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Sexually dimorphic ornamentation in modern spinicaudatans and the taxonomic implications for fossil clam shrimps 现代棘足动物的两性二形装饰及其蛤虾化石的分类学意义
IF 1.8 3区 地球科学 Q1 Earth and Planetary Sciences Pub Date : 2022-01-01 DOI: 10.4202/app.00892.2021
Xiaoyan Sun, Jin-hui Cheng
The phylogenetic studies of clam shrimps (Branchiopoda, Crustacea) demonstrated that the significance of several morphological characters for classification of branchiopod shells should be critically re-evaluated. Such a venture is partic-ularly important for integrating the taxonomy of fossil and extant branchiopods. One of the shell characters widely used in the branchiopod classification is the carapace ornamentation pattern. This character might, however, be significantly influenced by intraspecific variability and in particular the sexual dimorphism. In this study we investigate the pattern of ornamentation in extant branchiopods—including differences resulting from sexual dimorphism—in order to assess its value for branchiopod taxonomy. We examined 184 individuals representing 10 living species belonging to 7 genera, 5 families, and 2 suborders from China, and compared with the results of previous studies. Although some differences in ornamentation were related to reproductive modes, the basic ornamentation patterns or combinations were stable within each extant species. We found out that some taxa indeed display sexually dimorphic ornamentations, but their basic ornamentation patterns or combinations are stable within each species so they do not significantly influence the taxonomic identification. Integration of data on fossil and extant taxa indicates that similar ornamentation patterns can be observed on familial level of fossil spinicaudatan branchiopods and indicates therefore that characteristic ornamentation patterns can help to identify these taxa in the fossil record. In light of the new molecular phylogeny, we re-evaluated the phylogenetic relationship between fossil and extant spinicaudatan taxa. The resulting tree suggests: (i) paraphyly of the traditional Eosestherioidea, (ii) an affinity between Ozestheria and Triglypta , and (iii) an affinity between Cyzicus and Diestheria or Aquilonoglypta .
对蛤蛄(鳃足纲,甲壳纲)的系统发育研究表明,一些形态学特征对鳃足纲贝壳分类的意义应重新进行批判性评价。这样的冒险对于整合化石和现存的枝足类动物的分类学尤其重要。甲壳纹饰图案是鳃足类动物分类中广泛使用的贝壳特征之一。然而,这一特性可能受到种内变异性,特别是两性二态性的显著影响。在本研究中,我们研究了现存的枝足动物的纹饰模式,包括两性二态性造成的差异,以评估其在枝足动物分类中的价值。本研究调查了中国2亚目5科7属10种184个个体,并与已有研究结果进行了比较。虽然纹饰的差异与繁殖方式有关,但基本的纹饰模式或组合在各现存种内是稳定的。结果表明,某些分类群确实表现出雌雄二型的纹饰,但其基本纹饰模式或组合在每个物种内都是稳定的,因此对分类鉴定没有显著影响。通过对化石和现存分类群的综合分析,发现棘足纲分支足类化石在科系水平上具有相似的纹饰特征,表明其纹饰特征有助于化石记录中这些分类群的识别。根据新的分子系统发育,我们重新评估了化石与现存棘足动物类群的系统发育关系。结果表明:(i)与传统的Eosestherioidea有亲缘关系,(ii) Ozestheria与Triglypta有亲缘关系,(iii) Cyzicus与Diestheria或Aquilonoglypta有亲缘关系。
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引用次数: 3
First three-dimensional skull of the Middle Triassic ichthyosaur Phalarodon fraasi (Mixosauridae) from Svalbard, Norway. 挪威斯瓦尔巴群岛发现的中三叠世法拉西Phalarodon鱼龙(混合龙科)首个三维头骨。
IF 1.8 3区 地球科学 Q1 Earth and Planetary Sciences Pub Date : 2022-01-01 DOI: 10.4202/app.00915.2021
A. Roberts, Victoria Sjøholt Engelschiøn, J. Hurum
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引用次数: 3
A morganucodontan (Mammaliaformes) from the Upper Jurassic Morrison Formation, Utah, USA 美国犹他州上侏罗统莫里森组的一具摩根齿兽(哺乳动物目)
