvetica (L.) Spring, we report the first voucher-based record of S. kraussiana (Kunze) A. Braun from the eastern Carpathian Mountains in West Ukraine. A morphological comparison of S. kraussiana with S. helvetica and S. selaginoides occurring in this region is given. Selaginella kraussiana differs from them in habit, vegetative leaves and strobili structure.","PeriodicalId":45595,"journal":{"name":"Turczaninowia","volume":" ","pages":""},"PeriodicalIF":0.5,"publicationDate":"2022-09-30","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"47946573","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":0,"RegionCategory":"","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}
Pub Date : 2022-09-30DOI: 10.14258/turczaninowia.25.3.13
Ilja B. Kucherov, A. A. Zverev
Phytocoenotic behaviour of 14 nemoral or boreal-nemoral vascular plant species is analyzed in the middle- and southern-taiga subzones in comparison with that in the broadleaved-forest zone, based upon the set of 1669 relevés made in European Russia in 1996–2018. The nemoral species are mainly represented by shadow mesoeutrophic plants from the “corteges” of Tilia cordata s. l. and Fagus sylvatica. The more light-demanding plants from the Quercus robur “cortege”, either mesotrophic, or mesoeutrophic, more often demonstrate the boreal-nemoral type of zonal distribution. In the boreal-forest zone, linden and beech companions are typical for the southern-boreal nemoral-herb / sorrel spruce (Picea abies s. l.) forests on placors as well as the valley / riparian tall-herb forests, including the middle-boreal ones. The oak companions also inhabit grass-feathermoss pine (Pinus sylvestris) forests, besides the spruce forest types mentioned. Species from all the named “corteges” are also common in the small-leaved forests. The Betula companions which grow together with the nemoral plants in boreal forests could also be subdivided into the two florogenetical groups, namely a) the boreal, originally subalpine plants and b) the boreal-nemoral ones, originally oak companions. Species from the latter group often become widespread and dominant in the small-leaved forests; a good example is provided by Aegopodium podagraria. The rank correlation (Spearman rS) analysis shows the projective cover increase is connected with either the growing season warmth supply, or the climate oceanicity for both nemoral and boreal-nemoral species. These effects are most sound in the placor community sequence from broadleaved forests to sorrel and then bilberry-feathermoss spruce ones but weakened in the tall-herb forests on rich soil and completely levelled in the aspen (Populus tremula) forests. Carbonate bedrock outcrops contribute to nemoral species survival in the areas they inhabited during the climatically favourable epochs; they also favoured the migration of these species during the interglacials. Individualization of coenotic patterns of nemoral species at their distribution limits does not contradict to joint occurrence of such plants in relict / extrazonal locations.
{"title":"Phytocoenotical behaviour of nemoral and boreal-nemoral plant species in taiga zone communities","authors":"Ilja B. Kucherov, A. A. Zverev","doi":"10.14258/turczaninowia.25.3.13","DOIUrl":"https://doi.org/10.14258/turczaninowia.25.3.13","url":null,"abstract":"Phytocoenotic behaviour of 14 nemoral or boreal-nemoral vascular plant species is analyzed in the middle- and southern-taiga subzones in comparison with that in the broadleaved-forest zone, based upon the set of 1669 relevés made in European Russia in 1996–2018. The nemoral species are mainly represented by shadow mesoeutrophic plants from the “corteges” of Tilia cordata s. l. and Fagus sylvatica. The more light-demanding plants from the Quercus robur “cortege”, either mesotrophic, or mesoeutrophic, more often demonstrate the boreal-nemoral type of zonal distribution. In the boreal-forest zone, linden and beech companions are typical for the southern-boreal nemoral-herb / sorrel spruce (Picea abies s. l.) forests on placors as well as the valley / riparian tall-herb forests, including the middle-boreal ones. The oak companions also inhabit grass-feathermoss pine (Pinus sylvestris) forests, besides the spruce forest types mentioned. Species from all the named “corteges” are also common in the small-leaved forests. The Betula companions which grow together with the nemoral plants in boreal forests could also be subdivided into the two florogenetical groups, namely a) the boreal, originally subalpine plants and b) the boreal-nemoral ones, originally oak companions. Species from the latter group often become widespread and dominant in the small-leaved forests; a good example is provided by Aegopodium podagraria. The rank correlation (Spearman rS) analysis shows the projective cover increase is connected with either the growing season warmth supply, or the climate oceanicity for both nemoral and boreal-nemoral species. These effects are most sound in the placor community sequence from broadleaved forests to sorrel and then bilberry-feathermoss spruce ones but weakened in the tall-herb forests on rich soil and completely levelled in the aspen (Populus tremula) forests. Carbonate bedrock outcrops contribute to nemoral species survival in the areas they inhabited during the climatically favourable epochs; they also favoured the migration of these species during the interglacials. Individualization of coenotic patterns of nemoral species at their distribution limits does not contradict to joint occurrence of such plants in relict / extrazonal locations.","PeriodicalId":45595,"journal":{"name":"Turczaninowia","volume":" ","pages":""},"PeriodicalIF":0.5,"publicationDate":"2022-09-30","publicationTypes":"Journal Article","fieldsOfStudy":null,"isOpenAccess":false,"openAccessPdf":"","citationCount":null,"resultStr":null,"platform":"Semanticscholar","paperid":"42912466","PeriodicalName":null,"FirstCategoryId":null,"ListUrlMain":null,"RegionNum":0,"RegionCategory":"","ArticlePicture":[],"TitleCN":null,"AbstractTextCN":null,"PMCID":"","EPubDate":null,"PubModel":null,"JCR":null,"JCRName":null,"Score":null,"Total":0}