IF 1.8 3区 地球科学 Q1 Earth and Planetary Sciences Pub Date : 2022-01-01 DOI: 10.4202/app.00955.2021
Brian M Davis, K. Jager, G. Rougier, K. Trujillo, K. Chamberlain
A morganucodontan mammaliaform from Morrison We describe two skull fragments of a new morganucodontan from the Cisco Mammal Quarry (Upper Jurassic Morrison Formation), preserving portions of the palate and snout in excellent 3D detail as well as the complete upper postcanine dentition. Morganucodontans are best known by isolated elements and relatively complete skulls of several species of Morganucodon from the Lower Jurassic of Wales and China; this group is fundamental to our understanding of the early evolution of mammals. Cifellilestes ciscoensis gen. et sp. nov. possesses derived features of the snout paired with plesiomorphic construction of the molars; the distal premolars are complex and there is an unusually low count (two) of strongly imbricated molars. This character combination expands craniodental variation for the group. We sampled mudstone from the Cisco Mammal Quarry for ash-fall zircon analysis and obtained a date of 151.50 ± 0.28 Ma. This dates the locality to the earliest Tithonian and slightly younger than other major dated mammal-bearing localities in the Morrison Formation. Cifellilestes represents one of the youngest members of this group and extends the record of morganucodontans in North America by more than 30 Ma. Morganucodontans are a rare component of Late Jurassic faunas but display surprising dental diversity through variations in a tooth count and cusp morphology of a deeply conserved, generalized mammalian tooth pattern, which was fully established in brasilodontid (non-mammalian) ancestors at least 80 my prior. Morganucodonta, dentition, Jurassic, Morrison Formation.
我们描述了来自Cisco哺乳动物采石场(上侏罗纪莫里森组)的一个新的morganucodontan的两个头骨碎片,以极好的3D细节保存了部分上颚和鼻子以及完整的上犬齿。Morganucodontans最为人所知的是来自威尔士和中国下侏罗纪的几个morganucodondon物种的孤立元素和相对完整的头骨;这个群体是我们理解哺乳动物早期进化的基础。ciscoensis . et sp. 11 .具有衍生的鼻部特征与臼齿的半形结构配对;远端前磨牙是复杂的,有一个非常低的计数(两个)的强烈叠瓦磨牙。这一特征组合扩大了这一群体的颅齿变异。我们从思科哺乳动物采石场采集泥岩样本进行灰落锆石分析,得到了151.50±0.28 Ma的年代。这将该地区的年代确定为最早的泰桑期,比莫里森组中其他主要哺乳动物生长的地区略年轻。Cifellilestes代表了这个群体中最年轻的成员之一,并将北美morganucodontans的记录延长了30多万年。Morganucodontans是晚侏罗世动物群中罕见的组成部分,但通过牙齿数量和齿尖形态的变化显示出令人惊讶的牙齿多样性,这是一种深度保守的、广泛的哺乳动物牙齿模式,这在至少80年前的巴西齿类(非哺乳动物)祖先中完全建立起来的。mororganucodonta,齿列,侏罗纪,Morrison组。
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引用次数: 1
Reply to Bauer et al 2022 回复Bauer等人2022
IF 1.8 3区 地球科学 Q1 Earth and Planetary Sciences Pub Date : 2022-01-01 DOI: 10.4202/app.00995.2022
Christopher Paul
Cladistic analyses will be inaccurate if based on inaccurate data. Most blastozoan taxa were defined before cladistics was proposed. Redescription of precladistic taxa should be a priority. Cladistics recognizes similarities between taxa, not differences. Differences always exist, so the two approaches are asymmetrical. Any suggested similarity or homology needs testing, including output of computer-based analyses. New interpretations of plate homology between glyptocystitoid and hemicosmitoid rhombiferans had already rendered input data obsolete before the pan-dichoporite paper was published. Universal Elemental Homology (UEH) is not a scientific hypothesis. It makes no testable predictions; no system of naming plates can.
如果基于不准确的数据,分类分析将是不准确的。大多数囊胚动物分类在分类学提出之前就已被定义。对前灭绝分类群的重新描述应该是一个优先事项。分类学识别的是分类群之间的相似性,而不是差异性。差异总是存在的,所以这两种方法是不对称的。任何建议的相似性或同源性需要测试,包括基于计算机的分析输出。在这篇泛双孢石论文发表之前,关于形囊体类和半锥体类菱形虫板块同源性的新解释已经使输入数据过时。普遍元素同源(UEH)不是一个科学假设。它没有可检验的预测;任何命名系统都无法做到这一点。
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引用次数: 1
Dimorphism in tetragonitid ammonoid Tetragonites minimus from the Upper Cretaceous in Hokkaido, Northern Japan 日本北海道上白垩统四龙石氨样四龙岩的二态性
IF 1.8 3区 地球科学 Q1 Earth and Planetary Sciences Pub Date : 2022-01-01 DOI: 10.4202/app.01000.2022
D. Aiba
. Dimorphism in tetragonitid ammonoid Tetragonites minimus from the Upper Cretaceous in Hokkaido, Northern Japan. Acta
. 日本北海道上白垩统四龙石氨样四龙岩的二态性。学报
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引用次数: 2
Middle and Late Jurassic tube-dwelling polychaetes from the Polish Basin: diversity, palaeoecology and comparisons with other assemblages 波兰盆地中晚侏罗世管栖多毛体:多样性、古生态学及其与其他组合的比较
IF 1.8 3区 地球科学 Q1 Earth and Planetary Sciences Pub Date : 2022-01-01 DOI: 10.4202/app.01006.2022
J. Słowiński, O. Vinn, M. Jaeger, M. Zatoń
.
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引用次数: 5
Nazwy i logotypy stowarzyszeń i innych organizacji działających na rzecz g/Głuchych
IF 1.8 3区 地球科学 Q1 Earth and Planetary Sciences Pub Date : 2022-01-01 DOI: 10.17651/polon.42.4
Marta Wrześniewska-Pietrzak, Karolina Ruta-Korytowska
The article presents an analysis of the names and logotypes of companies, associations and other organiza-tions operating for the benefit of the d/Deaf in Poland. The analysis concerns the relationship between the name (its meaning) and the way of identifying the community of d/Deaf people. The description covers both the names and graphic elements, and their symbolism, which are present in the logotypes. The findings of cognitive methodology are used to read the correlation between the name, its logotype and the values identi-fied with the d/Deaf. The analysis covers 50 social chrematonims and logotypes of institutions from Poland who promote their activities on the Facebook social network or on Polish websites. The material collected allows one to notice that the most common way of identification with the d/Deaf community is the use of meanings and symbols of Polish Sign Language (PJM), which is one of the most important values creating the image of the Deaf as a linguistic and cultural minority.
本文分析了波兰为聋哑人服务的公司、协会和其他组织的名称和标志。分析了聋哑人群体的名称(其意义)与识别聋哑人群体的方式之间的关系。描述涵盖了名称和图形元素,以及它们的象征意义,这些都存在于标识中。认知方法的研究结果被用于阅读名称,它的标志和识别与聋哑人的价值观之间的相关性。该分析涵盖了50个在Facebook社交网络或波兰网站上宣传其活动的波兰机构的社会色彩和标志类型。收集到的材料使人们注意到,与聋人社区最常见的识别方式是使用波兰手语(PJM)的意义和符号,这是创造聋人作为语言和文化少数群体形象的最重要的价值观之一。
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引用次数: 0
Petrosal and cranial vascular system of the early Eocene palaeoryctid Eoryctes melanus from northwestern Wyoming, USA 美国怀俄明西北部早始新世古旋回动物黑纹旋回动物的岩质和颅骨血管系统
IF 1.8 3区 地球科学 Q1 Earth and Planetary Sciences Pub Date : 2022-01-01 DOI: 10.4202/app.00916.2021
Jean M. Wible
The petrosal and neighboring bones of the early Eocene palaeoryctid mammal Eoryctes melanus are described in tympanic and endocranial views based on CT scan data of the holotype. A second cranium of E. melanus has fragments of an osseous bulla, which have been interpreted as possibly formed by an independent entotympanic. The CT scans of the holotype reveal that the medial bullar wall is formed by an expanded rostral tympanic process of the petrosal, but the element(s) in the bullar floor remain unknown. The CT scans also allow for a comprehensive reconstruction of the cranial arterial and venous system. The arterial pattern differs from that in early eutherians by the absence of the arteria diploëtica magna and the bifurcation of the end branches of the stapedial artery dorsal to the tympanic roof. The venous pattern includes a large frontal diploic vein arising from the dorsal sagittal sinus on the midline and running through the frontal bone in a canal. Comparisons are made with other palaeoryctids, various Paleogene mammals (pantolestids, leptictids, apternodontids, apatemyids, and cimolestids) and the extant lipotyphlan Solenodon paradoxus. For the last taxon, the structure of the piriform fenestra and associated arteries is detailed. Cranial features support the monophyly of palaeoryctids and suggest possible lipotyphlan affinities.
基于CT扫描数据,对早始新世古周期哺乳动物黑斑古周期动物(Eoryctes melanus)的岩骨及其邻近骨骼进行了鼓室和颅腔图像的描述。另一种黑鳗的头盖骨有骨球的碎片,被解释为可能是由独立的内鼓室形成的。CT全型扫描显示,内侧壁是由扩大的岩骨吻侧鼓室突形成的,但仍不清楚黄底的组成部分。CT扫描也允许全面重建颅动脉和静脉系统。动脉形态与早期真兽不同,无diploëtica大动脉,镫骨动脉末端分支在鼓室顶背侧分叉。静脉形态包括一条大的额静脉,起源于中线上的背矢状窦,在椎管中穿过额骨。并与其他古兽类、各种古近系哺乳动物(甲兽类、瘦兽类、翼齿兽类、甲兽类和卷兽类)和现存的脂兽(lipotyphlan Solenodon paradoxus)进行了比较。对于最后一个分类群,梨状窗和相关动脉的结构是详细的。颅骨特征支持古周期动物的单系性,并提示可能的脂类亲缘关系。
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引用次数: 3
The paleoecology of the Late Miocene mammals from the Optima Local Fauna of Oklahoma, USA 美国俄克拉何马州Optima地方动物群中晚中新世哺乳动物的古生态学
IF 1.8 3区 地球科学 Q1 Earth and Planetary Sciences Pub Date : 2022-01-01 DOI: 10.4202/app.00941.2021
J. Frederickson, J. Cohen, M. Engel, Tyler C. Hunt, Greg A. Wilbert, O. Castañeda, N. Czaplewski
The Optima Local Fauna represents an important glimpse into the ecological transition between savannah and grassland during the late Miocene (Hemphillian) of what is now the southcentral Great Plains of North America. Though dominated by horses, herbivores from the Optima are morphologically diverse, bearing adaptations for both browsing and grazing lifestyles. Likewise, the carnivorans show similar ranges of size and presumed dietary behavior. In this study, we used carbonate isotope, mesowear, and tooth breakage and wear analyses to investigate the dietary complexity of mammals from a single site collected by the Oklahoma Museum of Natural History. Seventeen taxa were analyzed, including five perissodactyls ( Teleoceras hicksi, Dinohippus interpolatus, Neohipparion eurystyle, Nannippus ingenuus, and Astrohippus ansae ), four artiodactyls ( Texoceros guymonensis, Pediomeryx hemphillensis, Megatylopus matthewi, and Platygonus sp . ), a single proboscidean ( Mammut sp.), two rodents ( Dipoides indet. and Umbogaulus monodon ), and five carnivorans ( Agriotherium schneideri, Amphimachairodus coloradensis, Borophagus secundus, Eucyon davisi, Pliotaxidea cf. nevadensis ). Both stable isotope analysis and dental mesowear indicate a broad dietary partitioning occurred among the Optima herbivores, where the artiodactyls were identified as mixed feeders and the perissodactyls were recovered as grazers. In the carnivorans, the large felid Amphimachairodus coloradensis was a hypercarnivore with limited tooth breakage and an enriched δ 13 C signature, indicating low carcass utilization and a prey preference for horses. The canids had a more generalized diet, with B. secundus showing a greater proportional consumption of carcasses through a higher tooth breakage rate. The large ursid Agriotherium schneideri is here interpreted as an omnivore based on depleted δ 13 C values. Overall, we found evidence for a diversity of dietary niches in both carnivores and herbivores during the late Hemphillian in Oklahoma, likely driven by the expansion of grasslands in the region.
Optima当地动物群代表了中新世晚期(Hemphillian)大草原和草原之间生态转变的重要一瞥,现在是北美大平原的中南部。虽然以马为主,但来自Optima的食草动物在形态上是多样的,适应了浏览和放牧的生活方式。同样,食肉动物也表现出相似的体型范围和饮食行为。在这项研究中,我们使用碳酸盐同位素、中磨损、牙齿断裂和磨损分析来研究俄克拉何马州自然历史博物馆收集的单一地点哺乳动物的饮食复杂性。共分析了17个类群,包括5个异趾目(希克远角兽、内插恐龙、欧利波新河马、ingenuus Nannippus和Astrohippus ansae)和4个偶蹄目(古埃及特oceros guymonensis、hemphilensis Pediomeryx、matthewmegatylopus和Platygonus sp .)。一种长鼻目动物(Mammut sp.),两种啮齿类动物(Dipoides indet.)。以及5种食肉动物(Agriotherium schneideri, amphimacharodus coloradensis, Borophagus secundus, Eucyon davisi, Pliotaxidea cf. nevadensis)。稳定同位素分析和牙齿中磨损均表明,在Optima食草动物中存在广泛的饮食分配,其中偶蹄动物被确定为混合摄食动物,而异蹄动物被恢复为食草动物。在食肉动物中,大型猫科动物amphimacharodus coloradensis是一种具有有限牙齿断裂和丰富δ 13c特征的超级食肉动物,表明其较低的胴体利用率和对马的猎物偏好。犬科动物的饮食更广泛,第二齿犬科动物通过更高的牙齿断裂率,显示出更大比例的尸体消耗。大型熊科动物Agriotherium schneideri在此被解释为杂食动物,基于耗尽的δ 13c值。总的来说,我们发现了俄克拉荷马州亨菲利亚晚期食肉动物和食草动物饮食生态位多样性的证据,这可能是由该地区草原扩张所驱动的。
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引用次数: 0
Isometry in mesosaurs: implications for growth patterns in early amniotes 中龙的等距:对早期羊膜动物生长模式的影响
IF 1.8 3区 地球科学 Q1 Earth and Planetary Sciences Pub Date : 2022-01-01 DOI: 10.4202/app.00931.2021
Pablo A. Núñez Demarco, G. Piñeiro, Jorge Ferigolo
Isometry in Implications for growth patterns in early Mesosaurs were small amphibious tetrapods that lived in western Gondwana during the early Permian or even earlier, when temperate Carboniferous–Permian conditions initiated after the glaciations that affected the southern region of Pangea. In this contribution, we applied traditional linear regression morphometrics to analyse proportions of both the skull and limb bones in more than 100 mesosaur specimens. The analyses revealed that all mesosaur bones scale remarkably close to a model of geometrical similarity (isometry), and that this pattern is particularly strong in long bones and also in the skull. These results indicate that juvenile and adult mesosaurs do not display appreciable change in bone proportions, meaning that there are few or no noticeable differences between them during growth. The well-defined isometry, and particularly, the high interrelation between metatarsals and phalanges permit us to suggest that the mesosaur hind limb is subject to notable modularity. This evidence strongly argues that the differences previously described to support three mesosaur species in Western Gondwana, might instead reflect natural intraspecific variability, taphonomic features or even possible sexual dimorphism, as recently suggested. Our study also reinforces the general plesiomorphic structure of the mesosaur skeleton, which along with some cranial specializations for ecological fitness and the evidence of strong isometric growth as we demonstrate herein, may suggest new hypotheses of relationships for mesosaurs which thus would position them as more basal amniotes than previously thought.
早中龙是一种小型两栖四足动物,生活在早二叠纪或更早的冈瓦纳西部,在影响盘古大陆南部地区的冰期之后,温和的石炭纪-二叠纪条件开始了。在这篇文章中,我们应用传统的线性回归形态计量学分析了100多个中龙标本的头骨和肢骨比例。分析显示,所有的中龙骨骼都非常接近几何相似(等距)模型,这种模式在长骨和颅骨中尤为明显。这些结果表明,幼年和成年中龙在骨骼比例上没有明显的变化,这意味着它们在生长过程中几乎没有明显的差异。明确的等距,特别是跖骨和指骨之间的高度相互关系,使我们认为中龙后肢具有显著的模块化。这一证据有力地表明,之前描述的支持冈瓦纳西部三种中龙物种的差异,可能反映了自然的种内变异性、分型学特征,甚至可能是最近提出的两性二态性。我们的研究还强化了中龙骨骼的一般半形结构,以及我们在此展示的一些生态适应性的头骨特化和强等长生长的证据,可能提出了中龙关系的新假设,从而将它们定位为比以前认为的更基础的羊膜动物。
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引用次数: 2
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Acta Palaeontologica Polonica
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